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How to Identify the Hydrophyllaceae Family

Phacelia integrifolia scorpioid inflorescences with purple flowers are common among the Hydrophyllaceae family.
Phaclia integrifolia scorpioid inflorescences with purple flowers, common in the Hydrophyllaceae family
Page Last Updated May 5, 2026.

Introduction to the Hydrophyllaceae Family

When you learn to identify the Hydrophyllaceae family, you will quickly see some of their similarities with the Boraginaceae, which they were once included as a subfamily of until 2016.

Of all the small families split out of the Boraginaceae, this one looks the most like it, with its pretty blue and purple scorpioid cymes, especially my favorite genus Phacelia. I love their flowers, but I also love the stiff, often glandular hairs, divided leaves, and the interesting scents of the foliage in certain species, which some may consider unpleasant, but then again, for some reason I have always been biased; I tend to love smelly plants.

Common Botanical Description of the Hydrophyllaceae Family

If you’re new to plant morphology, this guide is a perfect beginner’s description, with no need to know any scientific jargon. Below this section is additional information on uses and morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems of the Hydrophyllaceae: They are always herbs that grow from taproots and are often stiff or sticky-hairy and may have a noticeable scent (pleasant or not). The leaves are found on stems and at the base, rarely one or the other, and are simple (undivided) or, more often, divided into pairs of leaflets in a compound leaf and may be arranged in opposite pairs or alternately along the stem. Leaf margins may be smooth, toothed, or lobed.

Flowers of the Hydrophyllaceae: Flowers contain both male (stamens) and female (ovary, style, stigma) parts in the same flower (bisexual). They are typically arranged in coiled clusters, and their parts are typically in 5s with 5 petals and a fused calyx with 5 lobes. The flowers are often blue to violet in color and often have scaly appendages inside their tubes.

Reproductive Features of the Hydrophyllaceae: There are 5 stamens, often attached inside the floral tube with widened or appendaged bases. The ovary is superior (sits above the attachment of the petals) with a single terminal style that is usually divided into 2 lobes.

Fruits of the Hydrophyllaceae: The fruits are dry capsules that typically split apart on two valves to release their seeds.

Uses of Hydrophyllaceae 

Several members of the Hydrophyllaceae are used ornamentally, especially in shade gardens, as well as for pollinator gardens and for use in erosion control. They also produce edible young shoots and roots that were historically used by Native Americans (e.g., Hydrophyllum species) and were also used medicinally for treating mouth sores and diarrhea.

Wildlife & Ecological Values of the Hydrophyllaceae Family

The Hydrophyllaceae family provides important nectar and pollen for native pollinators like bumblebees and long-tongued bees. They also are important early-season food sources and many aid in ecological succession by stabilizing soils and slopes.

Morphology of Hydrophyllaceae in North America

So far, I have only photographed the Phacelia genus, which is the most common and representative genus in the family, so all of these morphology photos are from that genus.

Learn how to identify the Hydrophyllaceae family with these morphology photos
Learn how to identify the Hydrophyllaceae family with these morphology photos

Some Hydrophyllaceae Species Found in North America

Phacelia bipinnatifida showy purple flower from Georgia, USA

Phacelia bipinnatifida—Purple Phacelia

The purple phacelia is a biennial herb up to 61 cm tall with showy rotate lavender-blue flowers up to 2 cm across, with appendaged stamens that often look fuzzy. It is typically found growing in cool moist woods and is native to the southeastern United States; this one was in a forest in Georgia.

Phacelia congesta or Blue curls showing scorpioid cymes common to the Hydrophyllaceae but in this case they are deeply congested

Phacelia congesta—Blue Curls

Blue curls is a leafy annual or biennial 30-90 cm tall with purple to lavender-blue deeply lobed flowers with stamens that extend beyond the lobes. Flowers occur in heavily congested scorpioid cymes, which uncurl as the flowers bloom. Leaves are deeply cut and may appear rough. It is native to New Mexico, Oklahoma, and Texas in the United States and south into northern Mexico.

Phacelia crenulata scorpioid inflorescence of pinkish-purple flowers

Phacelia crenulata—Notch Leaf Scorpionweed

This species is an aromatic annual, up to 80 cm tall, coated with stiff, glandular hairs. Leaves are oblong with wavy edges and get smaller up the stem. Flowers are purple or blue and may have a white throat. Stamens and style extend past the flower lobes. This native species has a disjunct distribution, found in the southwestern USA and northern Mexico but also in Peru, Bolivia, and Chile in South America.

Phacelia hirsuta plant with flowers showing its hirsute hairs common in the Hydrophyllaceae family

Phacelia hirsuta—Fuzzy Phacelia

Fuzzy phacelia is a small native annual known for its dense covering of stiff but non-glandular hairs. It only grows to about 45 cm tall, may or may not be branched, and the leaves are alternately arranged and deeply lobed to pinnately compound. This cute little Phacelia is an endemic native of the south-central United States; I found this one in Arkansas.

Phacelia integrifolia flowers closeup showing exserted stamens and bifid stigmas (two long points), common features in the Hydrophyllaceae family

Phacelia integrifolia—Gypsum Phacelia

People often easily identify this native Phacelia by its glandular hairs, which most find malodorous (but not me!) and sticky to the touch. It has leafy stems with simple leaves with crenate margins that often fold to the back of the leaf. It has the usual purplish to bluish scorpioid cymes of the family. It is native to the southwestern and south-central USA and northern Mexico, but most populations appear to be concentrated around New Mexico.

Phacelia linearis with saucer-shaped pink flowers but unusual linear leaves

Phacelia linearis—Linearleaf Phacelia

This Phacelia is native to the Pacific Northwest of the USA and British Columbia and Alberta, Canada. It is easily identified by its location as well as its linear leaves, which are not common in the genus and are just visible (blurry) beneath the flower in this photo.

Phacelia popei herbaceous annual, common among the Hydrophyllaceae, this one growing in the moisture of a septic field in a desert in New Mexico.

Phacelia popei—Pope’s Phacelia

This lovely Phacelia is one of my favorites, for its intense purple (sometimes pink) flowers in the usual scorpioid cymes. It is an annual that only grows to about 40 cm tall with leaves that range from “normal” to rather wide and pinnately divided. It is a somewhat rare species, found only in New Mexico and western Texas and just south of the border into northern Mexico.

Scientific Botanical Description of the Hydrophyllaceae

Habit & Leaf Form of the Hydrophyllaceae Family

The Hydrophyllaceae are annual, biennial, or perennial herbs growing from taproots with erect to prostrate stems, usually with an indumentum that is scabrid, hispid, or glandularly hairy, and sometimes the hairs can sting (e.g., as in Phacelia) and often have noticeable scents, from pleasant to unpleasant.

Leaves are simple to bipinnately compound, basal and cauline both (rarely only one or the other), and are arranged oppositely or alternately along the stem. They can be simple, but most are usually bipinnately divided. Secondary venation is typically palmate, and there are no stipules. Lamina margins may be smooth, toothed, or lobed.

Flowers of the Hydrophyllaceae Family

Flowers are bisexual and are arranged in terminal or axillary thyrsoidal, usually helicoid or scorpioid cymes that are sometimes congested. There are generally no bracts or bracteoles.

Flowers are usually 5(4)-merous. The calyx is fused at the base or nearly to the tip, with usually 5 lobes, linear to cordate, with valvate aestivation, and is generally persistent and enlarging in fruit.

The corolla also typically has 5 lobes, is rotate to cylindric, is generally deciduous, and often has paired appendages in the tube between the filaments. The corolla is also fused into a tube with lobes, often blue or blue-violet to purple but can also be white, pink, or yellow; corolla scales are present, small, or absent, and aestivation is contorted or imbricate.

Androecium of the Hydrophyllaceae Family

There are typically 5 stamens of equal or unequal length, with filaments that are often expanded and/or lobed at the base, often with scaly appendages, and they are typically epipetalous (adnate to the corolla tube).

Gynoecium of the Hydrophyllaceae Family

The Hydrophyllaceae gynoecium usually comprises a superior ovary, 1-2 carpels, with 2 swollen placentas with axile or (stalked) parietal placentation; usually placentation is intrusive parietal. Sometimes the ovary may appear 2-5-chambered via the intrusive, swollen placenta, but it does not appear lobed or only shallowly so. A nectar disc is often present at the ovary base, but it is sometimes reduced to glands. There is 1 terminal style, usually divided about halfway with punctate, capitate, or capitate-funneliform stigmas. There are 2 to many ovules that are epi-/apo-/pleurotropous, with integuments ca. 6 (-12) cells across.

Fruit of the Hydrophyllaceae Family

The fruit is always a dry membraneous capsule; it may be septicidal or loculicidal, generally with 2 valves, 1 to many seeds. Part of the persistent calyx is adnate to the fruit. Seeds ruminate by the inpushing of the exotestal cells/endothelium; exotestal cells are thickened on inner and radial walls; endotestal cells are persistent. The endosperm is variously copious to scanty.

Taxonomy of Hydrophyllaceae

There are about 320 species in 12 (16) genera in the Hydrophyllaceae family in the Boraginales order of the core eudicots (dicots). This family has been treated as distinct from Boraginaceae and then as the Hydrophylloideae subfamily of it. However, a 2016 revision based on modern phylogenetics has confirmed its status as a distinct family, including the removal of the Namaeae as its own family, the Namaeceae. For more information, check out Luebert et al. (2016) in the references for a great explanation and justification of the split. ‌

Genera of the Hydrophyllaceae:

Draperia (1), Ellisia (1), Emmenanthe (1), Eucrypta (2), Hesperochiron (2), Howellanthus (?), Hydrophyllum (10), Nemophila (13), Phacelia (206), Pholistoma (3), Romanzoffia (5), and Tricardia (1).

Key Differences From Similar Families

The Hydrophyllaceae had been included as a subfamily of Boraginaceae before, though it has recently been made a separate family again. They have very similar-looking scorpioid inflorescences in similar blue-to-violet colors. However, their fruits differ because Hydrophyllaceae always produce a dry, dehiscent capsule, while the Boraginaceae mostly produce indehiscent nutlets, and the Boraginaceae typically have a deeply 4-lobed ovary with a gynobasic style, while the Hydrophyllaceae have a mostly unlobed ovary with a terminal style with 1-2 stigmatic branches.

The Heliotropiaceae has also recently been split out of the Boraginaceae, and it too has scorpioid cymes. However, it can be distinguished from the Hydrophyllaceae by its terminal style with a unique cone-shaped stigmatic head and basal ring-shaped stigma that may be sterile at the tip (the stigma is not terminal) and fruits that are mostly fleshy drupes with 4 stones or are dry schizocarps.

The Namaeceae was, until recently, included as a tribe when the Hydrophyllaceae was the Hydrophylloideae subfamily in the Boraginaceae, but it too has now received family status. This small family is mostly shrubs or small trees (with the exception of Nama which is herbaceous), and they have only simple, cauline leaves, not basal. They also have 2 stylodia, or their style is united for 3/4 of its length with two stigmatic branches.

The Ehretiaceae and Cordiaceae are also recent splits from the Boraginaceae, but these families are both mostly shrubs and trees instead of herbs, and both mostly produce fleshy drupes for their fruits instead of dry capsules.

Distribution of Hydrophyllaceae

The Hydrophyllaceae are a family primarily located in western North America and western South America, from Arctic Alaska south through to southern Patagonia. They are especially common in drier areas of southwestern North America. While less common, they can also be found in eastern North America, but not eastern South America. The Hydrophyllaceae are not naturally found outside of the Americas but the Phacelia have been introduced to parts of Europe and Asia.

Distribution of Hydrophyllaceae in the Americas

Canadian Genera Include:

Ellisia monospecific NAM endemic native from BC east to ON; Hesperochiron 1 W NAM endemic sp. native to BC; Hydrophyllum 5 NAM endemics native to S Canada excluding SK and the maritime provinces; Nemophila 3 former NAM endemics native to BC and AB; Phacelia 13 spp. native to most of Canada except NU, QC, and the maritime provinces; Romanzoffia 2 W NAM endemics native to BC and AB.

USA Genera Include:

Draperia monospecific endemic of CA; Ellisia monospecific NAM endemic, and native to most of the central and eastern USA; Emmenanthe monospecific NAM endemic native CA, NV, UT, and AZ; Eucrypta 2 S NAM endemic spp. native to CA, NV, UT, AZ, NM, and TX; Hesperochiron 1 W NAM endemic sp. native to the western half of the US, E to MN, CO, and AZ; Howellanthus monospecific NAM endemic native to CA (Syn. of Phacelia?); Hydrophyllum 10 NAM endemics native to all of the continental states except for ME, TX, and FL; Nemophila 11 former NAM endemics native to most of the W USA and the SE USA; Phacelia 172 spp. native to all of the USA except FL; Pholistoma 3 S NAM endemics native to OR, CA, NV, and AZ; Romanzoffia 5 NAM endemics native to WA, ID, MT, OR, and CA; Tricardia monspecific USA endemic of CA, NV, UT, and AZ.

Mexico Genera Include:

Emmenanthe monospecific NAM endemic native to NW Mexico; Eucrypta 2 S NAM endemic spp. native to NW Mexico; Hesperochiron 1 W NAM endemic sp. native to NW Mexico; Nemophila 1? sp. NAM endemics native to NW Mexico; Phacelia many, ~100?? spp. native throughout Mexico; Pholistoma 3 S NAM endemic spp. native to NW Mexico.

Neotropical Genera Include:

Phacelia ~9 spp. native to Guatemala, Argentina, Bolivia, Peru, Uruguay, and Chile.

Patagonia Genera Include:

Phacelia ~5-7 spp. native throughout the Patagonia region.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, along with my own personal observations throughout North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • FNA (1993+). Flora of North America. https://floranorthamerica.org/Main_Page. Accessed 2022-current.
  • GBIF.org (2020+), GBIF Home Page. Available from: https://www.gbif.org
  • iNaturalist.org (2020+). https://www.inaturalist.org/. Accessed 2020-current.
  • Luebert, F., Cecchi, L., Frohlich, M. W., Gottschling, M., Guilliams, C. M., Hasenstab-Lehman, K. E., Hilger, H. H., Miller, J. S., Mittelbach, M., Nazaire, M., Nepi, M., Nocentini, D., Ober, D., Olmstead, R. G., Selvi, F., Simpson, M. G., Sutorý, K., Valdés, B., Walden, G. K., & Weigend, M. (2016). Familial Classification of the Boraginales. axon65(3), 502–522. https://plants.sdsu.edu/amsinckiinae/pdfs/Luebert_etal2016-Boraginales.pdf.
  • ‌Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019+). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Stevens, P. F. (2001+). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; Accessed 2020-present.
  • Walden, Genevieve K., Robert W. Patterson & Richard R. Halse. (2023), Hydrophyllaceae, in Jepson Flora Project (eds.) Jepson eFlora, Revision 12, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=157, accessed on April 25, 2026.
  • Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Ehretiaceae Family

Tiquilia canescens Shrubby Tiquilia - Learn to identify the Ehretiaceae family
Tiquilia canescens aka Shrubby Tiquilia, a native Ehretiaceae of North America.
Page Last Updated April 26, 2026

Introduction to the Ehretiaceae Family

The Ehretiaceae are a small family that was only somewhat recently (2016) pulled out of the closely related Boraginaceae family. I only discovered this somewhat recently since so many sources still refer to it as a subfamily of the Borage Family instead. This guide explains how to identify the Ehretiaceae family and how they differ from the new, narrower definition of the Boraginaceae.

My only experience with this mostly tropical family is with Tiquilia, which is a lovely, low-growing herb that is relatively common in the dry soils of the American Southwest. It has pretty, almost succulent leaves, and I always thought it would make a nice ground cover in a xeriscape. However, this means that all of my photos are of that genus. For people living in North America, this is by far the most common member of this family that you will see, although there are other shrubby, small tree, and parasitic species, especially once you get into Mexico. I will add more as I find them!  

Common Botanical Description of the Ehretiaceae Family

If you’re new to plant morphology, this guide is a perfect beginner’s description for learning to identify the Ehretiaceae family without needing to know any scientific jargon. Below this section is additional information on uses and morphology, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems of the Ehretiaceae: This family is mostly woody shrubs or trees, often with fibrous bark and whitish twigs, but perennial herbs and parasitic plants that don’t use chlorophyll are also seen. Leaves are simple (not compound) and arranged alternately along the stem on stalks, although sometimes they are reduced to tiny scales. Margins may be smooth or sometimes with rounded teeth.

Flowers of the Ehretiaceae: Flowers usually have both male (stamens) and female (ovary, stigma, and style) parts in the same flower (bisexual), but occasionally separate male and female flowers are seen on separate male and female plants (called dioecious). Flowers and sepals are usually fused at the base with 5 lobes.

Reproductive Features of the Ehretiaceae: There are typically 5 stamens that often stick out past the throat of the flower tube and are sometimes attached to the petals. The ovary is usually surrounded by a nectary disk, and it has a style on top that is typically divided into 2 branches.

Fruits of the Ehretiaceae: Fruits are mostly drupes (think of a cherry with their stony pits) with 1-2-seeded stones or 4 nutlets.

Uses of Ehretiaceae 

The Ehretiaceae, especially the Ehretia genus, are widely used for medicine (pain, fever, and dysentery), timber and furniture, and their edible drupes, especially in Asia, Africa, and parts of the Americas. They are used for treating ailments like fever, dysentery, and pain, with wood for furniture and edible fruits.

Morphology of Ehretiaceae in North America

As I currently only have the herbaceous-looking Tiquilia genus, which is not particularly representative of the family, I will add morphology photos when I acquire more photos.

Some Ehretiaceae Species Found in North America

Tiquilia canescens Shrubby Tiquilia from near Carlsbad National Park

Tiquilia canescens—Shrubby Tiquilia

The shrubby tiquilia looks a lot more like an herb, but technically you could call it a shrub or sub-shrub because it is woody at the base. It is a short, very low-growing, spreading plant with gray-green, fleshy, egg-shaped leaves that look almost succulent and are often rather congested on its branches. Its 5-lobed tubular flowers are white to pinkish. It is native to the American Southwest in California, Nevada, Arizona, Utah, New Mexico, and Texas and in Northern Mexico in Chihuahua, Sonora, and Baja California.

Tiquilia greggii Plumed Crinklemat in flower, Dog Canyon, New Mexico.

Tiquilia greggii—Plumed Crinklemat

The plumed crinklemat is another herbaceous-looking low-growing shrub, generally with wider internodes, giving it a more spindly appearance. The leaves are also gray-green and somewhat succulent, but what stands out most are the feathery calyxes that appear in rounded masses with usually only 1-2 flowers blooming at a time. The feathery calyx apparently persists and assists in dispersing the seeds. This one is more rare, and in the United States it is only found in southern New Mexico and the southwestern corner of Texas, being most common throughout northern and central Mexico instead.

Scientific Botanical Description of the Ehretiaceae

Learn to identify the Ehretiaceae family based on its new narrow definition after being removed from the Boraginaceae.

Habit & Leaf Form of the Ehretiaceae Family

This family is mostly woody shrubs or trees, often with fibrous bark, oxidizing stems, and whitish twigs and rarely with thorns (Rochefortia). Perennial herbs are also seen, especially Tiquilia. The indumentum is variable from hirsute to glabrescent. Also, plants in the Pholismateae tribe are achlorphyllous root parasites with glandular hairs and leaves that are reduced to scales.

Leaves are simple, alternately arranged, petiolate, with margins that are entire to crenate, or occasionally dissected (in Cortesia), and without stipules. In achlorophyllous root parasites, the leaves are typically reduced to scales.

Flowers of the Ehretiaceae Family

Inflorescences are axillary or sometimes terminal, usually laxly cymose or thyrsoidal, sometimes congested, and occasionally in few-flowered corymbs or solitary. In parasitic plants, they are condensed and more or less capitate.

Flowers are mostly bisexual, although occasionally plants are dioecious (Lepidocordia and Rochefortia). Flowers are 5-merous, usually with radial symmetry. The often long, persistent calyx has 5 lobes, fused or may be free to the base; aestivation is quincuncial or sometimes imbricate. The corolla is sympetalous and tubular with 5 spreading lobes; it is usually rotate, campanulate, or urceolate; aestivation is imbricate, and they are white, red, or sometimes blue.

Androecium of the Ehretiaceae Family

There are 5 stamens that are often epipetalous, fused to the floral tube at least at the base, and may be puberulent at the base; anthers are generally exserted.

Gynoecium of the Ehretiaceae Family

The ovary is bicarpellate, 1-4-locular from secondary division, superior, and generally subtended by a disk-like nectary. There is a bifid terminal style with an elongated clavate to capitate stigma. Placentation is apical to axile. There is usually 1 fertile ovule per carpel, apotropous, with integument 6-12 cells across, and epidermal cells anticlinally elongated or not. Parasitic members may have 10 or more ovules.

Fruit of the Ehretiaceae Family

Fruits are drupes, often surrounded by a persistent, accrescent calyx. They often dry and separate into 2 2-seeded pyrenes, 4 1-seeded pyrenes or schizocarps, or 4 nutlets. Seeds often have copious endosperm, but some seeds have none.

Taxonomy of Ehretiaceae

There are about 155 species in 10(7) genera in the Ehretiaceae family in the Boraginales order of the core eudicots (dicots). The Ehretiaceae family has frequently been treated as a subfamily of the Boraginaceae. A 2016 revision by the Boraginales Working Group, however, confirmed its status as a distinct family, and this is what is also listed on the Angiosperm Phylogeny Group, which I use as my most current authoritative source. This definition of the Ehretiaceae includes the Lennoaceae nested within it.

Genera:

Bourreria (52), Cortesia (?), Ehretia (66), Halgania (16), Keraunea (5), Lennoa (1), Lepidocordia (2), Pholisma (3), Rochefortia (9), and Tiquilia (28).

Key Differences From Similar Families

The Ehretiaceae was recently split from the Boraginaceae, but the Boraginaceae can be distinguished by usually being herbs and virtually always having a gynobasic style rather than a terminal one in the Ehretiaceae.

The Cordiaceae was also recently split from the Boraginaceae, and it too is mostly woody trees and shrubs with fleshy drupes, but they have characteristic 4-lobed rather than bifid stigmas; if they do have 2 stigmatic branches, then the fruit is enclosed in an accrescent calyx.

The Heliotropiaceae has also recently been split out of the Boraginaceae family, but it’s a family of more herbs or subshrubs. However, it does have some trees that also produce drupes and could be confused. But, it can usually be distinguished by its often deeply lobed ovary and unique cone-shaped stigma.

Distribution of Ehretiaceae

The Ehretiaceae are a mostly tropical and subtropical family throughout the tropics, though noticeably absent from eastern South America. There is also a strong presence of this family in the arid American Southwest.

Distribution of Ehretiaceae in the Americas

Canadian Genera Include:

Absent.

USA Genera Include:

Bourreria 3 spp. native to FL; Ehretia 1sp. native in TX; Pholisma 2 S NAM endemic spp. native in CA and AZ; Tiquilia 9 spp. native in all of W half of the USA, except for MT, also in MO.

Mexico Genera Include:

Bourreria 16 spp. native to all of Mexico; Ehretia 4 spp. native to all of Mexico; Lennoa monospecific endemic to Mexico and the N neotropics; Lepidocordia 1 N neoendemic sp. native to SW and SE Mexico; Pholisma 3 S NAM endemic spp. native to northern Mexico; Rochefortia 1 sp. native to much of Mexico except NW and C; Tiquilia ~12 spp native throughout all of Mexico.  

Neotropical Genera Include:

Bourreria ~30? spp. native CAM, Antilles, Colombia, Venezuela, Ecuador; Ehretia ~3 spp. native in CAM, Antilles, N+E Brazil, and N Argentina; Keraunea 5 spp. endemic to eastern Brazil; Lennoa monospecific endemic of Mexico, CAM, Colombia, and Venezuela; Lepidocordia 2 N neoendemic spp. native to Mexico, El Salvador, Honduras, Nicaragua, Venezuela, Guyana, and N Brazil; Rochefortia 9 neoendemic spp. native to Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela, and Peru; Tiquilia ~20 spp. native to Ecuador, Peru, Bolivia, N Chile, and NW+S Argentina, including 4? narrow endemics of the Galapagos.

Patagonia Genera Include:

Tiquilia 1? sp. native to S Argentina.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, along with my own personal observations throughout North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • FNA (1993+). Flora of North America. https://floranorthamerica.org/Main_Page. Accessed 2022-current.
  • GBIF.org (2020+), GBIF Home Page. Available from: https://www.gbif.org
  • iNaturalist.org (2020+). https://www.inaturalist.org/. Accessed 2020-current.
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019+). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Michael G. Simpson 2021, Ehretiaceae, in Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=84713, accessed on April 24, 2026.
  • Stevens, P. F. (2001+). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; Accessed 2020-present.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

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How to Identify the Boraginaceae Family

Anchusa officinalis inflorescence with violet blue flowers. Learn to identify the Boraginaceae family.
Anchusa officinalis inflorescence with violet blue flowers. Learn to identify the Boraginaceae family!
Page Last Updated May 15, 2026.

Introduction to the Boraginaceae Family

The Boraginaceae family has undergone major revisions over the years, including several small families and then more recently, having them removed. This description teaches how to identify the Boraginaceae based on the currently accepted, much narrower definition of the family that excludes Heliotropiaceae, Hydrophyllaceae, and others, which I will all cover separately.

What I love about this family is their often instantly blue-to-violet flowers almost always arranged in unique clusters called “scorpioid cymes.” They are truly beautiful; I even love their coarse hairs that can cause rashes in some people, but to me, it just makes them even more beautiful. But I might be biased too, since I think all flowers are beautiful in their own way.

Common Botanical Description of the Boraginaceae Family

If you’re new to plant morphology, this common botanical description is a perfect beginner’s description for learning to identify the Boraginaceae family, with no need to know any scientific jargon not explained in the description. Below this section is additional information on uses and morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions, taxonomic information, and genus-level distribution data in North, Central, and South America.

Leaves and Stems of the Boraginaceae: Most members are herbaceous plants, with occasional woody shrubs, trees, or vines, often accompanied by characteristic coarse hairs that can cause skin reactions. The leaves vary in size but are always simple (not compound) and are usually arranged alternately along the stem; however, in rare cases, the lower leaves may be in opposite pairs. Leaves are usually thin to lance-shaped, and most have entire margins, but in rare cases they may be toothed. Another characteristic feature of the family is that the leaves never have small leaf-like appendages called stipules at their base, often seen in other families.

Flowers of the Boraginaceae: The flowers of this family are mostly bisexual, containing both male (stamens) and female (ovary, style, stigma) parts in the same flower. What is most characteristic of this and closely related families is the flowers in coiled clusters called “scorpioid cymes.” Boraginaceae flowers are usually regular, meaning they can be divided in half on multiple planes of symmetry, and contain usually 5 sepals that may be free or joined at the base into a tube and are often covered with hairs. They also have 5 petals that are usually blue or purple (but may be pink, yellow, or white) and often have a collection of scales in the throat.

Reproductive Features of the Boraginaceae: The stamens can vary, being free or joined, but there are always five fertile stamens (with pollen-producing anthers), and they are always attached in the throat of the flower. The ovary is deeply four-lobed, and there is a single style attached at its base.

Fruits of the Boraginaceae: The fruits are always non-fleshy nutlets or sometimes dry, dehiscent fruits called schizocarps that split apart when mature.

Uses of Boraginaceae 

Many members of the family are used ornamentally for their lovely flowers, while others are used medicinally, and still others are used as a natural source of dye.

Common cultivated family members include forget-me-not (Myosotis), Symphytum, Borago, and more. Many species have been introduced as ornamental plants or weeds, e.g., the genera Anchusa, Borago, Cynoglossum, Echium, Myosotis, and Symphytum.

Morphology of Boraginaceae in North America

Learn to identify the Boraginaceae family with morphology photos
Learn to identify the Boraginaceae family with morphology photos

Some Boraginaceae Species Found in North America

Boraginoideae Subfamily

Anchusa officinalis inflorescence with violet blue flowers characteristic of the Boraginaceae

Anchusa officinalis—Common Alkanet

This beautiful introduced wildflower has long, lance-shaped leaves that become smaller on shorter petioles further up the stem. Most of the plant is covered in fine, stiff hairs, and it produces deep violet-blue flowers with five spreading, rounded lobes. It is native primarily to the Mediterranean region, but also to most of Europe and western Asia. It is now an occasional introduced weed in North America, like this one growing wild not far from human settlements in Peachland, British Columbia, Canada.

Buglossoides arvensis leaves and flowers closeup

Buglossoides arvensis—Corn Gromwell

An annual herb up to 40 cm long with stems that branch from the base, producing terminal racemes of pale blue to white flowers with forward-facing hairs on the outside and narrowly lance-shaped leaves that typically have a central groove on the upper surface. Native throughout Eurasia and northern Africa but widely introduced in Canada and the United States, as well as southern South America. This one was found in a well-maintained park in Vernon, British Columbia, Canada.

Lithospermum ruderale plant with flowers closeup showing leaves and flowers

Lithospermum ruderale—Wayside Gromwell

Unlike most members of the Boraginoideae subfamily, this perennial herb is native to North America, endemic to western North America from British Columbia east to Saskatchewan in Canada and from Washington State south to California and east to Montana south to Colorado in the United States. It also produces narrowly lance-shaped leaves with fine, coarse hairs and light yellow tubular flowers. It is common in dry soil with good drainage in sunny locations in its range. This one was in Princeton, British Columbia.

Cynoglossoideae Subfamily

Andersonglossum virdinianum plant with flowers closeup

Andersonglossum virginianum—Southern Wild Comfrey

This pretty wildflower has large comfrey-like leaves that give it one of its common names. The leaves are slightly hairy and tend to clasp the hairy stem at their base. It produces pale bluish flowers on erect, branching stems that often turn a violet color as they age. This is an uncommon endemic native of the eastern United States from Texas east to Florida and north to Vermont, where it normally grows in forested areas.

Cryptantha crassisepala scorpioid inflorescence with hairs and flowers

Cryptantha crassisepala—Thick-Sepaled Cryptanth

This very hairy annual herb only grows up to about 15 cm tall, with 2-6 cm long leaves and small white flowers with very thick and hairy sepals. However, examination of fruits is often needed for a positive ID. It is a southern North American endemic from California east to Kansas and Texas as well as throughout northern Mexico. It tends to grow in arid and semi-arid locations; this one was on a roadside in New Mexico.

Oreocarya aka Cryptantha flava inflorescences with flowers

Oreocarya aka Cryptantha flava—Yellow Cryptantha

Most sources still refer to Oreocarya as a separate genus, but authoritative sources like APG and USDA now consider it a synonym for Cryptantha, so we use both names here. This narrow endemic of Colorado, Wyoming, Arizona, Utah, and New Mexico in the United States is known for its cheerful yellow flowers and thinner but still very hairy sepals. This one was in the Wilson Arch area, Utah.

Oreocarya fulvocanescens var. fulvocanescens plant with flowers showing floral appendages

Oreocarya aka Cryptantha fulvocanescens—Gray Cat’s Eye

Another lovely Cryptantha endemic to the American Southwest, this time in Nevada, Utah, Colorado, Arizona, and New Mexico. It has white tubular flowers with conspicuous yellow appendages in their throats and the hairy sepals we are used to seeing in the genus. This one was found growing on a dry roadside outside of Cuba, New Mexico.

Myosotis arvensis forget me nots inflorescence with flowers

Myosotis arvsenis—Field Forget-Me-Not

An annual or short-lived perennial herb with stalkless, hairy leaves and pretty inflorescences of blue flowers with white to yellow throat appendages. The flowers are in typical scorpioid cymes, but they are best visualized while the flowers are still in bud as they elongate and straighten as they bloom. This species is originally native to northern Eurasia but is now widely introduced to northern North America. This was on the side of a logging road near Sechelt, British Columbia, Canada.

Myosotis macrosperma flowers and leaves

Myosotis macrosperma—Largeseed Forget-Me-Not

An annual herb with hairy, stalkless leaves with a prominent midrib and tiny white flowers in condensed cymes with hairy sepals and green bracts. This one is a North American native found in disturbed areas throughout the southeastern United States north into southern Ontario, Canada.

Lappula occidentalis plants with flowers and fruits

Lappula occidentalis—Western Stickseed

This hairy annual has stalked basal leaves that are mostly oblong with rounded tips and narrower, stalkless stem leaves with pointed tips. They are best known for their elongated inflorescences of tiny white to pale blue flowers, followed by fruits containing four nutlets, each with a row of slender prickles. It is native throughout western North America from Alaska south to northern Mexico and east to Manitoba, Canada, and Iowa, United States, with some disjunct populations in the eastern United States.

Scientific Botanical Description of the Boraginaceae

Learn to identify the Boraginaceae family based on the newer, more narrow definition of the family.

Habit & Leaf Form of the Boraginaceae Family

Most are annual or perennial herbaceous plants from taproots with occasional woody shrubs, trees, or self-supporting (usually) vines. They are often hispid or scabrid with coarse hairs that may contain SiO₂ or CaCO₃, which can cause adverse skin reactions in sensitive individuals. They may or may not have a basal aggregation of leaves. Helophytic, mesophytic, or xerophytic.

Leaves are minute to medium-sized, usually alternate or sometimes alternate and opposite below. They are simple, flat, herbaceous, petiolate to sessile, usually non-sheathing or sometimes sheathing in basal rosettes, not gland-dotted, and epulvinate. The lamina is entire, usually narrow, linear to lanceolate. Leaves lack stipules. Lamina margins are mostly entire, rarely crenate or dentate. Domatia may be present.

Flowers of the Boraginaceae Family

Plants are usually hermaphrodites or occasionally gynodioecious (Echium), with predominantly entomophilous pollination via Hymenoptera. Flowers are usually aggregated in coiled (scorpioid) cymes that may appear doubled in some species; coiled cymes typically elongate in fruit. Rarely are flowers solitary. Inflorescences are terminal or axillary.

Flowers are bracteate or not, usually regular or irregular and somewhat zygomorphic (e.g. Echium); usually 5-merous and tetracyclic. Free hypanthium is not present. Hypogynous disk present or absent. Perianth has a distinct calyx and corolla with 10 (11–13) parts in 2 whorls; mostly isomerous or sometimes anisomerous (Plagiobothrys).

Calyx 5 (usually) or 5–8 (sometimes Plagiobothrys) in 1 whorl; they may be free or connate basally with lobes shorter to longer than the tube; the degree of gamosepaly is 0.1–0.5; often covered with trichomes. Calyx persistent; imbricate, open in bud or valvate (rarely).

Corolla 5 parts in 1 whorl, often appendiculate with a corona of scales from the throat protecting the nectar. Petals are fused into a tube with lobes shorter to longer than the tube. Corolla is imbricate or contorted; rotate, campanulate to hypocrateriform or tubular; unequal, regular, or often bilabiate in Echium. They are often blue or purple in color but may also be pink, white, or yellow.

Androecium of the Boraginaceae Family

There are 5 unbranched androecial members made of exclusively fertile stamens that are adnate midway down or in the throat of the corolla tube; all are equal or unequal, free or coherent, and 1-whorled. Stamens are not didynamous or tetradynamous; they are isomerous with the perianth, arranged opposite the sepals, with filaments, or sometimes appearing sessile, inserted or exserted. Filaments are appendiculate or not. Anthers are cohering, free, or connivent; dorsifixed to basifixed; dehiscing via longitudinal slits; introrse; tetrasporangiate; appendaged or unappendaged.

Gynoecium of the Boraginaceae Family

The gynoecium is usually 2-carpeled or 4–5 carpeled (some Trigonotis). Pistil 2- or 4-celled (usually via false septa) or 8-10-celled (via false septa in some Trigonotis). The gynoecium is syncarpous with a superior ovary that is nearly always 2 locular but often appears 4 locular via false septa. The gynoecium is median. Style 1 is always gynobasic from an often deeply 4-lobed ovary. Stigmas 1–2: when simple, they are often 2-lobed but may also be capitate or minute; nearly always dry type; papillate. Placentation is basal. The ovules are 2 per locule, usually separating into 1-ovuled portions; they are positioned horizontally to ascending, epitropous, and have a dorsal raphe.  

Fruit of the Boraginaceae Family

The fruits are primarily non-fleshy nutlets, dehiscent, indehiscent, or a schizocarp with 4 or 8–10 (sometimes Trigonotis) mericarps.

Taxonomy of Boraginaceae

There are 1,793 species in 94 genera in the Boraginaceae family in the Boraginales order of the core eudicots (dicots). This family recently underwent major revisions, including the removal of the Hydrophyllaceae, Heliotropiaceae, Cordiaceae, Ehretiaceae, and others based on modern molecular phylogenetics. As a result, many genera have changed in the family, but the genera listed immediately below are accurate as per APG IV. For more information on the justification of this split of the Boraginales, I encourage you to check out Luebert et al. (2016) in the references below.

There are 3 currently accepted subfamilies:

  1. Boraginoideae is a large subfamily concentrated in the Mediterranean region, with some also in East Asia, Africa, and South America, but it is also introduced elsewhere. They are mostly perennial herbs, some annuals, and some shrubs; their flowers typically have a single line of symmetry and basal scales, the gynobase is flat, and the nutlets have a basal attachment scar.
  2. Cynoglossoideae is the largest subfamily with a nearly cosmopolitan distribution. It is a group of mostly herbs (annual to perennial) with some shrubs, leaves with obvious secondary venation, bractless flowers, a broadly pyramidal to flat gynobase, and often compressed nutlets that are ventrally attached with a large triangular scar.
  3. Echinochiloideae is a small subfamily with only 3 genera located in Mexico, south to northeastern Argentina, plus the Canary Islands, Africa, and east to western India. They are characterized by having densely hairy corolla throats and punctate stigmas (subterminal, sterile tips, bilobed or notched) with fruits that are laterally compressed nutlets.

Genera:

Boraginoideae: Aegonychon (3), Alkanna (68), Anchusa (35), Arnebia (36), Borago (5), Brunnera (3), Buglossoides (6), Cerinthe (6), Cynoglottis (2), Cystostemon (16), Echiostachys (3), Echium (68), Gastrocotyle (3), Glandora (8), Halacsya (1), Hormuzakia (3), Huynhia (2), Lithodora (3), Lithospermum (84), Lobostemon (28), Maharanga (43), Mairetis (1), Megacaryon (1), Melanortocarya (1), Moltkia (6), Moltkiopsis (1), Moritzia (3), Neatostema (1), Nonea (46), Onosma (247), Paramoltkia (1), Pentaglottis (1), Phyllocara (1), Podonosma (4), Pontechium (1), Pulmonaria (18), Stenosolenium (1), Symphytum (27), Thaumatocaryon (2), Trachystemon (1).

Cynoglossoideae: Adelinia (1), Afrotysonia (3), Amsinckia (13), Ancistrocarya (1), Andersonglossum (3), Anoplocaryum (4), Antiotrema (1), Asperugo (1), Bothriospermum (6), Brachybotrys (1), Caccinia (4), Chionocharis (1), Craniospermum (11), Cryptantha (104), Cynoglossum (83), Dasynotus (1), Decalepidanthus (9), Eremocarya (2), Eritrichium (78), Greeneocharis (2), Gyrocaryum (1), Hackelia (56), Harpagonella (2), Heterocaryum (1?), Iberodes (5), Johnstonella (17?), Lappula (81), Lasiocaryum (3), Memoremea (1), Mertensia (52), Microcaryum (1), Microparacaryum (4), Microula (34), Mimophytum (11), Myosotidium (1), Myosotis (157), Nesocaryum (1), Nihon (5), Omphalodes (11), Oncaglossum (1), Oreocarya (67? or syn. of Cryptantha), Pectocarya (13), Plagiobothrys (64), Pseudoheterocaryum (4? or syn. Suchtelenia), Pseudolappula (1), Rochelia (22), Selkirkia (4), Solenanthus (23), Suchtelenia (6), Thyrocarpus (4), Trichodesma (38), Trigonotis (70).

Echinochiloideae: Amphibologyne (1), Antiphytum (13), Echiochilon (16), Ogastemma (1).

Key Differences From Similar Families

The Hydrophyllaceae had been included as a subfamily of Boraginaceae before, though it has recently been removed and made a separate family again. They have very similar-looking scorpioid inflorescences in similar blue-to-violet colors. However, the Hydrophyllaceae always have dry capsule fruits, while the Boraginaceae have nutlets. The ovaries and seeds also differ.

The Heliotropiaceae has also recently been split out of the family, and it too has scorpioid cymes, though not all members do. However, it can be distinguished by having fewer bristly hairs that are always eglandular, leaves that are usually conduplicate, an ovary that is still often deeply lobed but has a cone-shaped stigma, and fruits that are drupes with 4 stones or are dry schizocarps.

The Ehretiaceae is another recent split. It is a small family with only 7 genera of mostly shrubs and trees as opposed to mostly herbs, and while the Boraginaceae have a gynobasic style, most of this family (excluding Tiquilia) has a terminal style with a bifid stigma.

The Cordiaceae is another recent split, but these are also mostly trees and shrubs with characteristic 4-lobed stigmas and fleshy drupes as opposed to mostly herbs with 2-lobed stigmas and producing nutlets in the Boraginaceae.

Distribution of Boraginaceae

The Boraginaceae are a cosmopolitan family found around the tropics, warm and cool temperate, and even arctic climates, although they are predominantly found in warm temperate climates, with a strong Mediterranean concentration. In the Americas, the family is found in Canada, the USA, Mexico, Central America, and South America.

Distribution of Boraginaceae in the Americas

Canadian Genera Include:

Boraginoideae: Anchusa 3 spp. intro to all S provinces except NL; Borago 1 sp. intro to all S provinces but ephemeral in ON, QC, NS, and NL; Buglossoides 1 sp. intro to BC, MB, ON, QC, and NS; Cerinthe 1 sp. intro to QC?; Echium 2 spp. Intro to all of S Canada except Labrador; Lithospermum 8 spp. native to BC, AB, SK, MB, ON, and QC and intro NB; Nonea 2 spp. intro to AB and QC; Pulmonaria 2 spp. intro to ON, QC, and NS; Pentaglottis 1 sp. intro to BC; Symphytum 3 spp. introduced in all of S Canada except Labrador. Cynoglossoideae: Adelinia monospecific W NAM endemic native BC; Amsinckia 6 spp. native to BC and introduced to YT, AB, and SK; Andersonglossum 1 NAM endemic sp. native to all S provinces exc. Labrador, also in YT; Asperugo monospecific intro BC, AB, SK, and MB (and Greenland); Cryptantha 11 spp. native to BC, AB, and SK; Cynoglossum 1 sp. intro to all of S Canada exc. NL and PE; Eritrichium 6 spp. native to YT and NT; Hackelia 6 spp. native to all of Canada except NU, NS, PE, and NL; Lappula 2 spp. native to YT, NT, BC, AB, SK, MB, and introduced to ON, QC, NS, NB, PE, and NL; Mertensia 7 spp. native to all of Canada, including the Arctic (and Greenland); Myosotis 9 spp. native to BC, AB, YT, NT, ON, QC, NB, NS, PE, and NL and intro to SK and MB (and Greenland); Omphalodes 2 spp. intro in BC, ON, and QC; Oreocarya (or ~ Cryptantha) 1 NAM endemic native to BC, AB, and SK; Pectocarya 1 sp. native to BC; Plagiobothrys 5 spp. native to BC, AB, SK, and MB, and intro to YT, ON, QC, and NB.

USA Genera Include:

Boraginoideae: Anchusa 5 spp. intro much of USA exc. NV, AZ, NM, NE, KS, OK, MO, AR, LA, MS, AL, GA, FL, SC, IN, WV, and VT; Borago 1 sp. intro to WA, OR, CA, UT, MT, ND, MI, IL, MN, WI, TN, OH, VA, WV, MD, PA, NJ, NY, CT, RI, MA, VT, NH, and ME; Brunnera 1 sp. intro to MO, OH, and NY; Buglossoides 1 sp. intro to most of the USA, excluding NV, AZ, and NM; Echium 7 spp. intro to most of USA, including AK but excluding NV, AZ, ND, MS, AL, and FL; Lithospermum 17-21 spp. native and intro to all of the USA; Nonea 3 spp. intro to TN, PA, NY, and ME; Pulmonaria 1 sp. intro to NY, VT. Cynoglossoideae: Adelinia monospecific W NAM endemic native to WA, OR, and CA; Amsinckia 10 spp. native to W USA MT S to NM and all W, and intro to ND, NE, OK, TX, WI, IL, MS, OH, PA, NY, CT, MA, NH, ME, VA, NC, SC, AK, and HI; Andersonglossum  3 NAM endemic spp. native, inc. 1 endemic to OR and CA, 1 endemic to E USA from MN S to LA, plus ND, SD, OK, and TX, with a 3rd spp. native to SD, IA, WI, IL, IN, NJ, NY, NH, ME, and Canada; Asperugo monospecific intro to CA, CO, and IL; Bothriospermum 1 sp. intro HI; Cryptantha 82 spp. native in W USA from ND S to TX and all W, inc. MA, native and intro AK, most of which are endemic; Cynoglossum 9 spp. intro and native to the entire USA and intro to HI; Dasynotus monospecific narrow endemic of ID; Eritrichium 3 spp. native to AK, WA, OR, ID, WY, MT, CO, UT, and NM; Eremocarya 2 spp. S NAM endemic native OR, CA, NV, UT, AZ, NM, TX; Greeneocharis 2 spp. native to WA, OR, CA (1 endemic), ID, NV, AZ, UT, and CO; Hackelia 29 spp. native to almost all of the USA except FL, including AK; Harpagonella 1 S NAM endemic native to CA and AZ; Johnstonella ? spp. native to CA, NV, UT, AZ, NM, and TX; Lappula 4 spp. native and intro most of the USA, including AK, but excluding LA, MS, AL, FL, GA, and VA; Mertensia 18 spp. native to most of the USA, including AK, but exc. TX, OK, LA, and FL; Mimophytum 1 S NAM endemic sp. native to TX; Myosotis 11 spp. native and intro most of the USA, inc. AK, exc. ND, and intro in HI; Omphalodes 2 spp. native and intro OR and TX (USDA, POWO says otherwise); Oreocarya (or ~ Cryptantha) 32 NAM endemics native W USA from ND S to TX and all states W; Pectocarya 8 spp. native to WA, OR, ID, WY, CA, NV, UT, AZ, NM, and TX; Plagiobothrys 43 spp. native to WA, OR, CA (12+ endemics), ID, MT, WY, CO, NV, UT, AZ, NM, ND, SD, NE, WI, MI, IL, AR, LA, NC, PA, MA, and ME, and intro and native to AK; Symphytum 3 spp. introduced in WA, OR, CA, ID, MT, WY, UT, CO, NM, MI, WI, MN, MO, AR, TN, KY, GA, NC, VA, WV, IN, OH, PA, MD, DE, NJ, NY, CT, RI, MA, VT, NH, ME, and AK. Echinochiloideae: Antiphytum 2 spp. native to NM and TX.

Mexico Genera Include:

Boraginoideae: Lithospermum ~43 spp. native to all of Mexico, including ~32 endemics. Cynoglossoideae: Amsinckia 3 spp. native to N Mexico; Borago 1 sp. intro to SW+C Mexico; Cryptantha 5-6 spp. native to N Mexico, including 3+ endemics; Cynoglossum 1 sp. intro to NE+SW Mexico; Eremocarya 2 S NAM endemic spp. native to NW Mexico; Hackelia 5 spp. native to all of Mexico (1 endemic); Greeneocharis 1 sp. native to NW Mexico; Harpagonella 2 S NAM endemic spp. native to BC, BCS, Guadalupe Is., and Son; Johnstonella 17 spp. native to most of Mexico except Chp, Tab, Cam, Yuc, and QR, most are endemic to Mexico; Lappula 1-2 spp. native to N Mexico; Mertensia 1 sp. native to NE Mexico; Mimophytum 11 S NAM endemic spp. native to Coa, NL, Tam, Dgo, SLP, Pue, Qro, Hgo, Gto, Mex, Tlx, Ver, NE, C Mexico, Ver, including 10 endemics; Oncaglossum monospecific endemic in NE, C, and SW Mexico; Oreocarya 3-4? spp NAM endemics native throughout the N half of Mexico; Pectocarya 6 spp. native to NW Mexico, including 1-2 endemics of BC; Plagiobothrys 7+ spp. native to NW+C Mexico. Echinochiloideae: Amphibologyne monospecific endemic of NE and SW Mexico; Antiphytum ~10+ spp. native to much of Mexico exc. Chp, Cam, Tab, Yuc, QR, including several endemics.

Neotropical Genera Include:

Boraginoideae: Anchusa 1 sp. intro to Argentina; Borago 1 sp. intro to Guatemala, Honduras, Greater Antilles, Colombia, Ecuador, Peru, C Chile, Uruguay, and Argentina; Buglossoides 1 sp. intro to Bolivia, Argentina, and Uruguay; Echium 1 sp. intro to C Chile, Argentina, Uruguay, and S Brazil; Lithospermum ~7 spp. native to Guatemala (1) and the rest are endemic to the Andes of Colombia, Venezuela, Ecuador, Peru, and Bolivia; Moritzia 3 neoendemic spp. native to Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, and SE+S Brazil; Symphytum 1 sp. intro to Dominican Republic, Haiti, C Brazil, and NE Argentina; Thaumatocaryon  2 E SAM endemic spp. native to S+SE Brazil, Paraguay, and NE Argentina. Cynoglossoideae: Amsinckia 2-3 spp. originally endemic to W SAM in Ecuador (1 still endemic), Peru, Bolivia, Chile, and Argentina; Cryptantha ~50 spp. native and endemic to W SAM in Peru, Bolivia, N+C Chile, and Argentina; Cynoglossum 1 sp. intro Jamaica, Haiti, Dominican Republic; Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia, SE Brazil, Argentina, C+S Chile, and Juan Fernandez Is.; Greeneocharis 1 sp. native NW+S Argentina; Hackelia 7 spp. native to Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru (1 endemic), Bolivia, and N Argentina; Iberodes 1 sp. intro to C Chile; Johnstonella ~6 spp. native to Peru, N Chile, and NW Argentina; Lappula 1-3 spp. intro to Venezuela and native N+S Argentina; Myosotis 1-2 spp. intro to Guatemala, Dominican Republic, Haiti, Colombia, Ecuador, Peru, Bolivia, N+C Chile, N Argentina, SE Brazil, Uruguay, and Trinidad-Tobago; Nesocaryum monospecific endemic of the Desventurados Islands off the coast of NC Chile; Pectocarya 6 spp. native to Ecuador, Peru, Bolivia, N+C Chile, and Argentina; Plagiobothrys 8+ spp. native to Colombia, Ecuador, Peru, Bolivia, C+S Chile, and NW+S Argentina; Selkirkia 4 SAM endemics, inc. 1 endemic to Juan Fernandez Is., 2 endemic to C+S Chile, and 1 endemic to Colombia and Ecuador; Trichodesma 1 sp. intro to Windward Is. Echinochiloideae: Antiphytum 2-3 spp. endemic to S Brazil and Uruguay.

Patagonia Genera Include:

Boraginoideae: Anchusa 1 sp. intro to S Argentina; Borago 1 sp. intro to C Chile and S Argentina; Buglossoides 1 sp. intro to S Argentina; Echium 1 sp. intro to Patagonia region; Lithospermum 1 sp. intro S Argentina. Cynoglossoideae: Amsinckia 1-2 spp. native throughout Patagonia; Asperugo monospecific intro to Patagonia region; Cynoglossum 1 sp. intro throughout Patagonia; Greeneocharis  1 sp. native S Argentina; Lappula 1 sp. native to Patagonia region; Myosotis 1-2 spp. native to S Chile and S Argentina and intro to Falkland Is.; Pectocarya 1 sp. native to S Argentina and SC Chile; Plagiobothrys 1 sp. native to C+S Chile and S Argentina; Selkirkia 2 SAM endemics native to SC and S Chile.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, along with my own personal observations throughout North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • FNA (1993+). Flora of North America. https://floranorthamerica.org/Main_Page. Accessed 2022-current.
  • GBIF.org (2020+), GBIF Home Page. Available from: https://www.gbif.org
  • iNaturalist.org (2020+). https://www.inaturalist.org/. Accessed 2020-current.
  • Luebert, F., Cecchi, L., Frohlich, M. W., Gottschling, M., Guilliams, C. M., Hasenstab-Lehman, K. E., Hilger, H. H., Miller, J. S., Mittelbach, M., Nazaire, M., Nepi, M., Nocentini, D., Ober, D., Olmstead, R. G., Selvi, F., Simpson, M. G., Sutorý, K., Valdés, B., Walden, G. K., & Weigend, M. (2016). Familial Classification of the Boraginales. axon65(3), 502–522. https://plants.sdsu.edu/amsinckiinae/pdfs/Luebert_etal2016-Boraginales.pdf.
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019+). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Stevens, P. F. (2001+). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/. This is the authoritative source on currently accepted families, subfamilies, tribes, and genera.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; Accessed 2020-present.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Campanulaceae or Bellflower Family

Campanula alaskana the Alaskan Bellflower, showing the bell-shaped flower this family is named for
Campanula alaskana the Alaskan Bellflower, showing the bell-shaped flower this family is named for
Page Last Updated May 14, 2026.

Introduction to the Campanulaceae Family

The Campanulaceae, or Bellflower family, is part of the Asterales order (related to sunflowers) of the core dicots. It is a widespread family found on every continent except Antarctica, and it is a popular garden ornamental for its lovely flowers. The first time I found one was on a grassy bluff next to the ocean, and I automatically knew what family it was in because of the pretty blue bell-shaped flowers with the long style that the family is best known for, although other colors and flower shapes do exist in the family.

 Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description, with no need to know any scientific jargon. Below this section is additional information on uses and morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems: Most members are perennial herbs, but some annuals, biennials, shrubs, or small trees exist in the family; mostly they live on land, but some are aquatic. One key identifying feature is the milky juice that is released when they are damaged.

Leaves are usually arranged alternately on the stem but are sometimes in opposite pairs or whorls. They are usually simple (not compound), linear, lance-shaped, oblong, egg-shaped, or rounded with variously toothed margins but are occasionally divided into opposite pairs of leaflets.

Flowers: Flowers are often medium to large and showing, appearing singly or in various spikes or umbrella-shaped clusters. Flowers are typically bell-shaped or star-shaped, giving them the common name of bellflowers. Most are in shades of blue, but they can also be found in white, yellow, red, pink, or purple. Flowers are often symmetrical but can be irregular or “two-lipped,” like in Lobelia.

Reproductive Features: Reproductive features are specialized to ensure successful pollination by insects. There are usually 5 stamens (male parts) that may be separate or joined in a long column surrounding the female parts (ovary, style). Most species have an inferior ovary located below where the petals attach. A single long style comes up through the center of the flower.

Fruits: The fruit is almost always a dry capsule that splits open through valves, irregular slits, or other ways to release the seeds. Very rarely is the fruit a fleshy berry (mostly in the Lobelioideae subfamily). Some seeds are equipped with wings to aid in dispersal.

Uses of the Campanulaceae Family

With plenty of showy flowers, this family is popular for garden ornamentals, especially from Lobelia, Wahlenbergia, Codonopsis, Jasione, and more than 120 species of Campanula. They are widely grown in gardens and landscapes around the world.

Morphology of Campanulaceae in North America

Learn how to identify the Campanulaceae family with morphology photos
Learn how to identify the Campanulaceae family with morphology photos

Some Campanulaceae Species of North America

Campanuloideae Subfamily

Campanula alaskana the Alaskan Bellflower, showing the bell-shaped flower the Campanulaceae family is named for

Campanula alaskana—Alaska Bellflower

Herbaceous perennial from rhizomes with weak stems with larger basal leaves and small, linear stem leaves. Flowers are large and showy, bell-shaped, nodding, and up to 3 cm long in blue-violet to lavender colors. It is often confused with Campanula rotundifolia but has broader leaves, and it is only found along the North Pacific from Washington State north to Alaska.

Campanula rotundifolia flowers showing bell-shaped flowers with a 3-lobed stigma, both common features of the Campanulaceae.

Campanula petiolata—Western Harebell

This was a recent taxon split from the much more widespread Campanula rotundifolia, with which it shares many characteristics but tends to be more upright with stronger stems. The long style with a 3-lobed tip visible in the photo is common among Campanula rotundifolia and close relatives like these. Some taxonomists do not recognize this or Campanula alaskana as separate species from Campanula rotundifolia.

Campanulastrum americanum American Bellflower inflorescence with flowers showing a single long style that is characteristic of the Campanulaceae family.

Campanulastrum americanum—American Bellflower

A tall, erect annual or biennial common in moist woods. Unlike most bellflowers, it has flat star-shaped flowers in elongated spikes, and it has alternate lance-shaped leaves compared to the often linear ones seen in Campanula species.

Triodanis biflora plant with flowers in Georgia

Triodanis biflora—Venus’s Looking Glass

This species is an annual herb of disturbed areas with alternate leaves that do not wrap around the stem. It has pinkish to purple bellflowers with widely spreading lobes that make them look like a star. Native to the southern and eastern United States, Mexico, and South America.

Triodanis perfoliata perfoliate leaves and flowers closeup

Triodanis perfoliata—Clasping Venus’s Looking Glass

This species is very similar to Triodanis biflora, with a similar native range, though perhaps a bit more widespread. It can usually be differentiated by the leaves shown here that clasp and nearly wrap all the way around the stem. However, they can hybridize, making identification difficult.

Lobelioideae Subfamily

Lobelia appendiculata two-lipped flowers closeup; though it has 5 lobes, they are clearly divided into an upper lip and a lower lip, common among the Campanulaceae

Lobelia appendiculata—Pale Lobelia

This species is an annual unbranched herb that grows up to 60 cm tall with oblong to egg-shaped leaves that clasp partway around the stem. It has pale blue to white two-lipped flowers (2 upper lobes and 3 larger lower ones) in a loose terminal spike. It is native to the south-central USA.

Lobelia cardinalis, a plant with bright red showy flowers

Lobelia cardinalis—Cardinal Flower

This herbaceous perennial grows up to 1.2 m tall and is found mostly in or near bogs, riverbanks, swamps, or wet forests. It has large, lance-shaped to oval leaves with toothed margins. Flowers are large and bright red with 5 deeply cut lobes. Plants with pink or white flowers occasionally occur. It is native to southeastern Canada through the eastern and southern USA, south to northern Colombia.

Lobelia spicata inflorescence with flowers

Lobelia spicata—Pale Spike Lobelia

This short-lived perennial is often found flowering below taller grasses in sunny or semi-shaded prairies, woodlands, and disturbed areas. They have simple, variously elongated-shaped leaves with shallow teeth and are most known for their sometimes densely flowered spikes of white or pale blue flowers, like those in the photo. Native to southern Canada and the eastern USA.

Scientific Botanical Description of the Campanulaceae Family

Habit & Leaf Form of the Campanulaceae Family

Mostly perennial herbs with some annual or biennial and a few shrubs or small pachycaul trees with heights from 8 cm to over 2 m tall (excluding trees). They are laticiferous, which is an important identifying characteristic. Variously hydrophytic to xerophytic, and when hydrophytic, they are rooted with emergent and submerged leaves. Other plants may or may not have a basal aggregation of leaves. Leaves usually alternate or sometimes are opposite or whorled.

Leaves are petiolate or subsessile, sheathing or non-sheathing, and when sheathing, they have free margins. Leaves are not gland-dotted; they lack a pulvinus and stipules and are usually simple or sometimes compoundpinnate. Lamina, when simple, may be dissected or entire. When entire, it is linear, lanceolate, oblanceolate, oblong, ovate, obovate, or orbicular. When dissected, it is pinnatifid or palmatifid. Lamina margins are crenate, serrate, or dentate.

Flowers of the Campanulaceae Family

Plants are always hermaphrodites. Pollination is entomophilous and conspicuously specialized via modifications of the style with sterile tissue covering the stigmas. Flowers are solitary or aggregated in cymes, racemes, spikes, and umbels, sometimes pseudanthial. Inflorescences are scapiflorous or not, terminal or axillary, and with or without involucral bracts. Flowers are medium to large; regular to very irregular; 5-merous; tetracyclic. Perianth has a distinct calyx and corolla of 10 or 16–20 (Michauxia) parts in 2 whorls.

Calyx 5 or 8–10 (Michauxia); 1 whorled; free or connate (depending on interpretation, with the tube nearly always being united with the ovary); basally appendaged (e.g., Campanula with adjoining pairs of sepals contributing to each appendage); spurred or not; imbricate or valvate. An epicalyx is sometimes present. Corolla 5 or 8–10 (Michauxia); 1 whorled; connate (usually) or free (Jasione); valvate; often campanulate with long tubular bells or open starry ones; bilabiate or regular; mostly blue, but also white, yellow, red, pink, or purple; spurred (e.g., Heterotoma) or not spurred.

Androecium of the Campanulaceae Family

There are 5 or 8–10 (Michauxia) androecial members made of exclusively fertile stamens that are free of the perianth or adnate low down on the corolla and free of one another or coherent, 1-whorled, and sometimes forming a column around the style. Stamens are isomerous with the perianth, opposite the sepals, and alternating with the corolla members; they are filantherous or laminar and filantherous (e.g., being laminate below the filaments in Wahlenbergia). Filaments are sometimes basally appendiculate (sometimes in Campanula). Anthers are cohering and sometimes terminating an androecial column (e.g., Centropogon, Burmeistera) or separate, dehiscing via longitudinal slits, introrse, and tetrasporangiate.

Gynoecium of the Campanulaceae Family

The gynoecium is 2, 3, 5 (8 in Ostrowskia) or 8–10 carpelled (Michauxia). The pistil is 2 or 3 celled or occasionally 5 (6–10) celled. The gynoecium is synstylovarious and with a usually inferior ovary or rarely superior. The ovary is 2, 3, or 5 (6–10) locular. Styles 1. Stigmas equal in number to carpels; wet or dry type; papillate or non-papillate; Group II or IV type. Placentation axile with 10–50 ovules per locule; horizontal; non-arillate; anatropous; unitegmic; tenuinucellate.  

Fruit of the Campanulaceae Family

The fruit of the Campanulaceae is almost always a non-fleshy, dehiscent septicidal, loculicidal, valvular, or irregularly splitting capsule or, rarely, a fleshy indehiscent berry. Seeds are small, endospermic, oily, and rarely starchy and are sometimes winged. 

Taxonomy of Campanulaceae

There are 2,380 species in 84 genera of the Asterales order of the core Eudicots (dicots).

The family is divided into five subfamilies, 2 of which are very widespread:

  1. Campanuloideae is a large subfamily characterized by polysymmetric flowers, stamens that sprawl at the bottom of the corolla tube after the anthers have dehisced, an inferior ovary, and long-hairy styles. These are widespread and found worldwide, with especially high diversity in the north temperate Old World.
  2. Lobelioideae is a subfamily of herbs or small trees with terminal, occasionally axillary, inflorescences and large to small resupinate flowers. Stamen filaments are connate at least apically, and anthers are connate. They are mostly tropical, especially common in the New World with a major center of diversity in the Andes and over 100 endemic species found on the Hawaiian Islands. They are not present in the Arctic and are absent from the Near East and Central Asia.
  3. Nemacladoideae is a small subfamily of tiny annuals (rarely perennial) with sub-opposite leaves, racemes without bracteoles, and small flowers that are not resupinate. Anthers are connivent; filaments are connate apically and may be free at the base. These are restricted to the southwestern USA (especially California) and northwestern Mexico.
  4. Cyphioideae is a group of perennial herbs (twining vines) and shrubs with tuberous roots. The fused corolla is split almost to the base into two groups, usually with three upper lobes and two lower lobes. Stamen filaments may be free or connate. Mostly in Southern Africa but also in East Africa and the Cape Verde Islands.
  5. Cyphocarpoideae is a small subfamily endemic to Chile and is made of annual or perennial spiny herbs with deeply lobed leaf margins and foliaceous bracts. The upper corolla lobe is sub-hooded, and the lower lobes show three ridges.

Genera of the Campanulaceae:

Campanuloideae: Adenophora (68), Asyneuma (37), Azorina (1? or syn. Campanula), Berenice (1), Campanula (452), Canarina (3), Codonopsis (49), Craterocapsa (5), Cryptocodon (1), Cyananthus (19), Cyclocodon (3), Cylindrocarpa (1), Echinocodon (1), Edraianthus (21), Favratia (1), Feeria (1), Githopsis (4), Gunillaea (2), Hanabusaya (1), Heterochaenia (4), Heterocodon (1), Himalacodon (1), Homocodon (2), Jasione (14), Kericodon (1), Legousia (6), Merciera (6), Michauxia (8), Microcodon (4), Muehlbergella (1), Musschia (3), Namacodon (1), Nesocodon (1), Ostrowskia (1), Pankycodon (1), Peracarpa (1), Petromarula (1), Physoplexis (1), Phyteuma (22), Platycodon (1), Prismatocarpus (27), Pseudocodon (8), Rhigiophyllum (1), Roella (22), Sachokiella (1? or syn. Campanula), Sergia (2), Siphocodon (2), Theilera (2), Theodorovia (1? or syn. Campanula), Trachelium (2), Treichelia (2), Triodanis (6), Wahlenbergia (263), Zeugandra (2).

Lobelioideae: Apetahia (? or syn. Sclerotheca), Brighamia (2), Burmeistera (129), Centropogon (212), Clermontia (23), Cyanea (81), Delissea (15), Dialypetalum (5), Diastatea (9), Dielsantha (1), Downingia (13), Grammatotheca (1), Heterotoma (1), Hippobroma (1), Howellia (1), Isotoma (10), Legenere (1), Lobelia (444), Lysipomia (32), Monopsis (15), Palmerella (1), Porterella (1), Ruthiella (4), Sclerotheca (10), Siphocampylus (237), Solenopsis (13), Trematolobelia (8), Unigenes (1), Wimmeranthus (1), Wimmerella (10).

Nemacladoideae: Nemacladus (26), Parishella (? syn. Nemacladus), Pseudonemacladus (1).

Cyphioideae: Cyphia (76).

Cyphocarpoideae: Cyphocarpus (4).

Key Differences From Similar Families

Campanulaceae is easy to distinguish by the combined presence of latex, simple leaves, and an inferior ovary. Sometimes plants from Acanthaceae, Lamiaceae, and Rubiaceae with long reddish or orange corolla tubes are placed in Campanulaceae, but they differ in leaves that are usually opposite and having free stamens and anthers.

Distribution of the Campanulaceae

The Campanulaceae are found from frigid zones to tropical climates with a cosmopolitan distribution excluding tropical Africa and Antarctica. Found from deserts to rainforests and the Arctic, but the majority of species are by far northern temperate species.

In the Americas, the Campanulaceae are found through Canada, including the Arctic (and Greenland), and south through the USA, Mexico, Central America, and South America.

Distribution of Campanulaceae in the Americas

NOTE: This data was gathered 4-5 years ago and may not reflect current genus names and status.

Canadian Campanulaceae Genera Include:

Campanuloideae: Campanula 19 spp., including 18 native in all of Canada, including the Arctic (and Greenland), and 1 sp. intro to BC; Githopsis 1 W NAM endemic sp. native to S Vancouver Island, BC; Heterocodon monospecific W NAM endemic native to BC; Jasione 1 sp. intro to BC?; Triodanis 1 sp. native to BC, ON, and QC. Lobelioideae: Downingia 2 spp. native to BC, AB, and SK; Isotoma 1 sp. intro to BC; Lobelia 2 spp. native to BC, ON, and QC, and ephemeral NB.

USA Campanulaceae Genera Include:

Campanuloideae: Asyneuma 1 sp. native to OR and CA; Campanula 34 spp., including 33 native to most of the USA and AK but excluding OK, AR, LA, MS, and 1 sp. intro to NV and NH; Clermontia 22-23 narrow endemic spp. of HI; Cyanea 80-81 spp. narrow endemic genera of HI; Githopsis 4 spp. W NAM endemic genera native MT, WA, OR, and CA (2 endemics); Heterocodon monospecific W NAM endemic native to WA, OR, CA, NV, ID, MT, WY, and CO; Jasione 1 sp. intro to WA, OR, NC, MD, PA, DE, NY, CT, NJ, RI, and MA; Legousia 1 sp. intro to CA and PA; Platycodon 1 sp. intro to NY, PA, NC, and HI; Triodanis 6 spp. native to all of the USA (5 endemics), and intro HI; Wahlenbergia 2 spp. intro to TX, LA, AR, MS, AL, GA, SC, NC, and FL. Lobelioideae: Brighamia 2 spp. narrow endemic genera of HI; Delissea 15 spp. narrow endemic genera of HI; Downingia 13 spp. native to WA, ID, MT, OR, WY, CA, NV, and UT, includes 9 USA endemics, mostly in CA; Hippobroma monospecific intro to FL and HI; Howellia monospecific W USA endemic of WA, ID, MT, OR, and CA; Legenere monospecific Americas disjunct endemic of CA + Patagonia; Lobelia 43 spp. native and intro to the entire USA and HI and native in AK; Palmerella monospecific narrow endemic of S CA + N BC Mexico; Porterella monospecific endemic of OR, ID, WY, CA, UT, and AZ; Trematolobelia 8 spp. narrow endemic genera of HI. Nemacladoideae: Nemacladus 18-26 W NAM endemic spp. native to OR, ID, CA, NV, UT, AZ, and NM, including many endemic to W USA.

Mexico Campanulaceae Genera Include:

Campanuloideae: Campanula 1-2 spp. native to NE Mexico and intro to SW+SE Mexico and Ver; Githopsis 1 W NAM endemic sp. native BC and Guadalupe Is.; Heterocodon monospecific W NAM endemic native to NW Mexico; Triodanis 1 sp. native throughout Mexico except the SE. Lobelioideae: Centropogon 4+ spp. mostly neoendemic, native to most of Mexico except the NW, and includes 1 endemic species; Diastatea 8 spp. including 7 endemics native to most of Mexico (exc. NW); Downingia 1 sp. native to NW Mexico; Heterotoma monospecific neoendemic native to NE+C+SW+NE Mexico; Hippobroma monospecific intro to all of Mexico; Lobelia ~86 spp., including ~20 endemic to all of Mexico; Palmerella monospecific narrow endemic of S CA + N BC Mexico; Wimmeranthus monospecific endemic of SW Mexico. Nemacladoideae: Nemacladus 10 W NAM endemic spp. native to NW Mexico, mostly in BC, including 1 endemic to BC; Pseudonemacladus monospecific endemic to NE Mexico.

Neotropical Campanulaceae Genera Include:

Campanuloideae: Triodanis 1 sp. native to Guatemala, Colombia, Ecuador, Peru, Bolivia, Paraguay, S+SE Brazil, Uruguay, and Argentina, and intro to Jamaica and Dominican Republic; Wahlenbergia 6 spp. neoendemics of E+S Brazil (inc. 1 Brazil endemic), Colombia, French Guiana, Peru, Ecuador, Bolivia, Paraguay, Argentina, C Chile, and Uruguay. Cyphocarpoideae: Cyphocarpus 3 spp. narrow endemic genera of N+C Chile. Lobelioideae: Burmeistera 129 spp. neoendemic genera of Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, and Peru; Centropogon ~211 spp. mostly neoendemic native to CAM (exc. Belize) and tropical SAM S to Peru, Bolivia, and C+SE Brazil, and the Lesser Antilles (2 endemics); Diastatea 2 spp. native to CAM (exc. Belize), Colombia, Venezuela, Ecuador, Peru, Bolivia, and NW Argentina; Heterotoma monospecific Mexico and neoendemic native to Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica; Hippobroma longiflora monospecific originally endemic Jamaica but widely introduced elsewhere; Lobelia ~50?? spp. native to CAM, the Antilles, all of SAM except Guyana and including Galapagos; Lysipomia 32 spp. Andes endemic genera of Venezuela, Colombia, Ecuador, Peru, and Bolivia; Siphocampylus 237 spp. neoendemic genera of Costa Rica, Panama, the Greater Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil (exc. N), Paraguay, N Argentina, and Uruguay.

Patagonia Campanulaceae Genera Include:

Campanuloideae: Campanula 1 sp. intro to S Argentina and Falkland Is.; Triodanis 1 sp. native to S Argentina; Wahlenbergia 1 sp. native to S Chile. Lobelioideae: Downingia 1 sp. native throughout Patagonia (also native in California); Legenere monospecific Americas disjunct endemic of California + Patagonia; Lobelia 5+ spp. native throughout Patagonia and Falkland Is. and intro to South Georgia Is.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, along with my own personal observations throughout North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • FNA (1993+). Flora of North America. https://floranorthamerica.org/Main_Page. Accessed 2022-current.
  • GBIF.org (2020+), GBIF Home Page. Available from: https://www.gbif.org
  • iNaturalist.org (2020+). https://www.inaturalist.org/. Accessed 2020-current.
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019+). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Stevens, P. F. (2001+). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; Accessed 2020-present.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].



Blog


How to Identify the Apiaceae / Carrot Family

Daucus carota inflorescence with flowers.
Daucus carota inflorescence with flowers. Flat-topped umbels of white flowers are quite common among the Apiaceae.
Page Last Updated May 13, 2026.

Introduction to the Apiaceae Family

The Apiaceae family is widespread globally but is particularly common in northern temperate zones. It is well known for its many edible foods and herbs, including carrots, celery, parsley, dill, and many, many more. Most plants have a characteristic smell and possess small flowers in umbels, making the family fairly easy to spot in the field.

The Apiaceae are part of the Apiales order of core dicot flowering plants. The very closely related Araliaceae family has many overlapping characteristics. In general, though, Apiaceae are usually herbs that produce dry fruits that split into 2 segments when mature (schizocarps), while the Araliaceae are usually shrubs or trees and usually produce berry-like drupes (soft fruits with a pit inside, like a cherry).  

Common Botanical Description

If you’re new to plant morphology, this guide is a great beginner’s description for learning to identify the Apiaceae family, with no need to know any scientific jargon. For researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below.

Leaves and Stems of the Apiaceae: Most plants are aromatic (malodorous to pleasant) annual, biennial, or perennial herbs. Their stems are often hollow between the nodes and are typically grooved or ribbed like celery and are often covered with hairs. The leaves are arranged alternately on the stem and often have sheaths or wrap around the stem. The leaves are often compound, made of many smaller leaflets.

Flowers of the Apiaceae: Flowers are usually rather small but are grouped into often large, flat-topped clusters called umbels for how they resemble the ribs of an umbrella. Each flower typically has five free white or yellow petals, or occasionally pink or purple. Flowers in the center of the cluster are often symmetrical, while those on the outer edge often have larger outer petals and smaller inner petals.

Reproductive Features of the Apiaceae: Flowers are bisexual, with both female (ovary, style, and stigma) and male (stamens) parts in the same flower. There are 5 free (unjoined) stamens. There is a single inferior ovary (located below where the petals attach), with two styles (pollen-collecting tubes) on top that often have thickened bases.

Fruits of the Apiaceae: Fruits are not fleshy but dry “schizocarps” that split into two parts, each containing a single seed when mature.

Uses and Cautions of Apiaceae 

Many members of the Apiaceae family are used for culinary purposes, including Daucus (carrot), Pastinaca (parsnip), Apium (celery), Petroselinum (parsley), Pimpinella (anise), Carum (caraway), Anethum (dill), Anthriscus (chervil), Foeniculum (fennel), and Levisticum (lovage), to name some of the more common ones. Ornamental ones include Eryngium, Angelica, Heracleum, Trachymene, and others; however, many ornamentals have been modified with enlarged bracts or sepals and are not representative of the family.

Many Apiaceae are also used medicinally in folk, herbal, and Chinese medicines to treat a wide variety of ailments.

Still other Apiaceae members have notoriously poisonous resins or alkaloids such as Cicuta, Conium (hemlocks), Aethusa (fool’s parsley), and others are phototoxic (Heracleum mantegazzianum). 

Morphology of Apiaceae in North America

Learn how to identify the Apiaceae family with common and scientific botanical descriptions and morphology photos.

Some of the Apiaceae Species Found in North America

Apioideae Subfamily

Anthriscus sylvestris inflorescence with flowers, a very typical inflorescence of the Apiaceae family

Anthriscus sylvestris—Cow Parsley

Herbaceous perennial, 60 – 170 cm tall with hollow, grooved stems that are green with purple splotches and tiny hairs. Leaves are compound and made of many leaflets (2-3 times pinnate) and appear fern-like. Flowers are white, small, in compound umbels like shown in the photo, with individual umbelets having hairy oval little bracts with red tips. It is native to Eurasia and Africa but widely introduced in the Americas.

Bifora americana inflorescence

Bifora Americana—Prairie Bishop

Herbaceous annual to 80 cm tall with deeply divided leaves with threadlike leaflets and tiny white flowers in compact compound umbels with individual petals having notched tips that help in their identification. Fruits are small, roundish dry fruits that split apart at maturity (schizocarp). This species is endemic to the southern USA, in Texas, Oklahoma, and Arkansas. This one was in Texas.

Chaerophyllum procumbens leaf made of numerous leaflets and flowers; compound leaves are common of the Apiaceae family

Chaerophyllum procumbens – Spreading Chervil

Herbaceous annual, 15 – 50 cm tall, with slender, branching, shiny, and finely hairy stems. Mostly hairless leaves are divided into many leaflets and look very fern-like (bipinnately compound). Flowers are in small terminal compound umbels, each with 1 to 7 small white flowers that bloom before the umbel finishes expanding, as shown in the photo. This Apiaceae species is endemic to eastern North America.

Chaerophyllum tainturieri inflorescence with flowers and fruits

Chaerophyllum tainturieri – Hairyfruit Chervil

Herbaceous annual to 80 cm tall with erect stems, opposite compound leaves (bipinnately like many Apiaceae) that are stiff-hairy. Flowers are in few-flowered umbels (not compound) in leaf axils, and it produces oblong fruits (schizocarps) also visible in the photo. It is native to the southeastern USA, with disjunct populations in Arizona and New Mexico.

Cicuta douglasii - deadly poisonous Apiaceae plant with flowers

Cicuta douglasii – Western Water Hemlock

Herbaceous semi-aquatic perennial from thick tuberous roots. Inner tubers and stem bases have horizontal chambers used to help identify them. Leaves are alternately arranged and divided (pinnate) into 3 – 10 cm leaflets with jagged margins. The flowers are arranged in compound umbels with numerous small white flowers. This Apiaceae is the most poisonous native North American plant. It is endemic to the Pacific Northwest, mostly in BC, Canada, where this one was.

Cicuta maculata - deadly poisonous plant with flowers

Cicuta maculata – Spotted Cowbane

Herbaceous perennial from rhizomes with a hollow, erect stem up to 1.8 m tall. It has compound leaves with several lance-shaped, pointy-tipped, toothed leaflets 2 – 10 cm long. Flowers are in compound umbels with many white or whitish flowers. The whole plant is poisonous and is native from northern Canada to southern Mexico.

Conium maculatum- deadly poisonous plant with flowers, inflorescence is also a common Apiaceae feature.

Conium maculatum—Poison Hemlock

Herbaceous biennial 1.5 – 2.5 m tall with hairless, smooth, green, hollow stems usually streaked with red or purple. Leaves are finely divided and lacy-looking (2 – 4 times pinnately compound) and about 50 cm long. Flowers are small and white in loose compound umbels. Native to Eurasia and Africa, widely introduced and invasive in North America.

Cymopterus longiradiatus plant with flowers. Narrow endemic of New Mexico and Texas.

Cymopterus longiradius – Trans-Pecos False Mountainparsley

Herbaceous perennial from a taproot with a basal rosette of 3 times pinnately compound leaves. Flowers are in compound umbels in yellow. The fruit is a schizocarp. This Apiaceae member is narrow native endemic of the Sacramento and Guadalupe Mountains of New Mexico and Texas, USA.

Daucus carota inflorescence with flowers, very tipical of the Apiaceae.

Daucus carota—Wild Carrot

Herbaceous biennial, 30–120 cm tall with a roughly hairy, stiff, solid stem. Finely divided leaves (tripinnate) are 5 – 15 cm long and alternately arranged. Flowers are small and whitish in large, dense, flat-topped terminal umbels, often with a central pink or purple flower visible here. Dry fruits (schizocarps) develop as the umbels contract and become congested. It is native throughout the temperate Old World and has been widely introduced in the Americas.

Daucus pusillus native American wild carrot inflorescence with fruits

Daucus pusillus—American Wild Carrot

Slender, herbaceous annual to 60 cm tall with hairy stems and finely dissected fern-like leaves with umbels of white flowers (lacking a central purple flower as in D. carota), with feathery bracts that extend beyond the flowers. The photo shows the contracted umbel in fruit. Native throughout the temperate Old World and widely introduced in the Americas.

Heracleum mantegazzianum invasive herbaceous plant with flowers

Heracleum mantegazzianum – Giant Cow Parsnip

Herbaceous perennial 2 – 5 m tall with a hollow, rigid, erect green stem with prominent red-purple splotches and coarse white hairs. Leaves are about 1 m wide, deeply lobed, with shredded (incised) margins. White or green-white flowers in compound umbels are flat-topped and massive, up to 1 m across. This Eurasian native is widely introduced and invasive. It causes phototoxicity and contact dermatitis.

Heracleum maximum herbaceous plant with flowers

Heracleum maximum—Cow Parsnip

Herbaceous perennial plant to 3 m tall with hollow, densely hairy stems that have few to no purple splotches. Leaves are 3-lobed and may be deeply so but not shredded (incised) on the margins and are up to 40 cm across. White flowers in compound umbels up to 30 cm across may be flat-topped or rounded, with outer flowers often much larger than inner flowers. Native to Canada & most of the USA, plus East Asia.

Lomatium grayi plant with flowers

Lomatium grayi—Gray’s Lomatium

Herbaceous perennial with divided, fernlike (pinnately compound) dark-green leaves. Compound umbels each with hundreds of yellow flowers appear on leafless stalks. Hairless fruits are elliptic with lateral wings. The plant smells of parsley. It is endemic to the western United States, and this one was found on Antelope Island, Utah.

Lomatium foeniculaceum plant with flowers

Lomatium foeniculaceum – Desert Biscuitroot

An herbaceous perennial herb up to 30 cm tall from a taproot. It’s stemless with hairy leaf stalks and peduncles that grow from the ground. Leaves are compound and fern-like (2–4x pinnate) and up to 30 cm long. Flowers are small, yellow, or purplish in compound umbels. The plant smells and tastes of parsley. It is native to western and central North America. This photo was taken at Gloss Mountain, OK, USA.

Lomatium multifidum inflorescence with flowers

Lomatium multifidum—Fern-leaved Desert Parsley

Herbaceous perennial 30 – 140 cm tall that may be hairless, hairy, or sandpapery to the touch, with a thin dry sheath at the base. Leaves are compound and fernlike (3x pinnate), 15–35 cm wide on a 3–30 cm petiole. Small yellowish flowers appear in compound umbels with tiny bracts. The dry fruit (schizocarp) has thick wings. This Apiaceae is native to western North America from BC, Canada, south to Baja California, Mexico.

Oenanthe sarmentosa aka Water Parsley inflorescence with flowers, which is very typical of the Apiaceae.

Oenanthe sarmentosa—Water Parsley

This perennial herb grows up to 1.5 m tall with compound leaves (pinnately divided but not as fern-like as others) up to 30 cm long on long stalks up to 35 cm long. Individual leaflets are toothed and lobed. Flowers are in somewhat dense compound umbels with white to red-tinged petals. It is native to western North America from Alaska south to California.

Osmorhiza berteroi plant with flowers

Osmorhiza berteroi—Mountain Sweet Cicely

Fragrant herbaceous perennial with branching stems to 1m tall. Leaves are divided into 3 leaflets with toothed or lobed margins and are up to 20 cm long with a long leaf stalk. Small whitish flowers appear in small terminal compound umbels with 4 – 10 flowers in each umbellet, with central flowers often having only anthers. The dry fruit (schizocarp) is elongated, ribbed, and bristly and up to 2.5 cm long. It is native to temperate regions of both North and South America.

Osmorhiza longistylis Aniseroot plant with flowers.

Osmorhiza longistylus—Aniseroot

This lovely herbaceous perennial grows to about 80 cm tall with small compound umbels made of small white flowers. It is easy to identify when you crush the leaves, as they smell of anise. This Apiaceae member is native to both North and South America.

Pastinaca sativa plant with umbels of fruits

Pastinaca sativa – Wild Parsnip

Herbaceous biennial/perennial to 1.2 m tall that is easy to identify because it smells like parsnip. Leaves are alternately arranged and compound (pinnate), made of yellow-green, diamond-shaped, and coarsely toothed leaflets. Numerous yellow flowers appear in compound umbels. The dry fruit (schizocarp) is made of 2 flattened, slightly winged sections (mericarps). It may cause photosensitivity. This native of Eurasia is widely introduced and invasive.

Scandix pecten-veneris plant with flowers and needle-like fruits, which, though they look different, they are schizocarps which are typical of the Apiaceae

Scandix pecten-veneris – Shepherd’s Needle

This small herbaceous annual only grows up to 50 cm tall with compound leaves that are finely divided and fern-like. It produces small white flowers in few-rayed umbels. What it is most known for is its distinctive needle-like fruits that give it its common name. This species is native to Eurasia and is introduced throughout the USA and parts of South America.

Sium suave plant with flowers, showing odd-pinnate compound leaves common in the Apiaceae.

Sium suave—Water Parsnip

Herbaceous perennial to 3 m tall with light green, hairless stems up to 5 cm thick with few branches. Leaves appear in basal rosettes or in clusters in aquatic habitats; those on the stem are alternately arranged and compound with opposite pairs and a single terminal leaflet (odd-pinnate). The flowers are in small compound umbels with 10 – 20 small white flowers. This Eurasian native is introduced in North America.

Torilis arvensis inflorescence with flowers, another common Apiaceae style inflorescence.

Torilis arvensis – Spreading Hedge Parsley

Herbaceous annual with slender, branching, rough-hairy stems to 1 m tall. Leaves are alternate and compound (pinnate) with deeply divided or coarsely toothed lance-shaped leaflets up to 6 cm long. Open compound umbels have few flowers per umbellet that have 5 unequal-sized petals in white with pink or red tinges. Fruits are greenish to pinkish, 3 – 5 mm long, and coated with straight or curved prickles. This species is native to Europe and has been introduced in North America.

Vesper bulbosus inflorescence with flowers and fruits forming

Vesper bulbosus – Bulbous Spring-Parsley

This herbaceous spring annual grows from a large swollen taproot and has compound (2x pinnate) leaves with pale gray-green leaflets. It produces purple to pink or whitish flowers in compact umbels follwed by distinctive fruits with conspicuous tan to purplish wings. This plant is endemic to the western United States.

Saniculoideae Subfamily

Eryngium yuccifolium inflorescence with flowers; while the umbels are common in the Apiaceae, its condensed flower heads are not.

Eryngium yuccifolium—Rattlesnake Master

Herbaceous perennial to 1.8 m tall with 15 – 100 cm long, stiff, 1 – 3 cm wide, sharp-tipped, waxy blue-green leaves with spiny margins. Flowers are in dense, round, terminal flowerhead-like umbels made of 10 – 40 tiny condensed flowers with a faint honey scent and a spiny green bract beneath them, with another cluster of spiny bracts beneath the round umbel, just visible in the photo. It is native to the tallgrass prairies of the central and eastern USA.

Sanicula canadensis leaves, stem, and flowers

Sanicula canadensis—Canadian Blacksnakeroot

Herbaceous biennial or perennial 0.3 – 1.4 m tall with compound leaves made of 3 (trifoliate) egg-shaped or elliptic leaflets. Small pale green flowers in tiny bur-like compound umbels have sepals that are longer than their petals. These are followed by bur-like dry fruits (schizocarps) that split into two parts (mericarps). It is native to eastern North America, west to Wyoming, USA.

Scientific Botanical Description of the Apiaceae

Below is for citizen scientists, researchers, or anyone who wants to dive deeper into the world of botany to learn more about the Apiaceae family. Note that most botanical terms have a hover-over definition to help you better understand what they mean.

Habit & Leaf Form of the Apiaceae

Plants of the Apiaceae are usually soft-stemmed aromatic annual, biennial, or perennial herbs, but some have tough stems, and a few are even woody, tree-like, or shrubby species in the tropics. Stems are hollow between leaf joints and are often ribbed (e.g., Angelica, celery). Sometimes they are switch-plants, occasionally with the principal photosynthesizing function transferred to stems (e.g., Platysace compressa) or phyllodinous (petiole or rachis performs leaf function) (e.g., Lilaeopsis). Leaves are usually well-developed but can be reduced in switch-plants. Plants are only occasionally succulent (e.g., Crithmum) and may or may not have a basal aggregation of leaves. They can be helophytic, mesophytic, or xerophytic (e.g., Eryngium). Plants occasionally are conspicuously heterophyllous (e.g., Apium inundatum).

Leaves are small to large and arranged alternately or alternately and oppositely (usually just oppositely on upper leaves). Leaves are usually herbaceous but may occasionally be leathery or rarely fleshy. Attachment to the stem is either petiolate or sometimes perfoliate or peltate and always more or less sheathing. Sheaths have free margins and may or may not be pulvinate. Leaves may be gland-dotted and are usually malodorous to pleasant-smelling, but are almost always scented. Leaf arrangement may be simple or compound ternate, imparipinnate, bipinnate, multiply compound, or rarely palmate. When simple, the lamina is usually dissected pinnatifid, but it may also be palmatifid (as seen in Sanicula, Astrantia, and Eryngium), spinose (in Eryngium), or sometimes entire. They are pinnately, palmately, or parallel-veined. Leaves are usually without stipules (but sometimes with stipular flanges).

Flowers of the Apiaceae

Apiaceae plants are usually hermaphroditic but may also be andromonoecious, polygamomonoecious, or dioecious (Acronema). Pollination is entomophilous. Numerous small flowers are almost always in a characteristic flat-topped terminal cymose umbel(s), often compound. Sometimes they are in cymose heads or are rarely reduced to a single flower. Inflorescences may be pseudanthial, often with sterile flowers at the periphery, and may or may not have involucral bracts. Flowers are usually actinomorphic but may be zygomorphic at the umbel edge.

Flowers are usually fairly small, possess bracts, and may be regular to slightly irregular in the corolla. Flowers are usually perfectly 5-merous (except for the gynoecium) and are tetracyclic. Free hypanthium is not present. The perianth usually has a distinct calyx and corolla (but the calyx is often very reduced). The perianth may be sepaline (corolla usually present) or petaline (calyx teeth sometimes present) with 4–10 parts in two (or one) isomerous whorls. The petals are unequal in size, with those pointing outwards from the umbel larger than those pointing in. The calyx, when detectable, has five parts in one whorl that may be free or connate (often reduced to a rim but never forming a tube). Sepals are persistent and may be lobulate, blunt-lobed, or toothed. The corolla also has five parts in one whorl. The white, yellow, pink, or purple petals are always free, valvate, and may be unequal or regular. 

Androecium of the Apiaceae

The androecium has five members (all fertile stamens) that are free of the perianth and each other. They are all equal or unequal and are found in 1 whorl. Their five stamens are isomerous with the perianth, opposite sepalous, and inflexed in bud. Anthers are dorsifixed or basifixed, introrse, tetrasporangiate, and dehisce via longitudinal slits.

Gynoecium of the Apiaceae

The gynoecium is 2-carpeled, and the pistil is one- or two-celled. The gynoecium is synovarious, median, and inferior. The ovary is 2(1) locular. An epigynous disk is present. Two apical styles are free to partly joined, with their bases thickened into one or two stylopodia that crown the ovary. Stigmas are wet type, non-papillate, and Group IV type. Placentation is either axile or apical. There are 1 or 2 ovules per locule (usually 2 with 1 abortive) that are pendulous and either epitropous or anatropous. Ovules have a ventral raphe, are non-arillate, are unitegmic, and are tenuinucellate or pseudocrassinucellate.  

Fruit of the Apiaceae

The fruits of the Apiaceae are non-fleshy, dry schizocarps with 2 mericarps that are united facially. The mericarps are 1-seeded in each part, with the integument sometimes united with the pericarp. Seeds are oily and endospermic.

Taxonomy of Apiaceae

There are 3,820 species in 446 genera in the Apiaceae family of the Apiales order of the core eudicots. Then, the Apiaceae family is further divided into four main subfamilies, with a couple of unplaced genera as well.

  1. Apioideae – The largest subfamily and a diverse group of annual to perennial herbs or subshrubs or rarely a small tree. Leaves are usually palmately divided but may be pinnate or simple and entire. Venation is pinnate, but some have parallel venation. Fruits are variable and may or may not be flattened, winged, or ribbed. Cosmopolitan distribution. 
  2. Azorelloideae – Annual herbs or small shrubs, often hummock-forming. Leaves are simple, trifid to palmately lobed, and stipulate. Fruit with wings or ribs where the wings or ribs are the largest. Mostly South American, particularly the Patagonia region.
  3. Mackinlayoideae – Annual herbs to shrubs. Leaves may or may not be pedately compound, palmate to simple, and are stipulate. Fruit is drupaceous and often laterally compressed. Mostly Old World Tropics.
  4. Saniculoideae – A diverse group of annual herbs and subshrubs to trees. Leaves may be tripinnately or palmately compound or simple and often palmately lobed, and some have hairy or spiny teeth. Inflorescences are usually simple umbels or capitula, and bracts may be foliaceous or petaloid. Fruits may be barely to strongly compressed dorsally or laterally, may have 2-3 wings, or may be scaly or spiny. Cosmopolitan distribution.    

Note that Angelica (Apioideae member) looks like it is being split so that the North American species may now/soon be considered Archangelica, while many Eurasian species have been moved to other groups as the genus gets reduced by ~100 species, so I give the species numbers in a range, and the distribution data for the USA and Canada below still call it Angelica until I can get enough sources in agreement.

Genera of the Apiaceae:

Apioideae: Aciphylla (45), Acronema (35), Actinanthus (1), Actinolema (2), Adenosciadium (1), Aegokeras (3), Aegopodium (12), Aethusa (1), Afroligusticum (14), Afrosciadium (18 Not in APG), Agasyllis (1), Alepidea (31), Aletes (4), Ammi (3), Ammodaucus (1?), Ammoides (2), Ammoselinum (3), Andriana (3 Not in APG), Anethum (2), Angelica (2-106), Anginon (12), Angoseseli (1), Anisopoda (1), Anisosciadium (3), Anisotome (17), Annesorhiza (23), Anthriscus (14), Aphanopleura (3), Apiastrum (1), Apium (12), Apodicarpum (1), Arafoe (1), Arctopus (3), Arcuatopterus (5), Arracacia (42), Artedia (1), Asciadium (1), Astomaea (2), Astrantia (11 Not in APG), Astrodaucus (3), Astydamia (1?), Athamanta (12?), Aulacospermum (15?), Austropeucedanum (1?), Autumnalia (2?), Azilia (1?), Berula (7), Bifora (3), Billburttia (2), Bilacunaria (6), Bonannia (1), Bunium (34), Bupleurum (224), Cachrys (7), Calyptrosciadium (2), Cannaboides (2 not in APG), Capnophyllum (4), Carlesia (1), Caropodium (3 Not in APG), Caropsis (1), Carum (20), Caucalis (1), Cenolophium (1), Cephalopodum (2), Chaerophyllopsis (1), Chaerophyllum (69), Chaetosciadium (1), Chamaesciadium (1), Chamaesium (10), Chamarea (5), Changium (2), Choritaenia (1), Chuanminshen (1), Chymsydia (2), Cicuta (4), Cnidiocarpa (2), Cnidium (8), Coaxana (2), Conioselinum (21), Conium (6), Conopodium (8), Coriandrum (2), Cortia (3), Cortiella (5), Cotopaxia (2), Coulterophytum (4), Crithmum (1), Cryptotaenia (6), Cuminum (4), Cyathoselinum (1), Cyclorhiza (4), Cyclospermum (3), Cymbocarpum (5), Cymopterus (42), Cynosciadium (1), Dactylaea (2), Dahliaphyllum (1 Not in APG), Dasispermum (7), Daucosma (1), Daucus (45), Demavendia (1), Dethawia (1), Deverra (10? or syn. of Deverra), Dichoropetalum (36 or syn. of Dichosciadium), Dichosciadium (1), Dickinsia (1), Dicyclophora (1), Dimorphosciadium (2), Diplaspis (3), Diplolophium (7), Diplotaenia (5), Donnellsmithia (19), Dracosciadium (2), Drusa (1), Ducrosia (6), Dystaenia (2), Echinophora (11), Ekimia (4), Elaeosticta (25), Eleutherospermum (1), Elwendia (31 not in APG), Enantiophylla (1), Endressia (2), Eremodaucus (1), Ergocarpon (1), Erigenia (1), Eurytaenia (2), Exoacantha (1), Ezosciadium (1 not in APG), Falcaria (1),Fergania (1), Ferula (230), Ferulago (1), Ferulopsis (1), Foeniculum (5), Frommia (1), Froriepia (1), Fuernrohria (1), Galagania (6), Geocaryum (3),Gingidia (12), Glaucosciadium (2), Glehnia (2), Glia (3), Gongylosciadium (1), Gongylotaxis (1 not in APG), Grafia (1), Grammosciadium (4), Halosciastrum (1), Haloselinum (1 not in APG), Hansenia (9 not in APG), Haplosciadium (1), Harbouria (1), Harperella (1? or syn. Ptilimnium),  Harrysmithia (2), Haussknechtia (1), Hellenocarum (3), Helosciadium (6? or syn. Apium), Heptaptera (8), Heracleum (92), Heteromorpha (8), Hladnikia (1), Hohenackeria (2), Homalocarpus (6), Homalosciadium (1?), Horstrissea (1), Hyalolaena (9), Hymenidium (40 not in APG), Hymenolaena (3), Itasina (1), Johrenia (7), Kadenia (2), Kafirnigania (1), Kailashia (3 not in APG), Kalakia (1), Kandaharia (1), Karatavia (1), Karnataka (1), Kedarnatha (6), Kelussia (1 not in APG), Kenopleurum (1 not in APG), Keraymonia (4), Kitagawia (8), Klotzschia (3), Komarovia (1),  Komarovia (?), Korshinskia (5), Kozlovia (2), Krasnovia (1), Krubera (1), Kundmannia (1), Ladyginia (3), Lagoecia (1), Lalldhwojia (4), Laser (7), Laserpitium (8), Lecokia (1), Ledebouriella (1), Lefebvrea (10), Leiotulus (10 not in APG), Lereschia (1), Leutea (9), Levisticum (1), Lichtensteinia (7), Lignocarpa (2), Ligusticopsis (21), Ligusticum (35), Lilaeopsis (12), Limnosciadium (2), Lipskya (1), Lisaea (3), Lithosciadium (2 not in APG), Lomatium (106), Lomatocarpa (4), Lomatocarum (1 not in APG), Magadania (2), Magydaris (2), Marlothiella (1), Mastigosciadium (1), Mathiasella (1), Mediasia (1), Meeboldia (6), Melanosciadium (6), Meum (1), Modesciadium (1?), Mogoltavia (2?), Molopospermum (1?), Musineon (6), Mutellina (4?), Myrrhidendron (5), Myrrhis (1), Nanobubon (3?), Naufraga (1?), Neoconopodium (2), Neogaya (1?), Neogoezia (5), Neomuretia (2?), Neonelsonia (1), Neoparrya (1), Niphogeton (18), Nirarathamnos (1), Normantha (1), Nothosmyrnium (2), Notiosciadium (1), Notobubon (13?), Oedibasis (4), Oenanthe (35), Oligocladus (1), Oliveria (1), Opoidia (1 or syn. Peucedanum), Opopanax (4), Oreocome (10), Oreocomopsis (3?), Oreonana (3), Oreoschimperella (3), Oreoxis (2-4), Orlaya (3), Ormopterum (2), Ormosciadium (1), Osmorhiza (12), Ostericum (11 or syn. Angelica), Ottoa (1), Oxypolis (4), Pachypleurum (2), Palimbia (3), Paraligusticum (1), Parapimpinella (1), Paraselinum (1), Parasilaus (2), Pastinaca (16), Pastinacopsis (1), Paulita (3), Pedinopetalum (1), Perideridia (14), Perissocoeleum (4), Petroedmondia (1), Petroselinum (1), Peucedanum (72), Phellolophium (2), Phlojodicarpus (3), Phlyctidocarpa (1), Physospermopsis (8), Physospermum (2), Physotrichia (6), Pilopleura (2), Pimpinella (156), Pinda (2), Pleurospermopsis (2), Pleurospermum (9), Podistera (4), Polemannia (3), Polemanniopsis (2), Polytaenia (3), Polyzygus (1), Portenschlagiella (1), Postiella (1), Prangos (49), Prionosciadium (23), Psammogeton (19), Pseudocannaboides (1?), Pseudocarum (2), Pseudoridolfia (1?), Pseudoselinum (1), Pseudotrachydium (5), Pternopetalum (21), Pterygopleurum (1), Ptilimnium (5), Ptychotis (2), Pycnocycla (15), Pyramidoptera (1), Registaniella (1), Rhabdosciadium (8), Rhizomatophora (1?), Rhodosciadium (15), Rhopalosciadium (1), Rhysopterus (1?), Ridolfia (1), Rivasmartinezia (2), Rohmooa (1), Rupiphila (1), Rutheopsis (2), Sajanella (1), Saposhnikovia (1), Scaligeria (6), Scandia (2), Scandix (12), Scaraboides (1?), Schoenoselinum (1?), Schrenkia (13), Schtschurowskia (2), Schulzia (5), Sclerochorton (1), Sclerosciadium (1?), Sclerotiaria (1), Scrithacola (1), Selinopsis (2), Selinum (11), Semenovia (31), Seseli (141), Seselopsis (2), Shoshonea (1), Siculosciadium (1?), Silaum (1?), Siler (4), Sillaphyton (1?), Silphiodaucus (2), Sinocarum (18), Sinolimprichtia (1), Sison (3), Sium (10), Sivadasania (1?), Smyrniopsis (2), Smyrnium (6), Spananthe (2), Spermolepis (11), Sphaenolobium (3), Sphaerosciadium (1), Sphallerocarpus (1), Spiroceratium (1?), Spuriopimpinella (5), Stefanoffia (2), Steganotaenia (3), Stenocoelium (4), Stenosemis (2?), Stenotaenia (5?), Stewartiella (1), Stoibrax (3), Symphyoloma (1), Synclinostyles (2), Szovitsia (1), Taenidia (2), Taeniopetalum (3?), Tamamschjanella (2?), Tamamschjania (1), Tana (1?), Tauschia (34), Tetrataenium (21), Thamnosciadium (1), Thapsia (8), Thaspium (4), Thecocarpus (2), Tiedemannia (2), Tilingia (3), Todaroa (1), Tongoloa (18), Tordyliopsis (1), Tordylium (20), Torilis (15), Trachydium (11), Trachyspermum (15), Trepocarpus (1), Tricholaser (2), Trigonosciadium (7), Trinia (13), Trocdaris (1?), Trochiscanthes (1), Tschulaktavia (1?), Turgenia (2), Turgeniopsis (1?), Vanasushava (1), Vesper (6?), Vicatia (5), Villarrealia (1?), Vinogradovia (1?), Visnaga (2?), Vvedenskya (1), Xanthogalum (3? or syn. Angelica), Xatardia (1), Xyloselinum (3?), Yabea (1), Zeravschania (14), Zizia (3), Zosima (4).

Azorelloideae: Asteriscium (9), Azorella (58), Bolax (2), Bowlesia (16), Diposis (3), Domeykoa (5), Eremocharis (9), Gymnophyton (6), Microsciadium (1?), Oschatzia (2), Pozoa (2), Schizeilema (?), Stilbocarpa (?).

Mackinlayoideae: Actinotus (20), Apiopetalum (2), Brachyscias (1), Centella (55), Chlaenosciadium (1), Mackinlaya (5), Micropleura (2), Pentapeltis (2), Schoenolaena (1), Xanthosia (20).

Saniculoideae: Eryngium (254), Hacquetia (?), Petagnaea or Petagnia (1?), Sanicula (49).

Unplaced: Hermas (9), Platysace (23).

Key Differences From Similar Families

Apiaceae are both similar to and closely related to the Araliaceae Family, and often, the two can be hard to differentiate. However, some more common differences include the fact that the Apiaceae are usually herbs vs. usually shrubs and trees in the Araliaceae; the leaves that usually lack stipules in the Apiaceae vs. usually present in the Araliaceae; and the fruit is a dry schizocarp with two mericarps vs a berry-like drupe in the Araliaceae.

Distribution of Apiaceae

The Apiaceae family is a cosmopolitan family found all over the globe from frigid to tropical zones. Still, most members of the Apiaceae are in the north temperate zone.

Distribution of Apiaceae in the Americas

Canadian Apiaceae Genera Include:

Apioideae: Aegopodium 1 sp. intro to all of S Canada except AB and Labrador; Aethusa monospecific intro to ON, QC, NB, and NS; Anethum 1 sp. intro to S provinces except NS, NB, PE, and NL; Angelica 9 spp. native to all of Canada (& Greenland) except SK and MB; Anthriscus 3 spp. intro to BC, ON, QC, NB, NS, and NL (exc. Labrador); Astrantia 1 sp. intro to NL (exc. Labrador); Berula 1 sp. native to BC and ON; Bunium 1 sp. intro NL?; Bupleurum 2 spp. native to YT, AB, NT, BC, and intro to QC and ON; Carum 1 sp. intro to all of Canada except YT and Labrador; Chaerophyllum 3 spp. intro to BC, native in ON, and QC?; Cicuta 4 N temperate spp. native to all of Canada including the Arctic; Cnidium 1 sp. native to BC, YT, and NT; Conioselinum 2 N temperate spp. native to BC, ON, NL, NB, NS, and PE; Conium 1 sp. intro to BC, AB, SK, ON, QC, NB, and NS; Conopodium 1 sp. intro to NL Island; Coriandrum 1 sp. intro to ON, QC, and NS; Cryptotaenia 1 sp. a former NAM endemic native in MB, ON, and QC and extirpated in NB; Cymopterus 1 NAM endemic sp. native to AB, SK, and MB; Daucus 2 spp. including 1 native to BC and 1 intro to all of the S provinces; Erigenia monospecific E NAM endemic sp. native to ON; Foeniculum 1 sp. intro to BC, ON, and QC via cultivation; Glehnia 1 NW NAM endemic sp. native to BC; Heracleum 4 spp., inc. 1 native to all of Canada except NU and 3 spp. intro to BC, YT, ON, QC, NB, NS, PE, and NL; Levisticum monospecific intro to AB, ON, QC, and NS; Ligusticum 6 N temperate spp. native to BC, ON, QC, NL, NS, NB, PE, and NU (& Greenland); Lilaeopsis 2 spp. native in BC and NS; Lomatium 20 W+C NAM endemic spp. native to BC, AB, SK, and MB; Musineon 1 C NAM endemic sp. native to AB, SK, and MB; Myrrhis monospecific intro to BC, ON, NS, and NL Island; Oenanthe 1 sp. native to BC; Osmorhiza 6 spp. native to all of Canada; Oxypolis 2 spp. NAM endemic genus native to BC and ON; Pastinaca 1 sp. intro to all of Canada except NU; Perideridia 1 NAM endemic sp. native to BC, AB, and SK; Petroselinum monospecific intro to BC and ON; Peucedanum 1 sp. intro to ON and NS; Pimpinella 2 spp. intro to AB, ON, QC, NB, NS, NL (exc Labrador); Podistera 2 spp. native to YT and NT; Sium 1 sp. native to all of Canada including the Arctic; Taenidia 1 E NAM endemic sp. native to ON and QC; Thaspium 3 spp. E NAM endemic genera native to ON and NS; Torilis 3 spp. intro to BC, ON, and QC; Yabea monospecific W NAM endemic species native BC; Zizia 2 N NAM endemic spp. native to all of S Canada exc. PE and NL, and inc. YT.

Saniculoideae: Eryngium 4 subcosmopolitan spp. introduced to BC, AB, SK, ON, and QC; Sanicula 9 spp. native to all of S Canada except Labrador.  

USA Apiaceae Genera Include:

Apioideae: Aegopodium 1 sp. intro to WA, OR, ID, MT, MN, WI, IL, MO, KY, TN, GA, SC, NC, VA, WV, MD, DE, IN, OH, MI, PA, NJ, NY, CT, RI, MA, VT, NH, and ME; Aethusa monospecific sp. intro to ID, AR, MN, WI, IL, IN, OH, KY, MI, WV, PA, DE, NJ, NY, CT, RI, MA, and ME; Aletes 4 S NAM endemic spp. native to CO, NM, TX, and WY, including 2 narrow endemics of CO (1 of which was also in WY but has been extirpated); Ammi 2 spp. intro to OR, CA, AZ, TX, LA, MS, AL, GA, FL, SC, NC, MO, SD, and PA; Ammoselinum 2-3 spp., including 1 endemic, native to CA, AZ, NM, TX, LA, OK, KS, AR, MO, MS, TN, NC, and intro to AL; Anethum 1 sp. intro to most of the USA, including HI but excluding ID, NV, UT, WY, NM, TN, MS, AL, GA, SC, FL, VT, and NH; Angelica 22 spp. native to all of the USA, including AK, but excluding ND, SD, NE, KS, and TX; Anthriscus 3 spp. intro to WA, OR, CA, ID, MT, WY, AZ, NE, OK, MN, MO, AR, LA, GA, SC, NC, TN, VA, WV, MD, DE, WI, IL, IN, OH, MI, PA, NJ, NY, CT, RI, MA, VT, and ME; Apiastrum monospecific SW NAM endemic native to CA and AZ; Apium 4 spp. intro in WA, OR, CA, ID, UT, AZ, TX, OK, SD, MO, LA, IL, TN, MS, FL, SC, NC, OH, WV, PA, NJ, NY, CT, and MA; Berula 1 sp. native to all of the W USA from MN S to LA and all states W except LA and including MI and NY; Bifora 3 spp., including 1 S USA endemic of TX, OK, and AR, and 2 spp. intro to AL, PA, MD, NJ, and RI; Bupleurum 4 spp. native AK and native and intro to most of USA except WA, NV, UT, ND, NE, OK, MN, IA, WI, RI, ME, MS, and SC; Carum 1 sp. intro to the N half of USA from OR E to NJ and all N except NE and including UT, CO, NM, LA, MO, TN, KY, WV, VA, MD, and NC; Caucalis monospecific intro to PA and HI; Chaerophyllum 4 spp. intro and native to most of the E USA from ND S to TX and all E except ND, SD, MN, CT, MA, VT, NH, and ME, and including AZ and NM; Cicuta 4 N temperate spp. native to all of the USA, Inc. AK; Cnidium 2 spp. including 1 native in AK and 1 intro to OR; Conioselinum 4 N temperate spp. native to most of the USA, including AK, but excluding ID, NV, ND, SD, KS, OK, TX, AR, MS, AL, SC, FL, TN, KY, WV, MD, and DE; Conium 1 sp. intro to all of the USA except MS and FL; Coriandrum 1 sp. intro to WA, OR, CA, NV, MT, ND, SD, AZ, NM, TX, OK, LA, FL, MO, IL, TN, SC, NC, VA, DE, MD, MI, OH, PA, NJ, NY, CT, RI, MA, and HI; Cryptotaenia 1 former NAM endemic sp. native to all of the E USA from ND S to TX and every state E; Cryptotaenia 1 former NAM endemic sp. intro to Colombia and Peru; Cuminum 1 sp. intro to TX and MA; Cyclospermum 1 former Mexico & neoendemic sp. intro to all of the S USA from NM E to NC and inc CA, NV, OR, IL, VA, WV, MD, PA, NY, and HI; Cymopterus ~40 spp. native to all of the W half of USA from ND S to TX, including MN; Cynosciadium monospecific S USA endemic of TX, OK, LA, AR, MO, IL, TN, MS, and AL; Daucosma monospecific narrow endemic of NM and TX; Daucus 2 spp., including 1 native to the W, S, and SE USA and 1 intro to all of the USA, including AK and HI; Erigenia monospecific E NAM endemic sp. native to NE, OK, MO, AR, and WI S to MS and all E except FL, SC, DE, NJ, CT, RI, MA, VT, NH, and ME; Eurytaenia 2 S USA endemic spp. of NM, TX (1 endemic), and OK; Falcaria monospecific intro to WY, SD, NE, OK, LA, IA, MO, WI, IL, WV, VA, MD, PA, NY, and CT; Foeniculum 1 sp. intro to HI and most of the USA except ID, MT, WY, CO, ND, SD, OK, MN, AL, VT, and NH; Glehnia 1 NW NAM endemic sp. native to WA, OR, and CA; Harbouria monospecific C USA endemic of WY, CO, and NM; Harperella monospecific SE USA endemic of AL, AK, MS, GA, SC, NC, VA, MD, and OK; Helosciadium 1 sp. intro to CA, PA, and NJ; Heracleum 3 spp. inc. 1 native and 2 intro to most of the USA exc TX, OK, AR, LA, MS, AL, FL, and SC, and it is native in AK; Levisticum monospecific intro to CO, NM, MO, MN, MI, OH, PA, NJ, NY, CT, MA, VT, and ME; Ligusticum 11 N temperate spp. native to all W and E states, including AK but excluding ND, S to TX, and MN, IA, LA, FL, WI, IL, MI, and VT; Lilaeopsis 6 spp. native to WA, OR, CA, AZ, TX, LA, AR, MS, AL, FL, GA, SC, VA, NC, MD, NY, NJ, MA, NH, ME, and AK; Limnosciadium 2 spp. SE USA endemic genus of TX, OK, KS, LA, AR, MS, MO, and IA; Lomatium 89+ W+C NAM endemic spp. native to all of the W+C USA from MN S to LA and all states W exc. LA, with most being endemics of the USA; Musineon 6 C NAM endemic spp. native to ID, MT (1 narrow endemic), WY, CO, NV, UT, NM, ND, SD, and NE, including 5 endemic to the USA; Myrrhis monospecific introduced to OR, MI, and PA; Neoparrya monospecific narrow SW USA endemic of NM and CO; Oenanthe 4 spp. native to AK, WA, OR, and CA and intro to MO, OH, MD; Oreonana 3 narrow endemic spp. of CA; Oreoxis 2-4 (USDA says 4) spp. narrow endemic genus of UT, CO, and NM; Osmorhiza 8 spp. native to all of the USA including AK but exc. FL and LA, including 1 US endemic; Oxypolis 4 spp. NAM endemic genera native to all of the USA except WA, ID, WY, NV, ND, SD, NE, and KS, inc. 2 endemic to S USA; Pastinaca 1 sp. intro in all the USA exc. MS, AL, GA, and FL; Perideridia 14 NAM endemic spp. native to W USA from MT S to NM, plus SD, KS, OK, MO, AR, IL, IN, KY, TN, AL, and MS and extinct in OH, includes 12 endemic to the USA; Petroselinum monospecific intro to WA, CA, NV, UT, ID, MT, KS, TX, LA, AR, MS, FL, GA, SC, NC, IA, MI, OH, PA, MD, NJ, NY, CT, RI, MA, and HI; Peucedanum 3 spp. intro to TN, PA, NY, WV, and MA, and 1 native to HI; Pimpinella 3 spp. intro to WA, MT, MN, WI, TN, IN, MI, OH, VA, WV, MD, DE, PA, NJ, NY, CT, MA, VT, and ME; Podistera 4 spp. native to CA, UT, CO, NM, and AK; Polytaenia 3 spp. E USA endemic genus of ND, NE, KS, OK, TX, MN, IA, MO, AR, LA, MS, AL, TN, KY, WI, IL, IN, and MI; Ptilimnium 5 USA & Caribbean endemic spp. native in most of SE USA (4 endemics) from KS S to TX and E to NJ except IN, OH, and inc SD, NY, CT, RI, and MA; Scandix 1 sp. intro to WA, OR, CA, AZ, TX, SD, AL, GA, FL, SC, NC, TN, OH, MI, PA, MD, NJ, NY, RI, and MA; Selinum 1 sp. intro to MA, VT, and NY?; Seseli 1 sp. intro to MD; Shoshonea monospecific narrow endemic of WY and MT; Sium 2 spp. native to all of the USA exc. OK but inc. AK; Smyrnium 1 sp. intro in AL; Spermolepis 9 spp. native in most of S+E USA (6 endemics) from ND S to TX except the NE states but inc. NM, AZ, CA, and HI; Taenidia 2 spp. E NAM endemic genus native in all the E half of USA from ND S to TX and all E except ME, NH, CT, FL, ND, and NE, includes 1 endemic of NE USA; Tauschia 10 spp. native to WA, ID, OR, CA, and TX, about half are endemic; Thaspium 4 spp. E NAM endemic genera native to all of E USA (1 endemic) from ND S to TX and all E exc ND, SD, NE, CT, MA, VT, NH, ME; Tiedemannia 2 SE USA & Caribbean endemic spp. native to TX, LA, MS, AL, GA, FL, SC, NC, and MD, extirpated in DE; Tilingia 1 sp. native to AK; Tordylium 1 sp. intro to AZ; Torilis 5 spp. intro to most of the USA except MT S to NM, also except NV, ND, SD, MN, DE, CT, RI, VT, NH, ME, and inc. HI; Trachyspermum 1 sp. intro to MI; Trepocarpus monospecific SE USA endemic of TX, OK, LA, AR, MO, KY, TN, MS, AL, GA, FL, and SC; Turgenia 1 sp. intro to WA, OR, and PA; Vesper 6 SW NAM endemic spp. native to CA, NV, ID, UT, AZ, WY, CO, NM, SD, NE, KS, OK, and TX, including 5 SW USA endemics; Visnaga 1 sp. intro to CA, OR?, AL, TX?, FL?, NC?, and PA?; Yabea monospecific W NAM endemic sp. native to WA, ID, OR, CA, NV, UT, AZ, and NM; Zizia 3 N NAM endemic spp. native in all of the USA exc. CA, AZ, and NM, includes 1 endemic to SE USA.

Azorelloideae: Bowlesia 1 sp. native to OR, CA, NV, AZ, NM, TX, OK, LA, MS, AL, and FL. Mackinlayoideae: Centella 2 spp. native to TX, LA, AR, MS, AL, GA, FL, SC, NC, VA, MD, DE, and NJ and intro to WA, OH, and HI. Saniculoideae: Eryngium 34 subcosmopolitan spp. native and intro to all of the USA except UT, WY, MT, and ND; Sanicula 22 spp. native in all of the USA except AZ and UT, and inc. HI.

Mexico Apiaceae Genera Include:

Apioideae: Aletes 1 S NAM endemic sp. native to NW Mexico; Ammi 1 sp. intro to SW+C Mexico; Ammoselinum 1-2 spp. native to BCN, BCS, Son, Sin, Chi, Dgo, Zac, Coa, NL, Tam, SLP, Gto, Qro, Ags, and Hgo; Angelica 2 spp. native to most of Mexico except BCN, BCS, Son, Sin, Chp, Tab, Cam, Yuc, and QR; Apiastrum monospecific SW NAM sp. native to BC, BCS, Son, and Sin?; Apium 1 sp. intro to NW Mexico; Arracacia ~24 Mexico & neoendemic spp. native through all of Mexico, including several endemics; Berula 1 sp. native to Nay, Jal, Col, Mch, Gro, Oax, Chp, Pue, Cam, Yuc, and QR;Brachyscias 1 sp. intro to NE Argentina; Chaerophyllum 3 spp. native to N+SW+C Mexico and Ver, including one endemic; Cicuta 2 N temperate spp. native to Chi, Coa, NL, Tam, Dgo, Zac, Gto, Qro, Ags, SLP, and Hgo; Coaxana 2 Mexico + CAM endemic spp. native to Nay, Jal, Col, Mch, Gro, Oax, Chp, Cam, Tab, Yuc, and QR, including 1 endemic of Gro + Oax; Conioselinum  1 N temperate sp. native to BC, BCS, Son, Sin, Chi, Coa, NL, Tam, Dgo, Zac, Ags, Gto, Qro, Hgo, SLP, Nay, Jal, Col, Mch, Gro, and Oax; Coriandrum 1 sp. intro to NW+C Mexico; Coulterophytum 4 narrow Mexico endemic spp. if Mch (1), Jal (2), Dgo + Sin + Nay (1); Cuminum 1 sp. intro to Chi, Coa, NL, Tam, Dgo, and Zac; Cyclospermum 1 former Mexico & neoendemic sp. native most of Mexico except BC, BCS, Son, and Sin where it is intro; Cymopterus 2-5 NAM endemic spp. native to much of Mexico except Mex, Cd Mex, Mor, Tlx, Pue, Ver, Tab, Cam, Yuc, and QR; Dahliaphyllum monospecific endemic of Nay, Jal, Col, Mch, Gro, and Oax; Daucus 3 spp., including 2 native to all of Mexico, including the Mexican Pacific Is., and 1 sp. intro to NE+C Mexico; Donnellsmithia 19 Mexico + N neoendemic spp. native through all of Mexico, including 17 endemic to Mexico; Enantiophylla monospecific Mexico & CAM endemic native to most of Mexico except Ver, Tab, Cam, Yuc, and QR; Ligusticum 1-2 N temperate spp. native to BC, BCS, Son, Sin, Chi, Coa, NL, Tam, Dgo, Zac, Ags, SLP, Hgo, Gto, and Qro; Lilaeopsis 1 sp. native to most of Mexico except Chp, Tab, Cam, Yuc, and QR; Lomatium 1-2 W+C NAM endemic spp. native to BC, BCS, Son, and Sin; Mathiasella monospecific NE Mexico endemic of Chi, Coa, NL, Tam, Dgo, SLP, Zac, Gto, Ags, Qro, and Hgo; Myrrhidendron 1 N neoendemic sp. native to Chp, Tab, Cam, Yuc, and QR; Neogoezia 5 spp. Mexican endemic genus of N+SW+C Mexico, and Ver; Neonelsonia monospecific neoendemic native to Nay, Jal, Col, Mch, Gro, Oax, Chp, Tab, Cam, Yuc, and QR; Oenanthe 1 sp. native to Nay, Jal, Col, Mch, and Gro; Osmorhiza 3 spp. native to most of Mexico except the SE, inc. 1 narrow endemic of NL; Ottoa monospecific N neoendemic sp. native in Nay, Jal, Col, Mch, Gro, Chp, Ver, Tab, Cam, Yuc, and QR; Petroselinum monospecific intro to Mex, Cd Mex, Mor, and Pue; Prionosciadium 23 Mexico & Guatemalan endemic spp. native to all of Mexico, including 21+ endemic to Mexico; Rhodosciadium 15 Mexico & Guatemala endemic spp. native to all of Mexico (14 endemics); Spananthe 1 Mexico & neoendemic sp. native in all of Mexico; Spermolepis 4 spp. native to BC, BCS, Son, Sin, Chi, Coa, NL, Tam, Dgo, Zac, Ags, Hgo, SLP, and Ver; Tauschia ~22 spp. native, mostly endemics, in all of Mexico; Torilis 1-2 spp. intro to NE Mexico; Vesper 1 SW NAM endemic sp. native to BCN and Son; Villarrealia monospecific narrow endemic of Coa and NL; Visnaga 1 sp. intro to SW Mexico; Yabea monospecific W NAM endemic native to NW Mexico.

Azorelloideae: Bowlesia 2 spp. native to N, C, and SW Mexico. Mackinlayoideae: Centella 2 spp. mostly native to Chi, Coa, Dgo, NL, Tam, Zac, Gto, Qro, Ags, SLP, and Hgo, but 1 sp. is intro; Micropleura 1 Mexico & N neoendemic sp. native to all of Mexico. Saniculoideae: Eryngium ~55 spp. native throughout all of Mexico; Sanicula 4-5 spp. native to all of Mexico, including 2 edemics. 

Neotropical Apiaceae Genera Include:

Apioideae: Ammi 2 spp. intro to Bermuda, Bahamas, Cuba, Hispaniola, Guatemala, El Salvador, Colombia, Argentina, S Brazil, and Uruguay; Ammoselinum 1 narrow endemic sp. of NE Argentina + Uruguay; Anethum 1 sp. intro to Guatemala, Bahamas, Greater Antilles, Leeward & Windward Is, Ecuador, Peru, NE Argentina, and S+SE Brazil; Anthriscus 1-2 spp. intro to C Chile and NW Argentina; Apium 9 spp., including 8 native to Bolivia, Chile (1 endemic), Juan Fernandez Is. (2 endemic), Desventurados Is, Argentina, Paraguay, S+SE Brazil, and Uruguay (1 endemic) and 1 intro to Guatemala, Bermuda, Cuba, Hispaniola, Puerto Rico, Trinidad-Tobago, Ecuador, and Peru; Arracacia ~35 Mexico & neoendemic spp. of Guatemala, El Salvador, Honduras, Costa Rica, Panama, Andes of Colombia, Venezuela, Ecuador, Peru, and Bolivia, and intro to Cuba, Hispaniola, Jamaica, Puerto Rico, and Trinidad-Tobago; Asciadium monospecific endemic of Cuba; Austropeucedanum monospecific endemic of NW Argentina; Berula 1sp. native to Guatemala; Bupleurum 1 sp. intro to NE Argentina; Caucalis monospecific intro to NE Argentina; Chaerophyllum 3 spp. native to Guatemala, Costa Rica, Colombia, Ecuador, Peru, Bolivia, and Argentina, including 2 endemics of the Andes; Coaxana 1 Mexico & CAM endemic sp. native to Guatemala and Honduras; Conium 1 sp. intro to Guatemala, Honduras, El Salvador, Cayman Is, Hispaniola, Colombia, Ecuador, Peru, Bolivia, S+SE Brazil, Argentina, and C+S Chile; Coriandrum 1 sp. cultivated and naturalized in Cuba, Hispaniola, Puerto RIco, Trinidad-Tobago, Juan Fernandez Is., Ecuador, Peru, Paraguay, S Brazil, and Argentina; Cotopaxia 2 N SAM endemic spp. of the high Andes of Ecuador and Colombia; Cyclospermum 3 spp. native to CAM (except Belize), Colombia, Venezuela, Ecuador, Galapagos, Peru, Bolivia, C+E+S Brazil, Paraguay, Chile, Argentina, and Uruguay, and intro to Bermuda, Greater Antilles (exc. Cayman Is.), Leeward & Windward Is., and Juan Fernandez Is., including 2 endemics of C-S SAM; Daucus 3 spp. including 2 native to Guatemala, El Salvador, Honduras, Colombia, Venezuela, Ecuador, Peru, Bolivia, S+SE Brazil, Chile, Argentina, and Uruguay and 1 sp. intro to Greater Antilles (exc. Cayman Is.), Leeward Is, Trinidad-Tobago, Juan Fernandez Is, Desventurados Is., Peru, Argentina, Chile, Uruguay, and S Brazil; Donnellsmithia 2 Mexico & N neoendemic spp. native to Guatemala, El Salvador, Honduras, Nicaragua, Panama, Colombia, and Venezuela; Enantiophylla monospecific Mexico & CAM endemic native to Guatemala, El Salvador, and Honduras; Foeniculum 1 sp. cultivated and naturalized in Bermuda, the Bahamas, Greater Antilles (exc. Cayman Is.), Leeward & Windward Is, Guatemala, El Salvador, Colombia, Ecuador, Peru, Chile, Paraguay, Argentina, S+SE Brazil, and Uruguay; Helosciadium 1 sp. intro to N+C Chile and NE Argentina; Heracleum 1 sp. intro to Haiti; Homalocarpus 6 narrow endemics of N+C Chile; Klotzschia 3 spp. endemic genus of Brazil; Lilaeopsis 6 spp. native to Cuba, Dominican Republic, Colombia, Ecuador, Peru, Bolivia, Paraguay, S Brazil (1 narrow endemic), Chile, Argentina, and Uruguay; Myrrhidendron 5 N neoendemic spp. native to Guatemala, Honduras, Costa Rica, Panama (1 narrow endemic), and N Andes of Colombia (1 narrow endemic) and Ecuador, including 4 endemic to neotropical zone; Neonelsonia monospecific neoendemic native to Guatemala and Andes of SAM in Colombia, Venezuela, Ecuador, and Peru; Niphogeton 18 neoendemic spp. of Costa Rica, Panama, and N Andes of Colombia, Venezuela, Ecuador, Peru, and Bolivia; Notiosciadium monospecific narrow E SAM endemic of NE Argentina and Uruguay; Oenanthe 1 sp. intro to Uruguay and NE Argentina; Oligocladus monospecific Argentina endemic found throughout Argentina; Osmorhiza 4 spp. native to Guatemala, Colombia, Peru, Bolivia, Argentina, and C+S Chile; Ottoa monospecific N neoendemic native to Guatemala, Costa Rica, Colombia, Venezuela, and Ecuador; Paraselinum monospecific endemic of Peru and Bolivia; Pastinaca 1 sp. intro to Greater Antilles, Ecuador, Peru, Bolivia, Argentina, C+S Chile, and Uruguay; Pedinopetalum monospecific endemic of the Dominican Republic; Perissocoeleum  4 spp. narrow endemics of Colombia & Venezuela; Petroselinum monospecific intro to Guatemala, El Salvador, Hispaniola, Puerto Rico, Ecuador, Peru, S Brazil, and Argentina; Pimpinella 2? spp. intro to Guatemala, Venezuela, N Argentina, and S Brazil; Prionosciadium 1 Mexico & Guatemala endemic sp. native in Guatemala; Ptilimnium 1 USA & Caribbean endemic sp. native to Cuba, Hispaniola, and Puerto Rico; Rhodosciadium 1 Mexico & Guatemala endemic sp. native to Guatemala; Ridolfia monospecific intro to Peru; Scandix 1 sp. intro to C Chile and NE Argentina; Seseli 1 sp. intro to C Chile; Sium 1 sp. intro to N+C Chile; Spananthe 2 spp. Mexico & neoendemic genera native CAM, Colombia, Venezuela, Ecuador, Peru (1 endemic), Bolivia, C +E Brazil, Haiti, and Trinidad-Tobago; Spermolepis 1 sp. endemic throughout Argentina; Tauschia 7-10 spp. native to Guatemala (2 endemics?), Colombia, Venezuela, and Ecuador; Tiedemannia 2 S USA & Caribbean endemic spp. native to the Bahamas and Cuba; Tordylium 1 sp. intro to NE Argentina; Torilis ~2 spp. intro to Peru, Bolivia, C Chile, N Argentina, S Brazil, and Uruguay; Visnaga 1 sp. intro to Colombia, Peru, C Chile, N Argentina, Uruguay, and S Brazil.

Azorelloideae of the Neotropics: Asteriscium 9 S SAM endemic spp. of Chile (3 endemics) and Argentina (3 endemics in NW); Azorella ~40 spp. native to Costa Rica, Venezuela, Colombia, and through the Andes S through Ecuador, Peru, Bolivia, Chile, and Argentina; Bowlesia 16 former neoendemic spp. native to French Guiana, Ecuador, Peru, Bolivia, Paraguay, Chile (3), Argentina (3 Argentina), Uruguay, and S+SE Brazil; Diposis 3 narrow endemics of C Chile (1), S Argentina (1), and Uruguay (1); Domeykoa 5 narrow endemics of Peru and N Chile; Eremocharis 9 narrow W SAM endemics of N Chile and Peru; Gymnophyton 6 Andes endemic spp. of N+C Chile (6 endemic), Bolivia, and NW Argentina; Pozoa  2 S SAM endemics of NW+S Argentina and C+S Chile. Mackinlayoideae: Centella 1-2 spp. native to CAM (exc. Panama), Bahamas, Turks-Caicos, Greater Antilles, Leeward & Windward Is., Trinidad-Tobago, Juan Fernandez Is., Galápagos, Colombia, Venezuela, Brazil (exc. N), Bolivia, Paraguay, C+S Chile, N Argentina, and Uruguay; Micropleura 2 spp. Mexico & neoendemic genus native to Guatemala, El Salvador, Honduras, Costa Rica, and Colombia (1 endemic). Saniculoideae: Eryngium 100+ subcosmopolitan spp. native throughout CAM, SAM, the Greater Antilles (exc. Cayman Is.), Leeward & Windward Is., and Trinidad-Tobago; Sanicula ~3 spp. native to CAM (exc. Belize), Colombia, Venezuela, Ecuador, Peru, Bolivia, C+S Chile, and NW+S Argentina.

Patagonian Argentina and Southern Chile Apiaceae Genera Include:

Apioideae: Ammi 1 sp. intro to S Chile and S Argentina; Anthriscus 1? sp. intro throughout Patagonia; Apium 2 spp. native throughout Patagonia & the Falkland Is.; Chaerophyllum 1-2 spp. native to S Chile, S Argentina, and Falkland Is.; Conium 1 sp. intro throughout Patagonia and Falkland Is.; Coriandrum 1 sp. intro to S Argentina; Cyclospermum 1 sp. native throughout Patagonia; Daucus 2 spp., including 1 native and 1 intro throughout Patagonia; Foeniculum 1 sp. intro throughout Patagonia; Helosciadium 1 sp. intro to S Argentina; Heracleum  1 sp. intro to S Argentina; Homalocarpus 1 narrow endemic sp. of N+C Chile found near N limit of Patagonia in SC Chile; Levisticum monospecific intro to S Chile; Lilaeopsis 1 sp. native throughout Patagonia and the Falkland Is.; Oligocladus monospecific Argentina endemic found in S Argentina; Osmorhiza 3 spp. native throughout the Patagonia region; Pastinaca 1 sp. intro to all of Patagonia; Petroselinum monospecific intro to S Argentina; Spermolepis 1 Argentina endemic in S Argentina.

Azorelloideae: Asteriscium 3-4 S SAM endemic spp. native throughout Patagonia; Azorella 15-20 spp. native throughout Patagonia and the Falkland Is., including several endemics of the Andes and Patagonia steppe; Bolax 2 Patagonian endemic spp. of S Chile, S Argentina, and the Falkland Is.; Bowlesia 4 endemic spp. native throughout Patagonia, now introduced in France and Pakistan; Diposis 2 SAM endemic spp. including 1 narrow endemic of S Argentina; Pozoa 1-2 S SAM endemics of the Patagonia region. Mackinlayoideae: Centella 1 sp. native to C+S Chile. Saniculoideae: Eryngium ~17 spp. native throughout Patagonia; Sanicula 2 spp. native throughout Patagonia.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get their definitions there.
  • Willis, Lyrae – Unpublished. Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as from personal observations of Apiaceae in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Delta: Watson, L., and Dallwitz, M.J. (1992 onwards). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • Flora of North America. (1993+). https://floranorthamerica.org/Main_Page.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx Accessed 2020–current.
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009 onwards). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/ Retrieved Winter 2020 – current.
  • USDA, NRCS. (2020). The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • WFO (2026): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Apocynaceae Dogbane and Milkweed Family

Asclepias variegata Redring Milkweed inflorescence with flowers. Learn how to identify the Apocynaceae family and the Milkweed subfamily
Asclepias variegata Redring Milkweed inflorescence with flowers. The milkweeds were once their own family but are now a subfamily of Apocynaceae.
Page Last Updated May 7, 2026.

Introduction to the Apocynaceae Family

The Apocynaceae family is an important family to learn how to identify. It is a family of often showy, beautiful flowers that are frequently fragrant but also often highly poisonous if ingested. The name “dogbane” comes from this toxic nature that has poisoned livestock and dogs. The Apocynaceae was recently revised to include the former Asclepiadaceae, or milkweed, family. There is an excellent reason to do so based on molecular phylogenetics, and the two families are similar in their morphology and other properties. However, there are a few notable differences in the Asclepiadoideae subfamily that I have included in detail in the descriptions below.   

Common Botanical Description

If you’re new to plant morphology, this section is a perfect beginner’s description to teach you how to identify the Apocynaceae family, with no need to know any scientific jargon. Below is some additional information on uses of the Apocynaceae and morphology pictures you can use to help identify family members and, finally, pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below the images in addition to genus-level distribution data.

Leaves and Stems of Apocynaceae: Annual herbs to shrubs, woody vines, and trees, all having a characteristic milky white juice (latex) they release when damaged. Leaves are usually simple (not compound) and may be thin to thick and leathery. They are usually arranged in opposite pairs or whorls around the stem. In some species, the leaves are modified into spines, and the stem may appear cactus-like.

Flowers of Apocynaceae: Flowers are often large, showy, symmetrical, and come in whites, reds, blues, and yellows and may be fragrant, foul-smelling, or odorless. Petals are joined at the base to form a funnel or trumpet shape, often with twisted or contorted lobes in bud. Most flowers have nectaries at their base to attract pollinators.

Reproductive Features of Apocynaceae: There are 5 male parts (stamens) that are often fused to the female parts (ovary, style, and stigma) to create a unique central structure called a “gynostegium.” There are typically two superior ovaries (sit above where petals attach), with an often large and conspicuous style head on top that acts as a platform to receive pollen.

Fruits of Apocynaceae: The fruits are often fleshy berries or drupes (like a cherry with a hard central pit) or dry capsules that split open when ripe. The Asclepiadoideae subfamily in particular is well-known for its large pod-like capsules (follicles) containing numerous seeds accompanied by long silky hairs (comose), which allow them to disperse in the wind.

Uses of Apocynaceae 

Most parts of many members of the Apocynaceae family are poisonous, hence the common family name of dogbane, since it has been used to poison dogs and other animals. The sap of most is a milky latex, which is often toxic but can sometimes be used for medicine and rubber.

Many ornamentals come from this family, including Oleander, Frangipani, Carissa, Plumeria, etc. The Asclepiadoideae subfamily of the Apocynaceae is an important source of cultivated succulents or vines from Asclepias, Hoya, Araujia, Ceropegia, Stapelia, Caralluma, Decabelone, etc. Carissa carandas is the source of the edible natal plum. 

Wildlife Values of the Apocynaceae Family

Asclepias species are required by the famous monarch butterfly, which uses them for reproduction, depositing its eggs, which hatch into caterpillars that eat the leaves. The monarchs are able to eat the toxic sap without ill effects.

Morphology of Apocynaceae in North America

The first table shows general Apocynaceae family morphology, but then there is a second one below it showing morphology specific to the Asclepiadoideae subfamily of the Apocynaceae.

Learn to identify the Apocynaceae family with morphology photos.
Learn how to identify the milkweed subfamily of the Apocynaceae family with morphology photos.

Some Apocynaceae Species Found in North America

Apocynoideae Subfamily

Adenium obesum Desert Rose flowers.

Adenium obesum – Sabi Star or Desert Rose

This is a drought-deciduous or evergreen shrub that is native to Africa but cultivated elsewhere. It has large swollen stems with a very enlarged swollen stem (caudex). Spiral leaves are clustered at branch tips. It has tubular pink flowers 4 – 5 cm wide with 5 petals.

Apocynum androsaemifolium plant with flowers showing opposite leaves and tubular flowers characteristic of the Apocynaceae family.

Apocynum androsaemifolium – Spreading Dogbane

This is a branching herbaceous perennial with opposite and more or less egg-shaped to somewhat elliptical leaves. Flowers are small, bell-shaped, whitish-pink, and found mostly in terminal clusters like these. Native throughout North America.

Apocynum cannabinum inflorescence with flowers.

Apocynum cannabinum – Indian Hemp

Erect, herbaceous branching perennial with usually purplish-green stems and with opposite lance-shaped to egg-shaped leaves that usually have white hairs on the lower surface. Flowers are small greenish to cream-colored, usually in terminal inflorescences. Native throughout North America.

Cascabela ovata leaves and drupe showing simple leaves and a fleshy drupe characteristic of some members of the Apocynaceae family.

Cascabela ovata—Yellow Oleander

A perennial shrub or tree with spiral, moderately thick, and leathery leaves with prominent veins (pinnate). Flowers are large, showy, tubular, and yellow. The fruit is large, ~rounded and somewhat plum-like but with multiple large stony seeds, turning purplish when mature. Native and common in Mexico and also Central America.

Nerium oleander fragrant flowers

Nerium oleander – Oleander or Nerium

This cultivated Apocynaceae species is originally from the Mediterranean. It has thick, leathery, lance-shaped leaves in whorls of 2 – 3. Flowers are deeply 5-lobed with a fringed corolla around the mouth of the floral tube as seen here in the center of the flower. Flowers are intensely fragrant (perfume).

Pachypodium rutenbergianum flowers, leaves, succulent stems, and spines

Pachypodium rutenbergianum—Madagascar Palm

This is a unique-looking tree 3 – 8 m tall with very thick succulent branches, long spines, and an enlarged stem (caudex). It has long, leathery, oblong to lance-shaped leaves. Flowers are white with a yellow center, petals spreading in a pinwheel shape. Native to Madagascar & cultivated elsewhere.

Trachelospermum jasminoides flowers and leaves

Trachelospermum jasminoides—Star Jasmine

This is a vigorous, medium-sized evergreen shrub or vine with elliptic to egg-shaped hairless, glossy, green, leathery leaves. Flowers are in few-flowered clusters on long stalks; they are very fragrant and hairy inside the floral tube. This Apocynaceae member is native to Asia but widely cultivated elsewhere.

Asclepiadoideae Subfamily

Asclepias arenaria Sand Milkweed Plant with flowers showing undulate leaf margins

Asclepias arenaria – Western Sand Milkweed

Erect herbaceous perennial with densely woolly leaves and stems. Leaves are broadly egg-shaped with wavy (undulate) margins seen in the photo. Flowers are found in the leaf axils and are greenish to cream-colored. This species is endemic to the Great Plains region of the USA, very drought-tolerant.

Asclepias asperula Antelope horns plant with curved follicles; follicles are characteristic of the Apocynaceae, especially in the Asclepiadoideae subfamily.

Asclepias asperula—Antelope Horns

This is a clump-forming, 1-2 ft tall perennial with long, narrow leaves that are usually folded lengthwise as shown in the photo. Flowers are usually in roundish terminal inflorescences with greenish flowers with maroon highlights. The fruit is a follicles usually curve like a horn (see pic). It is native to the southwestern USA and northern Mexico.

Asclepias curassavica Tropical Milkweed inforescence with buds and flower

Asclepias curassavica – Tropical Milkweed

A perennial native of South America but becoming widely introduced elsewhere. Flowers have purple or red corollas and yellow or orange coronas in the center. Leaves are long and narrow with acute tips or tapering to a point. It is not recommended to grow this plant outside of the tropics because it is confusing monarchs into overwintering in temperate climates instead of migrating.

Asclepias engelmanniana follicles and narrow leaves

Asclepias engelmanniana – Engelmann’s Milkweed

A tall, slender perennial with 1 – 4 branching stems with long and very narrow linear leaves, usually in groups of 2 – 3. Pale green flowers without horns are crowded in umbels in the upper leaf axils. This species is endemic to the Great Plains of the USA.

Asclepias incarnata Swamp Milkweed inforescences with flowers

Asclepias incarnata—Swamp Milkweed

A herbaceous perennial that is branched on the upper part of the stem and has lance-shaped to linearoblong leaves. The flower’s corolla is pink to red (rarely white), and the horn curves inward past the hood. This lovely plant is native to swamps and other wet areas of eastern North America.

Asclepias latifolia Broadleaf Milkweed plant with fruit, showing broad cabbage-like leaves

Asclepias latifolia – Broadleaf Milkweed

This unbranched, 2 – 3 ft tall herbaceous perennial has very large, broad, somewhat egg-shaped or val leaves with prominent veins. Leaves are densely packed, appearing cabbage-like when young. Flowers are pale greenish to yellow, often hidden in axils by the large leaves. Native to the western USA and northern Mexico, mostly in the Great Plains.

Asclepias pumila Plains Milkweed inflorescence with flowers.

Asclepias pumila – Plains Milkweed

This short herbaceous perennial is usually less than 30 cm tall with narrow leaves that are densely packed, appearing whorled but are not upon close inspection. Flowers are light pink or white in terminal inflorescences. This is another endemic of the Great Plains region of the USA.

Asclepias speciosa Showy Milkweed star-shaped corona of hoods and horns characterisitic of the subfamily of the Apocynaceae.

Asclepias speciosa – Showy Milkweed

Herbaceous perennial up to 1.2 m tall with oppositely arranged, elongated, simple leaves. Large, hairy, pale pink to pinkish-purple flowers appear in dense umbel-like clusters and have very prominent hoods and hooks forming a star shape. Native to western North America.

Asclepias subverticillata Horsetail Milkweed inflorescence with flowers

Asclepias subverticillata – Horsetail Milkweed

This erect, spreading, herbaceous perennial may or may not be branched and grows to 1 m tall with sparse, narrow, linear leaves with margins that roll backward on the edge (revolute) and grow in whorls of 3 – 5 at branch nodes. Star-like whitish or cream flowers grow in the upper leaf axils. It is native to the central and southwestern USA and northern Mexico.

Asclepias subverticillata Common Milkweed infructescence of rough-textured follicles; follicles are a characteristic fruit of the Apocynaceae.

Asclepias syriaca—Common Milkweed

A tall perennial up to 1 m tall with large egg-shaped to somewhat elliptic leaves with pale green on the lower surface. Flowers are in somewhat drooping umbels in leaf axils and are usually pink (rarely white), with horns that bend inwards. The follicles have a very roughly textured surface (shown in the photo). Native to the eastern USA and Canada.

Asclepias tuberosa Butterfly Milkweed inflorescence of bright orange flowers

Asclepias tuberosa—Butterfly weed

A bushy herbaceous perennial to 60 cm tall, with a hairy stem that may or may not branch in the top half. Leaves are linear to narrowly egg-shaped and may be rranged opposite or alternate on branches. Flowers are in terminal flat-topped umbels of yellow to red-orange flowers. Native to southeastern Canada and the eastern and southwestern USA.

Asclepias variegata Redring Milkweed inflorescence with flowers

Asclepias variegata—Red Ring Milkweed

Herbaceous perennial with usually unbranched stems with several pairs of opposite broadly oblong to ovate leaves. Flowers in 1 – 4 rounded, compact umbels with showy white and pinkish-red-tinged flowers with spreading hoods. Native to eastern North America, rare in the north, most common in the southeastern USA.

Asclepias verticillata Whorled Milkweed inflorescence with flowers

Asclepias verticillata – Whorled Milkweed

An erect perennial with branching only at the top of the plant (or unbranched) with numerous narrow linear leaves in whorls of 3 – 6. Flowers are in several umbels crowded in the upper axils with white to greenish flowers with somewhat spreading hoods and horns that are larger than the hoods. Native to eastern North America and parts of western North America.

Asclepias viridis inflorescence of green flowers with purple hoods and no horns; somewhat unusual in the milkweed subfamily of the Apocynaceae.

Asclepias viridis – Green Antelope Horns

A herbaceous perennial common on overgrazed pastures and roadsides. Simple leaves, often with wavy (undulate) edges, are arranged alternately on the stems. Flowers are green with purplish hoods, and they lack the horns usually present in the genus. This species is endemic to the south-central and some of the southeastern USA.

Cynanchum laeve leaves and flowers

Cyanchum laeve – Climbing Milkweed or Sand Vine

This is a climbing vine that is sparsely hairy with opposite heart-shaped leaves with palmate veins. Flowers are in clusters of 5 – 40 flowers in the leaf axils. The fruit is an egg-shaped follicle. Native to central and eastern USA, southeastern Canada

Funastrum clausum flowers and leaves

Funastrum clausum—White Twinevine

A tendril-climbing perennial vine with linear to oblong leaves that are hairy and somewhat succulent. Flowers appear in the leaf axils and have soft-hairy white petals. This species is native to the southern USA, Mexico, and the tropical Americas.

Rauvolfioideae Subfamily

Carissa macrocarpa Natal Plum flower and leaves; shrubs with tubuiar flowers are common in the Apocynaceae family.

Carissa macrocarpa—Natal Plum

Spiny evergreen shrub with leathery, glossy dark green leaves. Flowers are very fragrant and white with long narrow petals. The fruit is a large, reddish, edible drupe (looks like a plum). Native to Africa, cultivated in southern North America and tropical Americas.

Catharanthus roseus Madagascar Periwinkle plant with flower

Catharanthus roseus – Madagascar Periwinkle

An evergreen subshrub with hairless egg-shaped, somewhat glossy leaves arranged in opposite pairs. Flowers are white with yellow or red centers or dark pink with red centers. Native to Madagascar but frequently cultivated in North America.

Plumeria rubra Red Frangipani in its native habitat in the dry season

Plumeria rubra – Red Frangipani

A medium to large shrub or small tree with a succulent trunk and branches. Drought-deciduous or semi-evergreen with large leaves clustered at branch ends. Flowers are white to pink and very fragrant in terminal umbels. Native to Mexico and tropical America, widely cultivated elsewhere.

Rauvolfia tetraphylla Devil Pepper leaves and fruit

Rauvolfia tetraphylla – Devil Pepper

A much-branched shrub or small tree with 4 (3 – 5) whorled egg-shaped, elliptic, or ~oblong leaves per node. Flowers are small and white and appear in clusters in the leaf axils. The fruit is a small red 2-seeded drupe (shown in photo). Native to Mexico and the tropical Americas.

Vinca major Greater Periwinkle flowers and leaves

Vinca major – Greater Periwinkle

A scrambling vine with opposite semi-evergreen leaves with a waxy surface and tiny hairs, sometimes with hairy margins. Flowers are pink to violet, fused at the base, with pinwheel-like lobes. Native to the western Mediterranean, widely cultivated as a groundcover, and highly invasive in some areas.

Vinca minor Periwinkle plant with flower, a common vine. This subfamily is variable making it a bit harder to identify than most of the Apocynaceae.

Vinca minor – Periwinkle

A scrambling vine with opposite hairless evergreen leaves that are smaller and narrower than V. major and never have a hairy margin. Flowers are similar to V. major but a little smaller. This Apocynaceae member is native to Eurasia, widely cultivated and highly invasive in some areas.

Scientific Botanical Description of the Apocynaceae

Learn even more about how to identify the Apocynaceae family and the milkweed subfamily with this scientific botanical description.

Habit & Leaf Form of the Apocynaceae Family

The Apocynaceae are mostly annual or perennial shrubs, lianas, or herbs but sometimes are trees, and those trees may have buttress roots (a few in Tabernaemontana and Dyera). All forms are laticiferous, containing a milky latex characteristic of the family. They are self-supporting or climbing-stem twiners, root climbers, or scramblers; when twining, they usually twine anticlockwise. Some plants may be switch plants with succulent, photosynthetic stems. Others have peculiar vegetative forms like the leaves modified as passive pitcher traps in Dischidia rafflesiana. Leaves are usually well-developed but may be very reduced. They are persistent and simple. Leaves are herbaceous, leathery, membranous, or rarely modified into spines. They may be attached alternately, opposite, whorled (three per whorl), or rarely spiral. The leaf lamina is entire and is frequently pinnately veined but may also be one-veined or pinnately to palmately veined. Leaves are usually exstipulate but may be stipulate and sometimes are reduced to colleters found in the stipular position. Domatia occur in 18 genera as pits, pockets, or hair tufts. The leaf lamina is usually dorsiventral, sometimes bifacial, or isobilateral in Nerium oleander. The epidermis sometimes contains crystal idioblasts. Diverse forms of hairs are often found, and they may be eglandular or glandular, often with glandular, shaggy hairs at the lamina bases and on the petiole. Complex hairs are usually not present (except for Pachypodium).

Flowers of the Apocynaceae Family

Plants are hermaphrodites; pollination is entomophilous and often conspicuously specialized with trapping mechanisms using modified stylar heads or specialized anthers. Flowers are terminal or axillary and either solitary or aggregated in panicles, cymose (often umbelliform), or rarely racemose inflorescences. The flowers are often large and showy and are usually bracteate and bracteolate. They are regular, 4-5 merous, tetracyclic, and may be malodorous, fragrant, or odorless. A hypogynous disk is usually present except in the Asclepiadoideae, where it is always absent. The perianth has ten parts in two whorls with a distinct calyx and corolla, which is isomerous. The calyx has five parts in one whorl, is usually connate (at least basally), is regular, and is quincuncial (Apocynoideae), imbricate (Asclepiadoideae), or valvular (Asclepiadoideae). The corolla also has five parts in one whorl and may or may not be appendaged. The corolla is connate with the lobes shorter than or longer than the tube. It is usually contorted or rarely valvate, is funnel-shaped or salverform and regular, and comes in various shades of white, yellow, red, orange, pink, purple, or blue.

Androecium of the Apocynaceae Family

The androecium can vary depending on the Apocynaceae subfamily or tribe. Still, there are five members that are adnate and inserted near the base (always Asclepiadoideae), midway down, or in the throat of the corolla tube. They are usually united with the gynoecium (most Apocynoideae) and may form a gynostegium (always in Asclepiadoideae), or they are free of the gynoecium (most Plumerieae, Periplocoideae). They are free of one another or coherent (always one adelphous in Asclepiadoideae) and are one whorled and consist of exclusively fertile stamens. The stamens are isomerous with the perianth and alternate with the corolla members. The anthers are filantherous to sessile and may or may not be appendiculate. In the Asclepiadoideae, the filaments are almost always appendiculate, with their short filaments ornamented from their bases with nectaries. They are separate, cohering, or connivent (often sagittate in Apocynoideae and distinct from one another but attached adaxially to the stylar head in Asclepiadoideae). The anthers are basifixed or adnate, non-versatile, tetrasporangiate, introrse, and usually bilocular and bisporangiate. In Asclepiadoideae, the anthers are appendaged with horny wings and membranous connective appendages contributing to the coronal complex.  

Gynoecium of the Apocynaceae Family

The gynoecium almost always has two carpels but occasionally may have 2–5 (6–8) (especially in Pleiocarpa). Sometimes the ovary is interpreted as two locular when it is actually the separate ovaries of a syncarpous gynoecium. The pistil has 1–2 cells. The gynoecium is synstylovarious to syncarpous or synstylous (Asclepiadoideae, Plumerieae, Pleiocarpa) and is often superior or otherwise partly inferior. When synstylous, the carpel has 2–50 ovules. Placentation is marginal when synstylous. The gynoecium is usually transverse but always median in Asclepiadoideae. There is usually one style but always two in Asclepiadoideae that are partially joined (free below) by the dilated style head with lateral stigmatic surfaces that alternate with the stamens. There is one stigma that is usually massively expanded and may be contracted in the middle with a ring of hairs or a membrane below. Stigmas are wet or dry types, papillate or not, and either Group II, III, or IV types. Placentation when unilocular with two placentas is parietal and when bilocular (mostly) axile or apical. When bilocular (usually), there are 5–50 (1-4) ovules per locule, and when unilocular or incompletely bilocular, there are 2–100. Ovules are pendulous, anatropous, unitegmic, tenuinucellate (Apocynoideae), or pseudocrassinucellate (Asclepiadoideae).

Fruit of the Apocynaceae Family

The fruits of Apocynaceae are often a fleshy or non-fleshy dehiscent capsule or schizocarp with two mericarps of follicles, berrylets, nutlets, or drupelets. Sometimes it is a fleshy indehiscent berry or a drupe. In Asclepiadoideae, it is always a pair of follicles with thin papery placental flaps or may be a single follicle by abortion of the other carpel. Seeds are oily and may or may not be endospermic or not. They are usually flat and conspicuously hairy (comose in Apocynoideae and Asclepiadoideae) or not and may be winged or wingless.

Taxonomy of the Apocynaceae

The Apocynaceae contains 4555 – 5100 species in 400 genera contained within five subfamilies. It is part of the Gentianales order of the core eudicots. The Asclepiadoideae was recently included in Apocynaceae, greatly increasing the number of species and genera. As with most families, thanks to molecular phylogenetics clarifying our understanding of the plant families, the genera and species have been undergoing revisions. If major updates have been made, I will also update the lists here.

  1. Apocynoideae is a diverse subfamily of annual or perennial herbs, shrubs, lianas, and the occasional tree. They exude milky latex, but it is sometimes clear. Leaves are simple and entire but otherwise variable. They usually are exstipulate or have very small stipules. The stamens are almost always usually united with the gynoecium but do not form a true gynostegium; they also lack pollinaria. 
  2. Asclepiadoideae is a subfamily that can usually be easily recognized by the presence of a gynostegium with its one adelphous stamens and two styles that are partially joined (free below) by the dilated style head with lateral stigmatic surfaces that alternate with the stamens. The stamens are often appendaged at their bases, forming “hoods” and “horns.” There is no hypogynous disk, and the gynoecium is also always median; both of these characteristics are unlike the rest of the family. The fruit is always a follicle (1-2). Most species exude the same milky latex as most of the family.
  3. Periplocoideae is a family of herbs, shrubs, vines, or rarely small trees. Colleters are present in the sinuses of the calyx. The stamens are free of the gynoecium (unlike most of the family, excluding Plumerieae in Rauvolfioideae), but the triangular anthers are adnate to the style head.  
  4. Rauvolfioideae is split into eleven diverse tribes. In general, they are mostly trees and shrubs with some herbs or lianas. They also have variable fruit and floral morphology. Sometimes the stamens are free of the gynoecium (unlike most of the family except Periplocoideae). 
  5. Secamonoideae is a small subfamily of lianas or vines that climb by twining; rarely, they may be shrubs. Sometimes there are colleters on the adaxial leaf surface. 

Genera:

Apocynoideae: Adenium (6), Aganosma (9), Alafia (25), Allomarkgrafia (10), Allowoodsonia (1), Amalocalyx (1), Amphineurion (1), Anodendron (17), Apocynum (5), Artia (5), Asketanthera (4), Baharuia (1), Bahiella (2), Baissea (25), Beaumontia (9), Carruthersia (4), Cascabela (6 Not in APG), Chonemorpha (10), Cleghornia (2), Cycladenia (1), Dewevrella (1), Echites (14), Ecua (1), Elytropus (1), Epigynum (5), Eucorymbia (1), Farquharia (1), Forsteronia (45), Funtumia (2), Galactophora (6), Holarrhena (6), Hylaea (2), Ichnocarpus (3), Isonema (3), Ixodonerium (1), Kibatalia (15), Laubertia (4), Macropharynx (15), Malouetia (32), Mandevilla (183), Mascarenhasia (8), Mesechites (8), Micrechites (15), Motandra (3), Neobracea (8), Nerium (1), Odontadenia (23), Oncinotis (8), Pachypodium (23), Pacouria (3), Papuechites (1), Parepigynum (1), Parsonsia (85), Pentalinon (2), Pinochia (4), Pleioceras (5), Pottsia (3), Prestonia (63), Rhabdadenia (3), Rhodocalyx (2), Salpinctes (1), Sindechites (1), Spirolobium (1), Stephanostema (1), Stipecoma (1), Streptoechites (1), Strophanthus (39), Temnadenia (2), Thenardia (3), Thoreauea (3), Thyrsanthella (1), Tintinnabularia (3), Trachelospermum (10), Urceola (21), Vallaris (2), Wrightia (34)

Asclepiadoideae: Adelostemma (?), Aidomene (?), Anemotrochus (3), Anisopus (4), Anisotoma (3), Apteranthes (7), Araujia (13), Asclepias (210),Aspidoglossum (37), Aspidonepsis (5), Astephanus (2), Asterostemma (1), Australluma (2), Barjonia (7), Blepharodon (4-6), Calciphila (2), Calotropis (3), Campestigma (1), Caralluma (31 Not in APG), Ceropegia (473), Chlorocyathus (2), Cibirhiza (3), Cionura (1), Conomitra (1), Cordylogyne (2), Cosmostigma (3), Cristobalia (4 Not in APG), Cynanchum (259), Diplolepis (14), Dischidanthus (2), Dischidia (128), Ditassa (114), Dittoceras (?), Dolichopetalum (1?), Echidnopsis (36? = Ceropegia), Emicocarpus (1), Emplectanthus (3), Eustegia (5), Fanninia (1), Fischeria (17), Fockea (6), Funastrum (22), Glossonema (?), Glossostelma (12), Gomphocarpus (20), Gongronema (7), Gongronemopsis (9), Gonolobus (~140), Graphistemma (? = Cyanchum), Gunnessia (1?), Gymnema (53), Gymnemopsis (5?), Gyrostelma (1), Harmandiella (1), Hemipogon (9), Heterostemma (45), Heynella (1), Hoya (569), Hypolobus (1), Ibatia (36), Jobinia (25), Kanahia (2), Lachnostoma (16), Lagarinthus (?), Leptadenia (9), Lygisma (5), Macroscepis (21), Margaretta (1), Marsdenia (69-84), Matelea (181), Metastelma (94), Microloma (11), Minaria (22), Miraglossum (7), Monsanima (2), Nautonia (1), Neoschumannia (3), Nephradenia (5), Oncinema (1), Oreosparte (3), Orthosia (55), Oxypetalum (139), Oxystelma (2), Pachycarpus (39), Papuahoya (3), Parapodium (3), Pentacyphus (3), Pentasacme (6), Pentatropis (4), Peplonia (9), Pergularia (2), Petalostelma (12), Pherotrichis (4), Philibertia (45), Piaranthus (9), Pseudolachnostoma (12), Pycnorhachis (1), Rhyssolobium (1), Rhytidostemma (8), Riocreuxia (10), Rojasia  (1), Ruehssia (146), Sarcolobus (22), Schizoglossum (26), Schizostephanus  (2), Schubertia (6?), Scyphostelma (39), Sicyocarpus (1), Sisyranthus (15), Solenostemma (1), Stathmostelma (14), Stelmagonum (1), Stenostelma (6), Stephanotis (18), Stigmatorhynchus (2), Suberogerens (1),Talayotea (2), Tassadia (31), Telosma (9), Topea (2 Not in APG), Treutlera (1), Tweedia (6), Tylodontia (4), Vincetoxicum (264), Woodia (3), Xysmalobium (40).

Periplocoideae: Atherandra (1), Baroniella (10), Baseonema (1), Batesanthus (3), Buckollia (2), Camptocarpus (9), Cryptolepis (34), Cryptostegia (2), Decalepis (5), Ectadium (2), Epistemma (4), Finlaysonia (6), Gymnanthera (2), Hemidesmus (1), Ischnolepis (1), Maclaudia (1), Mondia (2), Myriopteron (1), Pentopetia (23), Periploca (17), Petopentia (2), Phyllanthera (10), Raphionacme (36), Sacleuxia (2), Sarcorrhiza (1), Schlechterella (2); Stomatostemma (1), Streptocaulon (5); Tacazzea (5), Telectadium (3), Zygostelma (1). 

Rauvolfioideae: Acokanthera (5), Allamanda (16), Alstonia (44), Alyxia (117), Ambelania (3), Amsonia (17), Ancylobothrys (8), Anechites (1), Aspidosperma (80), Bousigonia (3), Callichilia (6), Calocrater (1), Cameraria (7), Carissa (11-15), Carvalhoa (2), Catharanthus (9), Cerbera (6), Cerberiopsis (3), Chamaeclitandra (1), Chilocarpus (14), Clitandra (1), Condylocarpon (7), Couma (5), Craspidospermum (1), Crioceras (1), Cyclocotyla (1), Cylindropsis (1), Dictyophleba (6), Diplorhynchus (1), Dyera (2), Geissospermum (5), Gonioma (2), Hancornia (1), Haplophyton (2), Himatanthus (9), Hunteria (9), Kamettia (2), Kopsia (25), Lacmellea (24), Landolphia (63), Laxoplumeria (5), Lepinia (4), Lepiniopsis (2), Leuconotis (4), Macoubea (3), Melodinus (25), Microplumeria (1), Molongum (3), Mortoniella (1), Mucoa (2), Neocouma (2), Ochrosia (44), Orthopichonia (6), Parahancornia (7), Petchia (8), Picralima (1), Plectaneia (3), Pleiocarpa (7), Plumeria (18), Pteralyxia (2), Pycnobotrya (1), Rauvolfia (78), Rhigospira (1), Saba (3), Schizozygia (1), Skytanthus (3), Spongiosperma (6), Stephanostegia (2), Strempeliopsis (2), Tabernaemontana (126), Tabernanthe (2), Thevetia (3), Vahadenia (2), Vallesia (10), Vinca (7), Voacanga (22), Willughbeia (16),

Secamonoideae: Calyptranthera (13), Genianthus (17? or syn. of Secamone), Goniostemma (2), Pervillaea (5), Secamone (150), Secamonopsis (2), Toxocarpus (29), Trichosandra (1).

Distribution of the Apocynaceae

The Apocynaceae are mostly subtropical to tropical with a few temperate representatives. They are cosmopolitan in distribution, and they are widely found throughout the entire Americas, including some herbaceous Apocynaceae species that are found as far north as Arctic Canada. 

Distribution of Apocynaceae in the Americas

Canadian Apocynaceae Genera Include:

Apocynoideae: Apocynum 2 spp. native in almost all of Canada, including the Arctic but excluding NU and Labrador, 1 NAM hybrid sp. also native in this range. Asclepiadoideae: Asclepias 15 spp. native in all S provinces but intro in NL (absent in Labrador); Cynanchum 1 sp. intro in ON; Vincetoxicum 3 spp. intro to BC, ON, QC, and NB. Rauvolfioideae: Vinca 2 spp. intro to BC, ON, QC, NS, and NB.

USA Apocynaceae Genera Include:

Apocynoideae in the USA: Apocynum 2 spp. native to all of the USA including AK, plus 1 NAM hybrid sp. native in this range; Beaumontia 1 sp. intro HI?; Cycladenia monospecific SW USA endemic of CA, OR, AZ, and UT; Echites 1 sp. native to FL; Mandevilla 4 spp. native to AZ, NM, and TX; Nerium monospecific sp. intro to most of S border states exc NM, also in UT and CA; Pentalinon 1 Mexico, CAM & Caribbean endemic sp. native to S FL; Rhabdadenia 1 S NAM & neoendemic sp. native to FL; Thyrsanthella monospecific SE USA endemic of OK, TX, MO, AR, LA, IL, IN, KY, TN, MS, AL, GA, FL, SC, NC, VA, and DE; Trachelospermum 2 spp. intro TX, OK, MO, AR, LA, MS, AL, GA, FL, SC, NC, VA, MD, DE, TN, KY, IL, and IN. 

Asclepiadoideae in the USA: Araujia 2 former neoendemic spp. intro to CA, GA, and FL; Asclepias 75 spp. native to all the continental states and HI, with some intro in some states; Calotropis 2 spp. intro to CA, FL, and HI; Cynanchum 17 spp. native and intro to most of the USA exc WA, OR, WY, MT, CO, ND, and SD; Funastrum 5-7 spp. native to CA, NV, UT, CO, AZ, NM, TX, OK, AR, and FL; Gomphocarpus 1 sp. intro to CA?; Gonolobus 3 spp. native to AZ, KS, OK, TX, MO, AR, LA, IL, IN, KY, TN, MS, AL, GA, FL, NC, SC, VA, and MD; Hoya 1sp. intro to HI; Matelea 24 spp. (may be less now that genus has been reduced) sp. native to the most southern states; Metastelma ? sp. native to AZ, NM, TX, and FL (USDA-NRCS considers it a syn. of Cyanchum); Orthosia 1? sp. native to MS, FL, GA, and SC; Pherotrichis 1 sp. native to AZ; Suberogerens monospecific endemic of W+SE Mexico, and Ver; Vincetoxicum 3 spp. intro to CA, NE, KS, MN, MO, MI, WI, IL, IN, KY, OH, PA, NJ, MD, NY, CT, RI, MA, VT, NH, and ME.

Periplocoideaein the USA: Cryptostegia 2 spp. intro to TX and FL; Periploca 1 sp. intro to KS, OK, TX, TN, VA, PA, NJ, NY, CT, and RI.

Rauvolfioideae in the USA: Allamanda 1 former neoendemic sp. intro to FL; Alstonia 1 sp. intro to FL and HI; Alyxia 1 Pacific sp. native to HI; Amsonia 16 spp., including 14 native to all of S USA from CA E to NJ but excluding WV, which includes 7 endemic to the USA and 3 narrow endemics of AZ (2) and NM (1), and includes 2 spp. intro to NY, CT, and MA; Carissa 1 sp. intro to AZ, TX, and FL; Catharanthus 1 sp. intro to CA, TX, LA, MS, FL, GA, SC, NC, OH, and HI; Haplophyton 1 S NAM & N CAM endemic spp. native to AZ, NM, and TX; Ochrosia 4 spp. native to HI and intro to FL; Plumeria 1 sp. native FL; Pteralyxia 2 spp. endemic genera of HI; Rauvolfia 3 spp. inc. native in FL, and intro and native in HI; Tabernaemontana 2 spp. native to FL and intro HI; Thevetia 1 Neo sp. intro to TX, FL, and HI; Vallesia 2 Americas endemic spp., inc. 1 native and 1 intro to CA and FL; Vinca 3 spp. intro to most of USA except NV, OK, CO, WY, ND, and SD.

Mexico Apocynaceae Genera Include:

Apocynoideae in Mexico: Apocynum 1 sp. native to N Mexico, plus 1 NAM hybrid sp. native in this range; Cascabela 5 spp. native to all of Mexico, including 3 endemics; Echites 7 NAM, CAM & Caribbean spp. native to most of Mexico, exc. BC, BCS, Son, and Sin, and inc. 1 endemic to SW+C Mexico; Forsteronia 1-3? Neoendemic sp. native to SW+SE Mexico, Ver, Chp; Laubertia 1 Mexico endemic sp. in most of Mexico exc. Ver; Mandevilla ~22 spp. native (~1/2 endemic) to all of Mexico; Mesechites 1 Mexico & Neoendemic sp. native to Mexico except for BC, BCS, Son, Sin, Mex, Cd Mex, Pue, Tlx, and Mor; Nerium monospecific sp. intro to most of Mexico exc. BC, BCS, Son, Sin, and Ver; Odontadenia 1 neoendemic sp. native SW+SE Mexico, Chp; Pentalinon 1 Mexico, CAM & Caribbean endemic s.p native SW+C+SE Mexico; Pinochia 2 Mexico & N Neo endemic spp. native to SW+SE Mexico, Chp, and Ver; Prestonia ~5 soo native to most of Mexico exc. BC, BCS, Son, and Sin; Rhabdadenia 1 S NAM & neoendemic sp. native to Ver, Cam, Tab, Yuc, QR, and Chp; Thenardia 3 Mexico & Honduras endemic spp. native SW+C+SE Mexico, inc 2 endemics; Thoreauea 3 spp. endemic genus of SW Mexico + Ver; Tintinnabularia 2 Mesoamerica endemic spp. native Oax, Chp, and Ver, inc 1 narrow endemic of Ver.  

Asclepiadoideae in Mexico: Asclepias 70+ spp. native to all of Mexico; Calotropis 1 sp. intro to SW+C+SE Mexico; Cynanchum several spp. native to all of Mexico; Fischeria 1 Neoendemic sp. native to SW+C+SE Mexico; Funastrum 13 spp. native throughout all of Mexico, including 3 endemics; Gomphocarpus 1 sp. intro to SW Mexico; Gonolobus ~50 spp., about half of which are endemic and native to all of Mexico; Jobinia 1 sp. native Ver; Macroscepis 5 Mexico & neoendemic spp. native to all of Mexico, including 2 narrow endemics; Matelea ~70 spp, ~60% endemic to all of Mexico; Metastelma ~30 spp. native to all of Mexico, including Mexican Pacific Is., including many endemics; Orthosia 9 spp. inc. 6 endemics native to all of Mexico; Oxypetalum 1-2 spp. native to NE+SW+SE Mexico, and Ver; Pherotrichis 4 Mexico & N CAM endemic spp. native to most of Mexico except Ver, including 2 Mexican endemics of SW Mexico; Ruehssia ~40 Mexico & neoendemic spp. native to all of Mexico, with ~68% endemism; Stelmagonum monospecific endemic of SW Mexico; Talayotea 2 spp. endemic genera of most of Mexico except Ver;Tassadia 1 neoendemic sp. native to SW+SE Mexico, and Ver. 

Periplocoideae in Mexico: Cryptostegia 2 spp. intro to W+SE Mexico.

Rauvolfioideae in Mexico: Allamanda 1 former neoendemic sp. intro to SW+C Mexico; Alstonia 1 sp. native to SW+C+SE Mexico; Amsonia 4 spp. native to N Mexico; Aspidosperma 5-7+ neoendemic spp. native to SW+SE Mexico, Ver, including 2? endemics; Cameraria 1 former Mexico + N Neoendemic sp. native to SE Mexico; Carissa 1 sp. intro to SW+C Mexico; Catharanthus 1 sp. intro to SW+SE Mexico; Haplophyton 2 S NAM & N CAM endemic spp. native to most of Mexico except Ver; Plumeria 2 spp. native to all of Mexico; Rauvolfia 3-4 spp. native to all of Mexico; Tabernaemontana ~16 spp. native to all of Mexico, including 6 endemics; Thevetia 1 sp. native to most of Mexico except BC, BCS, Son, and Sin; Vallesia 8 Americas endemic spp. native to all of Mexico, inc. 5 narrow endemics; Vinca 1 sp. intro to SW+C+SE Mexico.

Neotropical Apocynaceae Genera Include:

Apocynoideaein the Neotropics:Allomarkgrafia 10 neoendemic spp. of Honduras S to Colombia, Ecuador, and Peru; Anodendron 1 sp. intro to Trinidad-Tobago; Asketanthera 4 Greater Antilles endemic spp. of Cuba, Dominican Republic, and Haiti; Bahiella 2 narrow endemic spp. of NE Brazil; Beaumontia 1 sp. intro to Costa Rica, Guatemala, El Salvador, and Honduras; Cascabela 3 spp. former native CAM S through tropical SAM to Peru, Bolivia, NE Argentina, SE Brazil (exc. NE Brazil), and intro to Bermuda, Bahamas, Antilles (exc. Cayman Is., Netherlands Antilles); Echites 13 NAM, CAM & Caribbean endemic spp. of CAM, Bahamas, Turks-Caicos, Greater Antilles, and Leeward Is., inc. 4 narrow endemics of Cuba (2), Costa Rica (1), Panama (1); Forsteronia 45 Neoendemic spp. native to Cuba, Jamaica, Trinidad-Tobago, CAM, and tropical SAM S to N Argentina (exc. N Chile); Funtumia 1 sp. intro to Cuba, Puerto Rico, Trinidad-Tobago, and El Salvador; Galactophora 6 SAM endemic spp. of Colombia, N+C Brazil, Venezuela, Guyana, Peru, and Bolivia; Hylaea 2 N SAM endemic spp. of S Venezuela and N Brazil; Laubertia 3 Neoendemic spp. native to Belize (endemic sp.), Colombia, Venezuela, N Brazil (endemic sp.), Ecuador, Peru, and Bolivia; Macropharynx 15 neoendemic spp. of Honduras, Costa Rica, Panama,  Colombia, Venezuela, Guyana, Suriname, Ecuador, Peru, Bolivia, Paraguay, Brazil, and N Argentina; Malouetia ~20 spp. native CAM (exc. El Salvador), Windward Is, Colombia, Venezuela, French Guiana, Suriname, Guyana, Brazil (exc. C Brazil), Peru, and Bolivia, and intro to Cuba; Mandevilla ~165 mostly Neoendemic spp. native to CAM, Cuba, Jamaica, Trinidad-Tobago, and tropical SAM S to N Argentina (exc. N Chile); Mesechites 8 neoendemic (1 also in Mexico) spp. native to CAM, Cuba, Hispaniola, Jamaica, Trinidad-Tobago, and tropical SAM S to N Argentina (excluding Uruguay and N Chile), inc. 5 endemics of Cuba (2), Hispaniola, Colombia, and Peru; Neobracea 8 narrow endemics of Cuba (7) and Cuba + the Bahamas (1); Nerium monospecific sp. intro to CAM and S Brazil; Odontadenia 23 neoendemic spp. native to CAM (exc. El Salvador), Hispaniola, Colombia, N+E+C Brazil, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, and Trinidad-Tobago, and intro to Windward Is.; Pentalinon 2 Mexico, CAM & Caribbean endemic spp. native to the Bahamas, Turks & Caicos, Greater Antilles, SW Caribbean, Leeward & Windward Is., Guatemala, Belize, El Salvador, Honduras, and Nicaragua; Pinochia 4 Mexico & N neoendemic spp. native to Guatemala, Belize, Nicaragua, Costa Rica, Panama, and Greater Antilles (exc. Cayman Is.), including 2 endemic to Greater Antilles; Prestonia 63 Mexico & neoendemic spp. native to CAM, tropical SAM S to N Argentina (exc. N Chile), Leeward & Windward Is, and Trinidad-Tobago; Rhabdadenia 3 S NAM & neoendemic spp. native to CAM, Bahamas, Greater Antilles, Leeward & Windward Is., Trinidad-Tobago, and tropical SAM S to N Argentina (exc. N Chile); Rhodocalyx 2 SAM endemic spp. of Peru, Bolivia, C+E+S Brazil, Paraguay, and N Argentina; Salpinctes monospecific endemic of S Colombia; Stipecoma monospecific SAM endemic of Bolivia and C+E Brazil; Strophanthus 1 sp. intro to Trinidad-Tobago; Temnadenia 2 spp. endemic genera of Brazil (exc. N); Thenardia 1 S Mexico + Honduras endemic sp. native Honduras; Tintinnabularia 2 Mesoamérica endemic spp. native to Guatemala, and Honduras (1 endemic).

Asclepiadoideaein the Neotropics: Anechites monospecific N neoendemic of Hoduras S to Colombia, Venezuala, Ecuador, and Peru, plus the Greater Antilles; Anemotrochus 3 Caribbean endemic spp. native to Bahamas, Turks-Caicos, Cuba, Hispaniola, and Jamaica; Araujia 13 former S SAM endemic spp. native Bolivia, C+SE+S Brazil, Paraguay, Uruguay, and Argentina, and intro in C Chile and elsewhere; Asclepias 12 spp. native to CAM, Bahamas, Turks-Caicos, Antilles (exc. Aruba), Trinidad-Tobago, Galapagos, and most of SAM excclduing Chile and intro in N Chile; Barjonia 7 SAM endemic spp. of Brazil, Suriname, and Bolivia; Blepharodon 4 SAM endemic spp. native Bolivia, C+S+SE Brazil, NE Argentina, and Paraguay; Calotropis 2 spp. intro to Bahamas, Turks and Caicos, Antilles, Trinidad-Tobago, Colombia, Venezuela, N+C+E Brazil, Paraguay, and Bolivia; Cristobalia 2 SAM spp. of Bolivia, Uruguay, N Argentina, and S Brazil; Cynanchum ~15 spp. native in CAM, Cuba, Jamaica, tropical SAM S to N Argentina (exc. N Chile); Diplolepis 9 S SAM endemic spp. native to N+C CHile, NW Argentina, including 4 narrow endemics of Chile, rest in Patagonia; Ditassa 114 SAM endemic spp. of Colombia S to N Argentina, except N Chile and Paraguay; Fischeria 17 Neoendemic spp. native from S Mexico S through CAM (exc. El Salvador), Cuba, Jamaica, Trinidad-Tobago, Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Paraguay, and NE Argentina; Funastrum 12+ former Americas endemic spp. native to the Antilles (exc. Aruba, Leewards, Netherlands Antilles), Trinidad-Tobago, CAM, and tropical SAM S to N Argentina (exc. N Chile), including 5 narrow endemics of Guatemala, Bolivia, Colombia, SE Brazil, and Galapagos, with 1 sp. now introduced to India; Gomphocarpus 1 sp. intro to CAM (exc. Belize), Cuba, Jamaica, Puerto Rico, Leeward Is, Colombia, Venezuela, Suriname, Brazil, Uruguay, NE Argentina, Ecuador, Peru, and Bolivia; Gonolobus ~100 spp. native to Greater Antilles (exc. Cayman Is.), Leeward & Windward Is., CAM, and tropical SAM S to N Argentina (exc. N Chile, Suriname, French Guiana); Gyrostelma monospecific endemic of C+SE Brazil; Hemipogon 8 SAM endemic spp. of Peru, Brazil, Bolivia, and Paraguay; Hoya 1 sp. intro to Cuba, Trinidad-Tobago, Puerto Rico, and Leeward Is.; Hypolobus monospecific endemic of NE Brazil; Ibatia 36 Neoendemic spp. of Honduras, Nicaragua, Panama, Cuba, Hispaniola, Puerto Rico, Lesser Antilles, Colombia, Venezuela, Guyana, Brazil, Paraguay, Ecuador, Bolivia, and N Argentina; Jobinia 25 Neoendemic spp. native to Guatemala, Honduras, Venezuela, Ecuador, Peru, Bolivia, C+S+E Brazil, N Argentina, and Uruguay; Lachnostoma 16 N SAM endemic spp. of Colombia, Ecuador, Peru, and Venezuela; Macroscepis 19 Mexico & neoendemic spp. native to CAM (exc. Belize), Colombia, Venezuela, Guyana, Ecuador, Peru, Bolivia, Paraguay, N Argentina, and Brazil; Matelea ~140 spp. native to CAM, Bahamas, Greater Antilles (exc. Cayman Is.), Leeward & Windward Is., Trinidad-Tobago, and tropical SAM S to N Argentina (exc. N Chile); Metastelma ~85-90 spp. native to the Bahamas, Turks and Caicos, Antilles, CAM, Colombia, Venezuela, Guyana, N+E+S Brazil, Ecuador, Peru, Bolivia, Paraguay, and N Argentina; Minaria 22 SAM endemic spp. of Brazil, Bolivia, and NE Argentina; Monsanima 2 narrow endemic spp. of E Brazil; Nautonia monospecific E SAM endemic of C+SE+S Brazil, Paraguay, and NE Argentina; Nephradenia 5 N SAM endemic spp. of Colombia, Venezuela, Guyana, N+E+C Brazil, and Bolivia; Orthosia ~48 neoendemic spp. native to CAM (exc. Belize), Colombia, Venezuela, E+S Brazil, Uruguay, N Argentina, Ecuador, Peru, Bolivia, Paraguay, Bahamas, and the Greater Antilles (exc. Cayman Is); Oxypetalum 139 mostly neoendemic spp. of Guatemala, Honduras, Costa Rica, Panama, Greater Antilles (exc. Cayman Is), Leeward Is, Colombia, Venezuela, Guyana, Brazil, Ecuador, Peru, Bolivia, Paraguay, Uruguay, and N Argentina; Pentacyphus 3 N SAM endemic spp. of high Andean Venezuela, Colombia, Ecuador, and Peru, including 1 narrow endemic of Caldas, Colombia; Peplonia 9 SAM endemic spp. of Peru, Bolivia, E+C+S Brazil, and Paraguay; Petalostelma 7 SAM endemic spp. of Brazil, Bolivia, Paraguay, and N Argentina; Pherotrichis 1 sp. native to Guatemala; Philibertia 41 W+S SAM endemic spp. of Ecuador, Peru, Bolivia, N+C Chile, and Argentina; Pseudolachnostoma 12 neoendemic spp. of Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Venezuela, N Brazil, and Trinidad-Tobago; Rhytidostemma 8 neoendemic spp. of Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana, C+N Brazil, Ecuador, and Peru; Rojasia monospecific SAM endemic of Bolivia, Paraguay, S Brazil, and NE Argentina; Ruehssia ~120+ spp. Mexico & neoendemic spp. native to CAM, much of the Antilles, and tropical SAM S to N Argentina (exc. N Chile); Schubertia 6? SAM endemic spp. of Colombia, Peru, Bolivia, Paraguay, Brazil, and N Argentina; Scyphostelma 39 neoendemic spp. of Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, and Bolivia; Stephanotis 1 sp. intro to Cuba, Hispaniola, Puerto Rico, Leeward Is, and Trinidad-Tobago; Tassadia 31 neoendemic spp. native to S Mexico, Cuba, Trinidad-Tobago, Belize, Nicaragua, Costa Rica, Panama, and tropical SAM S to Peru, Bolivia, Paraguay, and S Brazil; Topea 2 narrow endemic spp. of WC Brazil, Paraguay, and NE Argentina; Tweedia 6 S SAM endemic spp. of Bolivia, N+C Chile, Argentina, and Paraguay, spp. are mostly narrow endemics; Tylodontia 4 narrow endemic spp. of Cuba.  

Periplocoideae in the Neotropics: Cryptostegia 2 spp. intro to CAM (exc. Guatemala, Belize), the Bahamas, Antilles, Trinidad-Tobago, Colombia, Venezuela, and N+NE+C Brazil.

Rauvolfioideae in the Neotropics: Allamanda 15 spp. of former neoendemic genera native to tropical SAM S to Bolivia, NE Argentina, and S Brazil, excluding Ecuador and Paraguay and intro to Honduras, Nicaragua, Costa Rica, Panama, Guatemala, Belize, El Salvador, Trinidad-Tobago, and Galapagos; Alstonia 1 sp. native to CAM (exc. Belize) and intro in Trinidad-Tobago; Ambelania 3 SAM endemic spp. of Colombia, Venezuela, Peru, N+NE Brazil, French Guiana, Guyana, and Suriname; Anechites monospecific N neoendemic of Honduras S to Colombia, Venezuela, Ecuador, and Peru, and the Greater Antilles (exc. Cayman Is.); Aspidosperma ~78 neoendemic spp. native from S Mexico S through CAM and tropical SAM S to N Argentina (except N Chile), Hispaniola, Trinidad-Tobago, and Venezuelan Antilles; Cameraria 7 spp. native to Belize, Guatemala, Cuba, Hispaniola, Jamaica, and the Greater Antilles, including 6 single-island endemics of Cuba (4), Hispaniola, and Dominican Republic; Carissa 1 sp. intro to Honduras, Nicaragua, Bahamas, Jamaica, Puerto Rico, Leeward & Windward Is, and Trinidad-Tobago; Catharanthus 1 sp. intro to CAM, Bahamas, Turks-Caicos, Antilles (exc. Cayman Is., Venezuelan Antilles), SW Caribbean, the Galapagos, and French Guiana; Cerbera 1 sp. intro to Trinidad-Tobago; Chamaeclitandra monospecific African sp. intro to Trinidad-Tobago; Condylocarpon 7 Neoendemic spp. of Nicaragua, Costa Rica, Colombia, Venezuela, Brazil, French Guiana, Guyana, Suriname, Paraguay, Peru, Uruguay, NE Argentina, and Trinidad-Tobago; Couma 5 neoendemic spp. of Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Suriname, Guyana, French Guiana, N+NE Brazil, Ecuador, Peru, and Bolivia; Geissospermum 5 SAM endemic spp. of Venezuela, Guyana, Suriname, French Guiana, N+E Brazil, Peru, and Bolivia; Hancornia monospecific SAM endemic of Bolivia, Brazil, Paraguay, and Peru; Haplophyton 2 S NAM & N CAM endemic spp. native to Guatemala and intro in Cuba; Himatanthus 9 neoendemic spp. of Panama and tropical SAM S to Peru, Bolivia, C+SE Brazil; Kopsia 1 sp. intro to Trinidad-Tobago; Lacmellea 24 neoendemic spp. native to CAM (exc. El Salvador), and tropical SAM S. to Peru, Bolivia, and C+E Brazil; Landolphia 1 sp. intro to Trinidad-Tobago; Laxoplumeria 5 Neoendemic spp. of Panama, Colombia, Ecuador, Peru, Bolivia, N+C+SE Brazil, and French Guiana; Macoubea 3 neoendemic spp. of Costa Rica, Panama, Colombia, Peru, Bolivia, Venezuela, Guyana, Suriname, French Guiana, and N+NE Brazil; Microplumeria monospecific N SAM endemic of Colombia, Venezuela, and N Brazil; Molongum 3 N SAM endemic spp. of Colombia, Venezuela, and N Brazil; Mortoniella monospecific CAM endemic of Belize, Nicaragua, and Costa Rica; Mucoa 2 N SAM endemic spp. of Colombia, Venezuela, N Brazil, and Peru; Neocouma 2 N SAM endemic spp. of Colombia, Venezuela, Peru, and N Brazil; Pacouria 3 N SAM endemic spp. of Colombia, Venezuela, Guyana, Suriname, French Guiana, N+SE Brazil, Bolivia, and Peru; Parahancornia 7 NW SAM endemic spp. of Colombia, Venezuela, Guyana, Suriname, French Guiana, N Brazil, Peru, and Bolivia; Plumeria 18 former Americas endemic spp. native to CAM, Colombia, Venezuela, Guyana, Bahamas, Turks-Caicos, Antilles (exc. Aruba, Netherlands Antilles), and SW Caribbean, and now intro to Ecuador, Peru, Bolivia, Paraguay, Netherlands Antilles, and Trinidad-Tobago; Rauvolfia ~25 spp. native to CAM, Antilles (exc. Netherlands Antilles), tropical SAM S to N Argentina (exc. Uruguay, N Chile); Rhigospira monospecific N SAM endemic sp. of Colombia, Venezuela, N Brazil, and Peru; Skytanthus 3 narrow SAM endemic spp. of E Brazil (2), N+C Chile (1); Spongiosperma 6 N SAM endemic spp. of Colombia, Venezuela, and N+NE Brazil; Strempeliopsis 2 single island endemic spp. of Cuba (1) and W Jamaica (1); Tabernaemontana ~45 spp. native to the Antilles (exc Aruba, Netherlands Antilles), SW Caribbean, tropical SAM S to Peru, Bolivia, and NE Argentina; Thevetia 3 Mexico & neoendemic spp. native to CAM, Cuba, Colombia, Venezuela, N & C Brazil, Bolivia, Paraguay, and NE Argentina; Vallesia 6 Americas endemic spp. native to Cuba, Hispaniola, Jamaica, Belize, Guatemala, El Salvador, Honduras, Costa Rica, Colombia, Venezuela, N Brazil, Peru, Bolivia, Paraguay, and N Argentina, including 2 narrow endemics of Galapagos and Venezuela; Vinca 2 spp. intro to Guatemala, Costa Rica, Colombia, Venezuela, Bolivia, C Chile, and Uruguay.    

Patagonia Apocynaceae Genera Include:

Apocynoideae: Elytropus monospecific endemic C+S Chile, and S Argentina. Asclepiadoideae: Araujia 1? sp native to S Argentina; Asclepias 1 sp. native to S Argentina; Diplolepis 10 S SAM endemic spp. native to SC+S Chile and S Argentina, including 5 narrow endemics; Philibertia 1 SAM endemic sp. native to C Chile and S Argentina; Tweedia 2 S SAM endemic spp. native to C Chile and S Argentina. 

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as my own personal observations.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Flora of North America (1993+). https://floranorthamerica.org/Main_Page.
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout the fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current.

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Aristolochiaceae or Birthwort Family

Learn how to identify the Aristolochiaceae family, like this Aristolochia taliscana flower showing its unusual hairs used to temporarily trap flies for pollination
Aristolochia taliscana flower showing unusual hairs used to temporarily trap flies for pollination.
Page Last Updated May 5, 2026.

Introduction to the Aristolochiaceae Family

One important consideration when learning to identify the Aristolochiaceae family is that the Angiosperm Phylogeny Group (IV) recently combined two very small families into the Aristolochiaceae so it would not be paraphyletic. Since these new subfamilies are themselves very unique and different morphologically from the traditional Aristolochiaceae (which only included Asaroideae and Aristolochioideae), I am describing the subfamilies separately from the traditional Aristolochiaceae, which will be described in detail as one (differences between these two subfamilies can still be found in the taxonomy section below).

The Aristolochiaceae are a family with bizarre-looking flowers, and each genus has its own unique and often very bizarre morphology. Since I always believe the weirder the better, the first time I saw an Asarum and an Aristolochia, I was instantly in love with their unique beauty.

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to help you learn how to identify the Aristolochiaceae family, with no need to know any scientific jargon. Below this are morphology photos and pictures of some species found in North America to further help you with identification. But for researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below the images in addition to genus-level distribution data.

Leaves and Stems: These are mostly woody vines, shrubs, or herbs with simple (never compound) leaves on stalks (petioles) that are usually arranged in a spiral pattern and are often heart-shaped (cordate). Most have aromatic leaves that may have gland dots. The Hydnoroideae are leafless parasitic herbs that live almost entirely below the soil.

Flowers: The flowers are unusual and highly characteristic of the family, known for their often S-shaped floral tubes. They are often large and showy and often emit strong foul odors. They also often have unique and specialized hairs designed to trap flies. They are held there until the hairs wither, releasing them to escape and pollinate other flowers.

Reproductive Features: The bisexual flowers contain both male (stamens) and female (ovary, stigma, style) parts. There are typically 6-36 stamens, which are often fused to the female parts into a specialized central column called a gynostegium.

Fruits: Fruits are mostly dry capsules that split open when ripe to release their seeds. Rarely, they may be a fleshy berry or a nut.

Uses of the Aristolochiaceae Family 

A few Aristolochia and Asarum species are routinely cultivated as ornamentals. Aristolochia and some Asarum contain a toxin known to be carcinogenic to humans and rats, so they are never ingested in any way. However, Pipevine Swallowtail butterflies are immune to the toxin and lay their eggs on Aristolochia spp, and their caterpillars eat the leaves when they hatch. The fruits of Hydnora, on the other hand, have edible, fragrant pulp, which attracts animals, including porcupines, monkeys, jackals, rhinoceros, and armadillos, and are sometimes eaten by humans.

Morphology of Aristolochiaceae in North America

Learn how to identify the Aristolochiaceae, or Birthwort Family, with morphology photos.

Species of Aristolochiaceae I have Covered So Far

Aristolochioideae Subfamily

Aristolochia taliscana flower, bud, and heart-shaped leaf

Aristolochia taliscana – Mexican Birthwort

This is a lush medium green vine with heart-shaped (cordate) leaves with a deep cleft and a rounded to notched tip. Flowers are solitary in leaf axils with an S-shaped petal-like calyx in burgundy, purple, or brown, usually with yellow markings. The limb is covered with conspicuous purple hairs that trap pollinating insects. Native to western Mexico.

Aristolochia watsonii flower and arrow-shaped leaf with bizarre flowers charafcteristic of the Aristolochiaceae

Aristolochia watsonii – Watson’s Dutchman’s Pipe

This one has very deeply arrow-shaped (sagittate) green leaves with purple markings. The calyx tube is straight or S-shaped, and the limb has maroon spots and no elaborate hairs. It is native to the southwestern USA and northwestern Mexico.

Asaroideae Subfamily

Note that the genus Hexastylis was recently combined with Asarum based on phylogenetic research, though not all sources have accepted this change. Since it is still not widely accepted, and their floral morphology is different, I have kept it separate for now.

Asarum caudatum western wild ginger showing heart-shaped leaves and bizarre flowers characteristic of the Aristolochiaceae

Asarum caudatum – Western Wild Ginger

This long favorite of mine has classic heart-shaped leaves with very long stalks from the ground (no plant stem). Flowers are solitary and burgundy or greenish-yellow, with a 3-part calyx with very long lobes that taper to a fine point. It is native to the Pacific Northwest, from BC, Canada, south to California, and east to Idaho and Montana, USA.

Hexastylis arifolia Little Brown Jug showing heart-shaped leaves and bizarre little brown jug flowers characteristic of the Aristolochiaceae

Asarum (Hexastylis) arifolia Little Brown Jug

This low herbaceous perennial has triangular, arrow-shaped, heart-shaped, or diamond-shaped leaves that are often variegated (this one shows slight variegation). Flowers are jug-shaped (urceolate) and brown, purple, or reddish with three lobes that may or may not be flared (visible at the base of the leaf stalk). Native to the southeastern USA.

Hexastylis shuttleworthii showing heart-shaped leaves characteristic of the Aristolochiaceae

Asarum (Hexastylis) shuttleworthii – Shuttleworth Ginger

This one has round to heart-shaped & usually variegated leaves. Its flowers are similar to A. arifolia but much larger. It’s a narrow endemic of the southern Appalachians & the surrounding area, eastern USA.

Scientific Botanical Description of the Aristolochiaceae

Note that, with the addition of the two very unique subfamilies, I have kept my original description of Aristolochiaceae (=Aristolochioideae + Asaroideae) separate from the Hydnoroideae and the Lactoridoideae to make it easier to describe the species in larger groups. The Lactoridoideae is a rare endemic monospecific subfamily not encountered in North America, but I still include it since it is in the neotropical zone.

Habit & Leaf Form of Aristolochiaceae

Habit & Leaf Form of Traditional Aristolochiaceae

Shrubs, lianas, or herbs, but mostly woody vines bearing essential oils. Plants are green and photosynthesize. They are always perennial and do not have a conspicuous aggregation of leaves. They are usually climbing stem twiners, but sometimes they are self-supporting. Plants are mesophytic. Leaves are alternate, spiral, flat, and either herbaceous or herbaceous and membranous. They are petiolate, sheathing to non-sheathing, simple, aromatic, and may be gland-dotted, pellucid, or punctate. Lamina is usually entire or sometimes dissected, palmatifid, or trilobed and is often cordate. They are either palmately or pinnately veined and are cross-venulate. Leaves are exstipulate, but sometimes the first 1–2 leaves of suppressed axillary branches simulate stipules.  

Habit & Leaf Form of Hydnoroideae

Ectoparasitic herbs with a very peculiar vegetative form that lacks leaves and is more or less fungoid in habit. The vegetative component consists of a coarse, rhizome-like pilot root with many slender, unbranched haustorial roots that parasitize the roots of host plants. Plants are succulent and not green. 

Habit & Leaf Form of Lactoridoideae

Shrubs bearing essential oils with small, simple, alternate, distichous, petiolate, and gland-dotted leaves. The lamina is entire, obovate, emarginate, and pinnately veined

Flowers of Aristolochiaceae

Flowers of Traditional Aristolochiaceae

Plants are hermaphrodites. Pollination is entomophilous by Diptera, and the mechanism is conspicuously specialized via an elaborate system for trapping flies within the perianth tube using articulated hairs that end up withering to release the flies so they can visit another flower. Flowers can be solitary or aggregated in cymes, racemes, or spikes, and they are often axillary but sometimes terminal. Flowers are very unusual in shape and can be small, medium, or large. They are often malodorous and smelling of carrion but may also be odorless. They are actinomorphic to zygomorphic and tricyclic to pentacyclic.

The perianth has 3 or 6 parts that are joined in one whorl or two (then isomerous), and they either have a distinct calyx and corolla or are petaline. Only in Saruma is the corolla whorl conspicuous and well developed. The calyx has three parts in one whorl: connate, entire or blunt-lobed, and campanulate or tubular, with the tube often S-shaped. The calyx is either bilabiate, unequal, or regular; it can be persistent or not, and valvate or valvateinduplicate. The corolla, when present, has three parts in one whorl but, otherwise, is usually reduced or absent.

Flowers of Hydnoroideae

Plants are hermaphrodites, and pollination is via beetles. Flowers are solitary and arise endogenously from the pilot roots on short stalks that barely emerge above the ground, often with the lower parts remaining beneath the soil. They are medium to large in size, malodorous, regular, and either tricyclic or tetracyclic. A short free hypanthium is present. The perianth is sepaline or petaline or may be interpreted as tepals. It has 3–4(5) parts joined in one whorl. The lobes are valvate, often fleshy, often have retrorse bristles, and are white, red, pink, or brown in color. 

Flowers of Lactoridoideae

Plants are polygamodioecious with axillary flowers that are solitary or aggregated in cymes. Flowers are small, 3-merous, and tetracyclic. There is no free hypanthium or hypogynous disk. The perianth has three parts in one whorl and is sepaline.

Androecium of Aristolochiaceae

Androecium of Traditional Aristolochiaceae

The androecium has 4–36 members in one or two whorls that are free of the perianth but may be free of or united with the gynoecium or coherent via the gynostemium and one adelphous. All members are fertile, usually with 6 stamens, but there may be 4–36. Stamens are isomerous with the perianth to polystemonous, and they may have filaments or sessile anthers. Anthers are cohering or separate from one another, basifixed or adnate, non-versatile, and dehiscing via longitudinal slits. They are extrorse or both extrorse and introrse only in Heterotropa. Anthers are sometimes appendaged apically with the expanded connective that assumes stigmatic functions associated with the gynostemium, but they may also be unappendaged.

Androecium of Hydnoroideae

The androecium has 3–10 members that are free of the perianth but inserted on the hypanthium. They are one whorled and coherent and united into an ovoid synandrium of a thick annulus in Hydnora. The androecium is either made of fertile stamens or, in Prosopanche, includes staminodes. In Prosopanche 3–4(5), small fleshy staminodes alternate with the stamens. There are 3–4(5) stamens that are oppositisepalous and isomerous with the perianth. They either have sessile anthers in Hydnora or are filantherous in Prosopanche, where the very short filaments arise from the hypanthium, and the anthers are connate to form a dome or cap with a small central opening. Anthers may be separate from one another or coherent in Prosopanche. They are dehiscing via longitudinal slits or transversely and are extrorse.

Androecium of Lactoridoideae

There are 6 androecial members that are free of the perianth and one another and are two-whorled. The androecium is either exclusively all fertile stamens or sometimes includes staminodes. When present, three non-petaloid staminodes are internal to the fertile stamens. There are 6(3) stamens that are diplostemonous, narrowly laminar, and short. Anthers are adnate, non-versatile, and dehiscing via longitudinal slits. They are extrorse, bilocular, and almost as long as the blade. Anthers have short terminal appendages. 

Gynoecium of Aristolochiaceae

Gynoecium of Traditional Aristolochiaceae

The gynoecium has 4–6 carpels and is synstylovarious, syncarpous, or in Hexastylis (Asarum), it is synovarious. The gynoecium is usually inferior but sometimes may be partly inferior. The ovary is either 4–6 or 1 locular (where the septa sometimes incompletely intruded). An epigynous disk is either present or absent. There are 1 or 4–6 free or partially joined terminal styles. Placentation when unilocular is parietal, and when plurilocular it is axile. There are 50-100 ovules in a single cavity if unilocular and 20-50 when plurilocular. Ovules possess a funicle, are pendulous or horizontal, and are anatropous.

Gynoecium of Hydnoroideae

The gynoecium is 3(4) carpelled and is either partly or completely buried in the soil. The gynoecium is synstylovarious to syncarpous and inferior. The ovary is one locular but becomes occluded by the ingrowth of the accrescent placentas. There are 1 or 3 sessile and commissural (Prosopanche) stigmas that are mostly 3-lobed in Hydnora. Placentation is parietal (Prosopanche) or apical (Hydnora). There are 50-200 undifferentiated ovules that are orthotropous and tenuinucellate.

Gynoecium of Lactoridoideae

The gynoecium is three-carpelled and isomerous with the perianth. It is superior and semicarpous with the carpels in a single whorl and more or less basally connate. The carpel is shortly stylate with a decurrent stigma and 6 (4–8) ovules. Placentation is marginal, and the placenta is intruded. The ovary is three locular basally. Ovules have long funicles, are biseriate, anatropous, bitegmic, and weakly crassinucellate.

Fruit of Aristolochiaceae

Fruit of Traditional Aristolochiaceae

The fruit is usually non-fleshy, but sometimes it has a fleshy endocarp. Usually, they are dehiscent septicidal, valvular (usually basally but only rarely at the top), or an irregularly splitting capsule. Rarely are they an indehiscent berry or a nut, or in Saruma, they are a schizocarp with 4-6 mericarps of follicles. Seeds are endospermic, ruminate or not, and are oily. The embryo is rudimentary to weakly differentiated at the time of seed release.

Fruit of Hydnoroideae

The fruit is fleshy inside but with a more or less woody pericarp. It is a dehiscent circumscissile capsule (Prosopanche) or an indehiscent capsule or berry. Fruits contain 500–2000 tiny seeds that are endospermic.

Fruit of Lactoridoideae

The fruit is a non-fleshy aggregate with the fruiting carpels coalescing into a secondary syncarp, a dehiscent follicle. Seeds are endospermic and oily.

Taxonomy of Aristolochiaceae

The Aristolochiaceae family has 682 species in 7-9 genera of the Piperales order in the Magnoliids clade of basal angiosperms. This clade is not part of either the monocots or the dicots, which is why it is considered a basal angiosperm since it diverged early on.

APG IV has combined the former Hydnoraceae and Lactoridaceae because their exclusion would make the Aristolochiaceae paraphyletic.

  1. Aristolochioideae – Plants are mostly lianas or occasionally shrubs or herbs. Inflorescences are usually axillary, and the flowers or at least the floral buds, are zygomorphic. The gynoecium has 4-6(2-3) carpels that are constricted apically. Stigmas are wet or dry and are commissural in Aristolochia. Fruits are dehiscent and open laterally, acropetally or basipetally septicidal, or rarely adaxially if a schizocarp or berry. Seeds are winged.
  2. Asaroideae – Plants are rhizomatous perennial herbs. Flowers are solitary and terminal, the gynoecium is inferior to half inferior, and the stigma has multicellular papillae. The fruit of the Asarum is an irregularly dehiscent capsule and a follicular schizocarp in Saruma
  3. Hydnoroideae – Unusual parasitic herbs that resemble fungi and lack leaves entirely, living mostly underground with only fleshy, foul-smelling flowers emerging above the soil.
  4. Lactoris or Lactoridoideae – a monospecific subfamily with a narrow endemic shrub with very small flowers and dry fruits that split into pod-like segments.

Genera:

Aristolochioideae: Aristolochia (569), Thottea (46).

Asaroideae: Asarum (138), Saruma (1).

Hydnoroideae: Hydnora (10), Prosopanche (7).

Lactoridoideae: Lactoris monospecific (Lactoris fernandeziana).

Key Differences From Similar Families

The unique flowers of most members of the Aristolochiaceae make them hard to confuse with other families. The subfamilies Aristolochioideae and Asaroideae are so unique they are rarely confused, and Lactoroideae is a rare microendemic from Juan Fernández Island. The Hydnoroideae, on the other hand, despite their uniqueness, do have some superficially similar-looking families. However, the other families all contain features lacking in Hydnoraceae. They include the Apodanthaceae with its very small unisexual flowers; the Balanophoraceae with its fungus-like inflorescence with numerous small flowers; the Cytinaceae with its spicate inflorescence; the Mitrastemonaceae with its whorl of opposite decussate leaves and a superior ovary; and the Orobanchaceae with its zygomorphic flowers on racemes or spikes.   

Distribution of Aristolochiaceae

The Aristolochiaceae family is widespread in warm temperate to tropical areas around the world except in Australasia, and they are not present in arctic areas. The Lactoridoideae is a rare microendemic genus confined to Juan Fernández Island off the coast of Chile.

Distribution of Aristolochiaceae in the Americas

Canadian Genera Include:

Aristolochioideae: Aristolochia 2 spp. including 1 native to ON and 1 intro (sometimes referred to as Isotrema) to ON and QC. Asaroideae: Asarum 2 spp. native to BC, MB, ON, QB, and NB.

USA Genera Include:

Aristolochioideae: Aristolochia 16 spp., mostly native, some intro to most of the USA except WA, OR, NV, UT, ID, MT, WY, CO, ND, SD, NE, MN, and VT. Asaroideae: Asarum (inc. Hexastylis) 17 spp. native in most of the USA except NV, AZ, UT, WY, CO, NM, NE, and TX.

Mexico Genera Include:

Aristolochioideae: Aristolochia 47 spp. native throughout all of Mexico, including the Mexican Pacific Islands. Asaroideae: Asarum 1 sp. native Yuc.

Neotropical Genera Include:

Aristolochioideae: Aristolochia ~200 spp. native throughout Mexico, CAM, Bermuda, the Bahamas, Cuba, Hispaniola, Jamaica, Puerto Rico, the Leeward & Windward Is., and tropical SAM S to Santiago, C Chile, and N Argentina, and it is intro in the Venezuelan Antilles and Galapagos. Hydnoroideae: Prosopanche 7 neoendemic spp. native to Costa Rica, Peru, Bolivia, Argentina, Paraguay, NE & S Brazil. Lactoridoideae: Lactoris monospecific endemic of Juan Fernandez Island off the coast of Chile. 

Patagonia Genera Include:

Hydnoroideae: Prosopanche 1 sp. native to S. Argentina.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as my own personal observations.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Flora of North America. (1993+). https://floranorthamerica.org/Main_Page
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx Accessed 2020–current.
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009 onwards). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Stevens, P. F. (2001+). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Amaranthaceae Family

Amaranthus palmeri inflorescence showing tiny flowers in bloom. Learn how to identify the Amaranthaceae family with descriptions and morphology photos.
Amaranthus palmeri inflorescence showing tiny flowers in bloom.
Page last updated April 25, 2026.

Introduction to the Amaranthaceae Family

The Amaranthaceae family is part of the Caryophyllales order of core eudicots (dicots). When you learn to identify the Amaranthaceae family, you will see it is made of mostly herbs or shrubs. Many are often found growing in disturbed habitats, and many of them are introduced weeds, sometimes considered noxious. However, many edible plants come from the Amaranthaceae, including commercially important ones. Apart from being weedy, the family is known for having flowers that are very small and often hidden by more conspicuous bracts.

Most of us will recognize this family as weeds, often noxious ones, like certain Amaranth, Salsola, Iresine, and Bassia, to name a few. They are hardy herbs or sub-shrubs that can grow in deserts, saline or alkaline soils, and extreme heat. I have always thought that instead of waging war on amaranth with glyphosate (to which most are now resistant, which is why we have “Roundup Ready” crops so they can still kill the amaranth without killing the crops), we should grow it and eat it instead. It will grow in marginal lands already badly degraded by conventional farming (chemicals and tilling), and it needs very little water and very little fertilizer. You can also harvest about 60-80% of the leaves (eat them like spinach; they are delicious) without it impacting seed production. Seeds can be eaten in porridge or ground into a gluten-free flour for baking with (mmm… zucchini bread and carrot cake). It is far more nutritious than the corn and wheat fields it competes with. Lesson: We should eat our weeds instead of poisoning ourselves while trying to kill them!

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to teach you how to identify the Amaranthaceae family, with no need to know any scientific jargon. Below is additional information on uses and morphology, as well as pictures to help identify family members and individual species found in North America. But for researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below the images in addition to genus-level distribution data.

Leaves and Stems of the Amaranthaceae: Most members are annual or perennial herbs or shrubs, sometimes described as sub-shrubs due to their somewhat woody nature. Stems often have thickened joints (nodes) and may become fleshy (succulent) in some species (especially Chenopodioideae). Leaves vary in size from tiny scales to large, well-developed leaves but are also occasionally absent. Leaves are usually simple (not compound) and may be in opposite pairs or alternately arranged along the stem.

Flowers of the Amaranthaceae: Flowers are usually very small to minute and grow in spikes, heads, tight clusters, or branched groups. They are most often accompanied by conspicuous papery leaf-like structures (bracts) that stay on the plant for a long time. Rather than petals, most flowers are made of thin and papery parts that look more like scales or sometimes sepals, and they sometimes remain attached to the fruits as they grow, developing into wings, spines, or bumps to help protect or distribute them.

Reproductive Features of the Amaranthaceae: These are very diverse in the family, ranging from flowers with both male and female parts (bisexual) to separate male and female flowers on separate plants (dioecious). And since the flowers are tiny, these features are seldom easily visible, so we’ll skip it here. But you can check out the scientific botanical description below for more information.

Fruits of the Amaranthaceae: Fruits may be dry, bladder-like fruits (utricles) or capsules that may split open from the top, but sometimes they are small, fleshy berries or drupes (think cherry—fleshy with a stony pit). In some species, the fruit stays enclosed in and is dispersed with the flower parts. The seeds are known for containing starch, which helps them grow and makes them useful as a human food source.

Uses of Amaranthaceae 

The Amaranthaceae family has a few cultivated ornamentals that are from Amaranthus, Gomphrena, and Iresine, and some noxious weeds are notable from Amaranthus, Iresine, and Acnida. Amaranthus species are also used as food in many parts of the world. In North America, Native American peoples used to cultivate it for both the greens, eaten as a vegetable, and the seeds, which were often ground into flour. Amaranth seeds and flour are gaining renewed interest as they make a suitable replacement for wheat for those with grain allergies and gluten intolerance. Other commercially grown species include the sugar beet and Swiss chard from Beta vulgaris and spinach (Spinacia oleracea).

Morphology of Amaranthaceae in North America

Learn how to identify the Amaranthaceae family with simple descriptions and morphology photos.

Some Amaranthaceae Species Of North America

Amaranthoideae Subfamily

Achyranthes aspera inflorescence showing conspicuous bracts characteristic of the Amaranthaceae family.

Achyranthes aspera—Chaff Flower

This is a tall herb, up to 2 m tall, with silky or short-hairy stems and variable leaves, often somewhat egg-shaped to rounded. Sessile flowers are in a long spike surrounded by persistent awned bracts and bracteoles. These pantropical herbs are introduced to the southern USA and Mexico.

Amaranthus arenicola Sand Amaranth inflorescence

Amaranthus arenicola—Sand Amaranth

This dioecious annual (male and female flowers on separate plants) is common in sandy habitats, hence its common name. It has simple leaves and grows 2 m tall. Flowers occur in congested clusters (as shown in the photo). Native to the central and south Great Plains of the United States.

Branched inflorescence of Amaranthus palmeri, branched spikes like these are characteristic of the Amaranthaceae.

Amaranthus palmeri—Palmer’s Amaranth

This species is a fast-growing, aggressive herb that grows to 1.5+ m tall with diamond-shaped leaves on long stalks. Flowers are in terminal spikes or drooping branched inflorescences, as shown in the photo. Native to southern North America but now widely introduced and considered one of the most noxious weeds in the world. It is also edible and highly nutritious and can be used as a food crop in marginal environments.

Amaranthus retroflexus plant with inflorescences

Amaranthus retroflexus—Red Root Amaranth

It is a tall herb, growing to 3 m tall; is pubescent and hairy; and has rhombic or ovate leaves with long petioles. Flowers are in terminal and axillary spikes or panicles. It is believed to be native to eastern and central North America.

Camphorosmoideae Subfamily

Bassia scoparia infructescence with leaves and immature fruits

Bassia scoparia—Summer Cypress

This annual herb grows up to 1.5 m tall and has linearlanceolate leaves that often curl and a stem that is often reddish. Inconspicuous flowers grow in the leaf axils with much more conspicuous 0.5-1 cm long bracts. This Amaranthaceae member is a highly invasive Eurasian introduced species.

Chenopodioideae Subfamily

Atriplex canescens Four Wing Saltbush inflorescences with flowers

Atriplex canescens—Four-Wing Saltbush

This lovely woody shrub has the occasional spine on its branches, as well as plenty of more or less stalkless, somewhat oblong leaves and tiny flowers in congested spikes, as shown here. It produces conspicuous, winged greenish bracteoles that expand in fruit and turn papery and beige, becoming wings. This common Amaranthaceae member is native and widespread in the western USA.

Atriplex confertifolia Spiny Saltbush inflorescences with flowers; compact globular inflorescences are fairly common in the Amaranthaceae.

Atriplex confertifolia—Spiny Saltbrush

This one is similar to A. canescens but with shorter, wider leaves, unwinged fruits, and many more spines on its branches. It’s also native to dry areas of the western USA, especially in the Great Basin region.

Chenopodiastrum simplex leaf closeup, large leaves are not common among the Amaranthaceae family.

Chenopodiastrum simplex—Giant-Seeded Goosefoot

This herb has larger leaves than most of the family; they are 3–10 cm long and more than half as wide, with irregular, large pointy teeth, lobes, or lobules (of varying depths), mostly hairless stems and leaves, and small flowers without petals. It is native to much of North America.

Chenopodium album inflorescence with flowers

Chenopodium album—White Goosefoot

This plant is a very common erect annual with visible lines down its stems in green or reddish-purple. Their leaves are diamond-shaped to eggshaped, and their flowers are in compact cymes in terminal or lateral spikes. It was introduced from Eurasia and is widespread in North America.

Krascheninnikovia lanata infructescence with fruits and silky hairs that help them fly in the wind

Krascheninnikovia lanata—Winterfat

This is a small shrub with flat lance-shaped leaves. Leaves and stems are both white and woolly, turning reddish. Its fruits (utricles) are densely woolly and visible all winter, giving it its common name of “winterfat.” The silky hairs help them disperse in the wind. It is endemic to western North America from Canada south to northern Mexico.

Gomphrenoideae Subfamily

Froelichia floridana infructescence with fruits

Froelichia floridana—Large Cottonweed or Plains Snakecotton

A small annual with a single erect or ascending stem and leaves on the bottom third of the plant that are lanceolate to linear with silky-wooly hairs. Flowers are in hairy spikes, and the fruits are flask-shaped utricles. Native to the central and eastern USA and northern Mexico.

Gomphrena serrata inflorescence

Gomphrena serrata—Prostrate Globe Amaranth

An erect or prostrate perennial or annual, with a pilose stem and obovate to oblong pilose leaves that are sessile to petiolate. Flowers in a head covered by white lanceolate bracteoles. Native to the southeastern USA, south to the tropical Americas.

Iresine diffuse inflorescence with flowers

Iresine diffusa—Juba’s Bush

This is a branched erect to clambering annual to perennial with elongated egg-shaped leaves that are 3 – 14 cm long and hairless to woolly-hairy. Flowers are in woolly-hairy branched open inflorescences (males) or compacted ones (females). This species is native to South America and has been introduced into North America.

Salsoideae Subfamily

Halogeton glomeratus plant without flowers

Halogeton glomeratus—Salt Lover

This well-branched annual has narrow fleshy blue-green leaves tipped with long spines and flowers that grow densely along the branches with large, conspicuous, waxy, winged bracts. This species is native to Central Asia and introduced in western North America. It is often easy to spot because it is a halophyte, or salt-loving species that grows where many other plants cannot.

Salsola tragus in fruit. This introduced Amaranthaceae species is one of our most common tumbleweeds.

Salsola tragus aka Kali Tragus—Tumbleweed

This is a low-growing annual with branched and tangled stems and firm, more or less linear & usually very spine-tipped leaves. Flowers are surrounded by long spiny bracts and a disk of winged white/pink sepals. This Amaranthaceae member is from Russia and widely introduced in western North America. It is often our most common tumbleweed, breaking easily at the base when mature to roll across the plains and disperses its seeds.

Scientific Botanical Description of the Amaranthaceae

Habit & Leaf Form of Amaranthaceae

Most are annual or perennial (sometimes biennial) herbs or subshrubs, some shrubs, and a few vines and small trees. Sometimes they are succulent and nearly always halophytic in the Chenopodioideae, where they can even be more or less cactoid with succulent photosynthetic stems (Salicornia). Many species have thickened nodes on their stems. The wood of the perennial stem has a typical anomalous secondary growth, and only the subfamily Polycnemoideae has normal secondary growth.

Leaves may be well-developed, much reduced, or absent, and they may be minute to large in size. Usually they have no basal aggregation of leaves except sometimes in Ptilotus and the Chenopodioideae. The leaves are arranged alternate or opposite, and they are usually herbaceous but can also be fleshy or membranous. They are simple and may be petiolate or sessile, connate or not, sheathing or not, and flat or terete, and they are exstipulate and epulvinate. Leaves can be extremely variable in shape, but they mostly have entire or occasionally toothed margins. The lamina is either one-veined or pinnately veined. Various hair shapes and features are often present, and domatia are common. 

Flowers of Amaranthaceae

Plants are usually hermaphroditic but otherwise may be monoecious, andromonoecious, gynomonoecious, dioecious, androdioecious, or rarely polygamomonoecious. Flowers are solitary, paired (axillary), or aggregated in terminal or axillary cymes, spikes, heads (often with conspicuous persistent bracts and bracteoles), panicles, or thyrses, and they may or may not be pseudanthial. Flowers are small to minute in size; they are bracteate and bracteolate, regular, and tricyclic when hermaphrodite; otherwise, sometimes, they are cyclic or two-cyclic. Free hypanthium is usually absent except sometimes in the Chenopodioideae. The hypogynous disk is absent except in Chenopodioideae, where it is usually present. The perianth is dry and scarious, sepaloid or petaloid, or may be vestigial or absent (sometimes in Chenopodioideae). When present, there are 3–5 (1–6) parts to the perianth. The perianth is typically interpreted as a calyx with 3–5 (1–6) parts that may be free, partially connate, or connate (almost always in Chenopodioideae). The calyx is regular, imbricate, usually non-fleshy except in Chenopodioideae, persistent in fruit, and may or may not be accrescent, enclosing the fruit with wings, tubercles, or spines. 

Androecium of Amaranthaceae

The androecium has 2-5 (1–10) members that are either free of the perianth or adnate to the perianth or disk. All members are usually unequal except in the Chenopodioideae and some Ptilotus, where they are all equal. They are always in one whorl and are usually basally coherent, 1 adelphous, with filaments usually connate all or part of their length into a membranous tube, or in Chenopodioideae, they are often free from one another. The androecium can be made of exclusively fertile stamens (sometimes, or usually, in Chenopodioideae) or may often include staminodes, with some members lacking anthers or with petaloid or pseudostaminodial scales alternating with the true androecial members. When present, there are 1–3 or 5 staminodes in the same series as fertile stamens, and they may or may not be petaloid. There are 3-5(1) stamens that are oppositisepalous and usually isomerous with the perianth. The anthers dehisce via longitudinal slits, and they are introrse, unilocular to bilocular, and either bisporangiate or tetrasporangiate.  

Gynoecium of Amaranthaceae

The gynoecium is 2–3(4–5) carpelled, often 5 carpelled in Chenopodioideae, and the pistil is 1-celled. The gynoecium is synovarious, synstylovarious, or syncarpous, and it is usually superior but can sometimes be partly inferior. The ovary is 1-locular and sessile to stipitate. The gynoecium may be non-stylate or stylate. When present, there are 1-3 (2-4) styles that may be partially joined. There are 1-3 stigmas (1-3) that are of the dry type, papillate, and Group II type. Placentation is always basal. There are 1-2 or 5 ovules in the single cavity, and they are ascending or pendulous, non-arillate, campylotropous, bitegmic, and crassinucellate.

Fruit of Amaranthaceae

The fruit may be fleshy or non-fleshy. It may be a dehiscent, irregular, or often circumscissile capsule, or it may be an indehiscent berry, drupe, nut, capsule, or, often, a utricle. Sometimes in the Chenopodioideae, the fruit may be enclosed in the fleshy hypanthium or perianth, and sometimes, the gynoecia also combine to form a multiple fruit. Seeds are non-endospermic and contain starch. Perisperm is present (abundant, mealy) or sometimes absent in Chenopodioideae. Seeds have starch.

Taxonomy of Amaranthaceae

There are currently about 2000 – 2500 species in about 180 accepted genera of the Amaranthaceae family, which is part of the Caryophyllales order of core eudicots. There are currently 6–10 subfamilies, as this family is being reworked. Here I describe it with the 10 divisions I had, which APG shows in their list of genera, even though on the website it shows several have become tribes instead. Once the family work is complete, I will update the list again, but for now it reflects the most current data I have (May 5, 2026).     

  1. Amaranthoideae – Annual or perennial herbs to shrubs, occasionally small trees. The inflorescence is branched or not but is usually spike-like or capitate. The bracts are disarticulating, and bracteoles may or may not be papyraceous or scarious. The perianth is typically scarious, and it possesses staminodes or pseudostaminodes. Cuticle waxes lack platelets, and hairs are uniseriate
  2. Betoideae – Annual to perennial herbs or occasionally subshrubs or vines. Bracteoles are usually present; the perianth has 5 (3) parts and is persistent and accrescent in Beta. The androecium has 5 (1) members, and the gynoecium is sometimes partly inferior. The fruit is a circumscissile capsule or pyxidium.
  3. Camphorosmoideae – Often shrubby plants but sometimes annual herbs, often with C4 photosynthesis; leaves are terete, and ‘prickles’ are often present. Their hairs have swollen bases, styles are filiform with papillae all around. The perianth is fleshy and spiny.
  4. Chenopodioideae – Annual herbs or short-lived perennial herbs to shrubs that are more or less succulent and often have swollen nodes. They prefer dry and/or saline temperate to subtropical habitats. Flowers are small and have a greenish perianth that often becomes more elaborate after flowering and is accrescent with the usually indehiscent fruits.
  5. Corispermoideae – Annual herbs without C4 photosynthesis. The inflorescence is spicate, simple, compact, or sometimes globular, and lacking bracteoles. Leaves are mostly alternate, sessile, or petiole-like. Hairs are branched or stellate. Perianth has 0-5 membranous tepals that are not persistent.
  6. Gomphrenoideae – Annual to perennial herbs or subshrubs with a perianth that is free to connate. Anthers are bisporangiate, monothecal, and their filaments are more or less connate. Their stigma is either capitate or bilobed.
  7. Polycnemoideae – Annual to perennial herbs or small shrubs that often inhabit saline habitats. Leaves are needle-like or succulent, bracts are disarticulating, and bracteoles are large. Flowers are axillary, the perianth is petaloid but never modified, and the androecium is basally connate. This subfamily is distinguishable from all other members of Amaranthaceae by normal secondary growth. 
  8. Salicornioideae – Annual or perennial herbs to low shrubs, usually with articulated stems. Plants are glabrous, and leaves are usually opposite, more or less terete or scaly or reduced to a rim. The inflorescence is dense, spicate, and leafless. The perianth has 3-4 (2-5) parts and is typically at least partially connate. In fruiting, the perianth may remain membranous or become spongy, crustaceous, or horny.
  9. Salsoideae – Herbs, subshrubs, shrubs, or sometimes trees. Stems and leaves are often succulent, leaves are terete, and bracteoles are present. Scarious wings usually develop from the perianth in fruit. Anthers have colored appendages, and their stigmas are flattened.
  10. Suaedoideae – Annual to perennial herbs to shrubs that are usually glabrous and usually have well-developed terete leaves. The inflorescence is spicate axillary cymes that are loose, leafy, and possess bracteoles. Styles are filiform; stigma may be capitate but is papillate all around. The perianth is persistent and usually accrescent and enlarged or winged when in fruit.

Genera of the Amaranthaceae:

Amaranthoideae: Achyranthes (33), Aerva (7, 6 extant), Allmania (3), Allmaniopsis (1), Amaranthus (93), Arthraerua (1), Bosea (3), Calicorema (2), Celosia (46), Centema (2), Centemopsis (13), Chamissoa (3), Charpentiera (6), Chionothrix (2), Cyathula (44), Cyphocarpa (?), Dasysphaera (4), Deeringia (10), Digera (1), Eriostylos (1), Evelynastra (1), Hebanthodes (1), Henonia (1), Herbstia (1), Hermbstaedtia (14), Indobanalia (1), Kyphocarpa (4), Lagrezia (14), Lecosia (2, not in APG), Leucosphaera (1), Lopriorea (1), Marcelliopsis (3), Mechowia (2), Neocentema (2), Nothosaerva (1), Nyssanthes (4), Omegandra (1), Ouret (8), Paraerva (2), Pleuropetalum (3), Pleuropterantha (3), Polyrhabda (1), Pseudosericocoma (?), Psilotrichopsis (1), Psilotrichum (26), Ptilotus (127), Pupalia (4), Rosifax (1), Saltia (1), Sebsebea (1), Sericocoma (3), Sericorema (2), Sericostachys (2), Siamosia (1), Stilbanthus (1), Trichuriella (1), Volkensinia (1), Wadithamnus (1).  

Betoideae: Acroglochin (3), Aphanisma (1), Beta (10), Hablitzia (1), Oreobliton (1), Patellifolia (1).  

Camphorosmoideae: Bassia (20), Camphorosma (5), Chenolea (2), Didymanthus (1), Dissocarpus (4), Enchylaena (1?), Eremophea (2), Eriochiton (1?), Grubovia (6), Kirilowia (? = Bassia), Kochia (? = Bassia), Londesia (? = Bassia), Maireana (58), Malacocera (4), Neobassia (2), Neokochia (2), Osteocarpum (5), Panderia (?), Roycea (3), Sclerolaena (78), Spirobassia (1), Stelligera (? = Sclerolaena), Threlkeldia (2).

Chenopodioideae: Ambrina (?), Archiatriplex (1), Atriplex (250), Axyris (7), Blitum (12), Carocarpidium (?), Ceratocarpus (1), Chenopodiastrum (10), Chenopodium (133), Dysphania (47), Eokochia (1), Exomis (1), Extriplex (2), Grayia (4), Halimione (?), Holmbergia (1), Krascheninnikovia (3), Lipandra (1), Manochlamys (1), Microgynoecium (1), Neomonolepis (1), Oreosalsola (12? Not in APG), Oxybasis (14), Phyllotheca (2?), Proatriplex (1), Sclerochlamys (?), Spinacia (2), Stutzia (2), Suckleya (1), Teloxys (1)

Corispermoideae: Agriophyllum (6), Anthochlamys (5), Corispermum (75)

Gomphrenoideae: Alternanthera (107), Froelichia (15), Froelichiella (1), Gomphrena (144), Guilleminea (7), Iresine (34), Pedersenia (8), Pfaffia (33), Pseudogomphrena (?), Pseudoplantago (2), Quaternella (3), Tidestromia (6), Xerosiphon (2).

Polycnemoideae: Hemichroa (1), Nitrophila (4), Polycnemum (6), Surreya (2).

Salicornioideae: Allenrolfea (3), Arthrocaulon (3), Arthroceras (1), Halocnemum (3), Halopeplis (3), Halostachys (1), Heterostachys (2), Kalidium (8), Mangleticornia (1), Microcnemum (1), Salicornia (53), Tecticornia (48)

Salsoideae: Agathophora (1), Akhania (3), Anabasis (30), Arthrophytum (7), Brachylepis (?), Caroxylon (134), Climacoptera (39), Cornulaca (6), Cyathobasis (1), Gamanthus (5?), Girgensohnia (5), Halanthium (5), Halarchon (1), Halimocnemis (21), Halocharis (6), Halogeton (3), Halothamnus (22), Halotis (?), Haloxylon (12), Hammada (2), Horaninovia (8), Iljinia (1), Kali (? = Salsola), Kaviria (9), Lagenantha (2?), Micropeplis (1?), Nanophyton (8), Noaea (7), Nucularia (1), Ofaiston (1), Petrosimonia (11), Physandra (1?), Piptoptera (1), Pyankovia (3), Rhaphidophyton (1), Salsola (39), Seidlitzia (3?), Sympegma (2), Traganopsis (1), Traganum (2), Turania (4? Not in APG).

Suaedoideae: Baolia (1), Bienertia (4), Fadenia (1?), Suaeda (94). 

Key Differences From Similar Families

Amaranthaceae differs from the similar but very small (only two genera) Achatocarpaceae family because Amaranthaceae usually have a herbaceous habit vs. always being woody and always with normal secondary growth in Achatocarpaceae. Amaranthaceae also usually have dry fruits rather than the succulent berries found in the Achatocarpaceae.  

Distribution of Amaranthaceae

The Amaranthaceae family is a temperate to tropical, truly cosmopolitan family missing only from Antarctica and the most extreme Arctic areas. Widespread throughout North America, Central America, the Caribbean, and South America.  

Distribution of Amaranthaceae in the Americas

Canadian Amaranthaceae Genera Include:

Amaranthoideae: Achyranthes 1 sp. intro in ON; Amaranthus 20 spp. intro in BC, NT, NB, NS, PE, and NL Island, with some native in AB, SK, MB, ON, and QC. Camphorosmoideae: Bassia 2 spp. intro to all S provinces except NL, NS, and PE. Chenopodioideae: Atriplex 18 spp. native to all of Canada and Greenland; Axyris 1 sp. intro to all of S Canada exc PE and NL; Blitum 4 spp., including 2 native and 2 intro in all of Canada inc Arctic but ephemeral PE and NL (exc Labrador); Chenopodiastrum 4 spp. inc 1 native to all S provinces exc NL and 3 spp. intro into S Canada, YT, and NT; Chenopodium 19 spp. native to YT, NT, BC, AB, SK, MB, ON, QC, and NS, and intro in NB, PE, NL, NU (and Greenland); Dysphania 4 spp., including 3 intro to BC, ON, QC, NB, and NS and 1 native to ON, QC, and MB; Krascheninnikovia 1 sp. native to YT, AB, SK, and MB; Lipandra monospecific intro to BC, SK, MB, ON, QC, and NB; Oxybasis 6 subcosmopolitan spp. including 3 native to all of Canada including the Arctic but excluding PE and NL, and 3 spp. intro to E Canada; Spinacia 1 sp. intro and ephemeral in YT, NT, AB, and MB; Stutzia 1 NW NAM endemic sp. native to AB and SK; Suckleya monospecific N-C NAM endemic sp. native to AB and SK. Corispermoideae: Corispermum 6 spp., mostly native to BC, AB, SK, MB, ON, QC, YT, and NT. Gomphrenoideae: Froelichia 1 sp. intro to ON. Polycnemoideae: Polycnemum 3 spp. intro to BC and ON. Salicornioideae: Salicornia 5 spp. native to all of Canada except Labrador, it’s found in coastal and inland saline areas. Salsoideae: Salsola (inc. Kali) 3 spp. intro to all S provinces. Suaedoideae: Suaeda 4 spp. native to all of Canada except for Labrador.

USA Amaranthaceae Genera Include:

Amaranthoideae: Achyranthes 4 spp. including 2 intro to TX, LA, FL, AL, KY, WV, IN, OH, and MD, plus 2 spp. endemic to HI; Aerva 1 sp. native and believed extinct in HI; Amaranthus 45 spp. both intro and native in all of the USA, including HI and intro in AK; Celosia 4 spp., including 1 native to TX and FL and 3 intro to most of E USA from WI S to MS and all E exc MI, GA, SC, RI, MA, NH, ME, and inc MO, LA, KS, UT; Charpentiera 5 spp. endemic to HI; Digera monospecific sp. intro to NJ; Hermbstaedtia 1 sp. intro to MD. Betoideae: Aphanisma monospecific SW NAM endemic sp. native to CA; Beta 2 spp. intro to MT, OR, CA, UT, TX, MO, AL, SC, NC, VA, WV, PA, MI, NY, CT, MA, RI, NH, and ME. Camphorosmoideae: Bassia 6 spp. including 2 native and 4 intro to most of the USA except AR, GA, and FL, and intro to HI; Enchylaena 1 sp. intro to CA; Neokochia 2 W USA endemic spp. of OR, CA, ID, NV, MT, WY, CO, UT, AZ, NM, and TX; Spirobassia monospecific intro to PA, NJ, MD, NY, and CT. Chenopodioideae: Atriplex 94 spp. mostly native with some intro, found in most of the USA, including AK, but excluding AR and TN, and intro HI; Axyris 1 sp. intro to MT, CO, ND, WI, and MO; Blitum 4 spp. inc. 3 native and 1 intro to most of USA except AR S to LA and E to NC S to FL plus VA, MD, and DE; Chenopodiastrum 2 spp. including 1 native to most of the USA and AK but excluding OR, AZ, AL, GA, SC, and FL where intro and 1 sp. intro scattered throughout USA; Chenopodium ~40 spp. native and intro to all of the USA inc AK and HI; Dysphania 11 spp. native and intro to the entire USA, intro in AK and HI; Extriplex 2 SW NAM endemic spp. native to CA, inc 1 narrow endemic; Grayia 4 W USA endemic spp. from MT S to NM and all W; Krascheninnikovia 1 spp. native to W USA from ND S to TX and all states W; Lipandra monospecific sp. intro to OR, WI, IL, MI, PA, NJ, NY, CT, and ME; Neomonolepis monospecific former SW NAM endemic of CA, NV, and OR; Oxybasis 6 spp., including 2-3 native to most of USA except the SE states and 2-3 spp. intro to much of the same range but including FL, AL, IL, IN, KY, WV, VA, NC, and VT; Proatriplex monospecific S-C USA endemic of UT, AZ, CO, and NM; Spinacia 1 sp. intro to WA, OR, CA, UT, TX, KS, SC, NC, VA, OH, PA, CT, MA, NY, NH, and ME; Stutzia 2 NW NAM endemic spp. native to CA, NV, MT, WY, CO, ND, SD, and NE; Suckleya monospecific N-C NAM endemic sp. native to MT, ND, WY, CO, NM, AZ, NE, OK, and TX; Teloxys monospecific sp. intro to MI, NY, and AK. Corispermoideae: Corispermum 9 spp. native and intro in most of the USA except ME, VT, NH, PA, MD, DE, WV, VA, TN, NC, SC, LA, MS, AL, and FL, and it is native in AK and extinct in CA. Gomphrenoideae: Alternanthera 11 spp. including 1 native to FL and the rest intro to CA, AZ, NM, TX, OK, MO, AR, LA, IL, KY, TN, MS, AL, GA, SC, NC, VA, MD, PA, NJ, NY, and HI; Froelichia 6 former Americas endemic spp. native to most of the USA except WA, OR, ID, MT, NV, UT, ND, VT, RI, and ME, including 2 narrow endemics of TX, and intro CA?; Gomphrena 8 spp. native to AZ, NM, TX, LA, FL, GA, and VA, and intro to PA, MD, OH, NY, MA, and HI; Guilleminea 1 sp. native to CA, AZ, NM, TX, OK, CO, SC, MD, AR, and LA; Iresine 6 former Americas endemic spp. native to AZ, NM, TX, KS, OK, MO, AR, LA, IL, IN, KY, TN, MS, AL, GA, FL, SC, NC, MD, and PA and intro to HI; Tidestromia 5 S NAM & Caribbean endemic spp. native to CA, NV, UT, AZ, CO, NM, TX, LA, OK, KS, SD, MO, IL, and PA. Polycnemoideae: Nitrophila 2-3? SW NAM endemic spp. native WA, OR, CA, NV, UT, and AZ, including 1 narrow endemic of the Mohave Desert; Polycnemum 1 sp. intro to IN, NY, MD, DE, and NH. Salicornioideae: Allenrolfea 1 SW NAM endemic sp. native to OR, ID, CA, NV, UT, AZ, NM, and TX (other 2 are endemic to Argentina); Arthroceras monospecific SW NAM endemic sp. native to CA; Salicornia 5 spp. native most of the USA including AK but exc. AZ, OK, AR, TN, KY, IN, WI, WV, PA, and VT, and it’s intro in HI and MI, and extirpated IL, found in coastal & inland saline areas. Salsoideae: Caroxylon 1 sp. intro in CA; Halogeton 1 sp. intro in all of W USA from MT S to NM and all W, plus SD and NE; Salsola 6 spp. intro to all of the USA including HI; Suaedoideae: Suaeda 13 spp. intro and native to most of the USA except IA, AR, TN, WV, and VT, and it is native in AK.

Mexico Amaranthaceae Genera Include:

Amaranthoideae: Achyranthes 2 spp. inc 1 intro in all of Mexico and 1 native to S Mexico; Amaranthus ~35? spp. native through all of Mexico and intro to Mexican Pacific Is.; Celosia 3? spp. native to most of Mexico except where intro in Mex, Cd Mex, and Tlx; Chamissoa 1 Mexico & neoendemic sp. native in all of Mexico; Cyathula 2 spp. native to Nay, Jal, Col, Gro, Mch, Oax, Chp, Pue, Ver, Tab, Cam, QR, and Yuc; Lagrezia 1 sp. Mexico endemic of Nay, Jal, Col, Mch, Gro, Oax, Chp, Tab, Cam, Yuc, and QR (most of genus is Madagascar); Pleuropetalum 2 Mexico & neoendemic spp. native to N+SE+SE Mexico, Ver. Betoideae: Aphanisma monospecific SW NAM endemic sp. native to BC, BCS?, and Mexican Pacific Is.; Beta 2 spp. intro to BC, BCS, Son, and Sin. Camphorosmoideae: Bassia 2 spp. intro to BC, BCS, Son?, and Sin?. Chenopodioideae: Atriplex ~30 spp. native through all of Mexico, including Mexican Pacific Is.; Blitum 4 spp., inc. 2 native and 2 intro to BCN, BCS, Son, Sin, Chi, NL, Tam, Dgo, Zac, SLP, Gto, Qro?, and Hgo?; Chenopodiastrum 1 sp. intro to BC, BCS, Son, Sin, Nay, Jal, Col, Mch, Gro, and Oax; Chenopodium ?? spp. native throughout all of Mexico; Dysphania 8 spp., inc. 7 native and 1 intro, found in all of Mexico and intro Mexican Pacific Is.; Extriplex 1 SW NAM endemic sp. native BCN and BCS?; Krascheninnikovia 1 sp. native to N Mexico in BC, BCS, Son, Sin, Chi, Coa, NL, Tam, Dgo, Zac, Qro, Ags, Gto, and SLP; Neomonolepis monospecific former SW NAM endemic native to BCN and BCS?; Oxybasis 4 3 subcosmopolitan spp., including 2 native (1 endemic) to Chi, Coa, NL, Tam, Dgo, Zac, Ags, SLP, Gto, Qro, Hgo, Mex, Cd Mex, Mor, Nay, Jal, Col, Gro, Mch, and Oax, and 2 spp. intro to BC, BCS, Son, and Sin. Corispermoideae: Corispermum 1 sp. native to Chi, Dgo, Zac, SLP, Hgo, Gto, Qro, NL, Tam, and Coa. Gomphrenoideae: Alternanthera ~6 spp. native throughout all of Mexico; Froelichia 6 former Americas endemic spp. native in all of Mexico, including 1 narrow endemic of BCS; Gomphrena ~16 spp. native to all of Mexico; Guilleminea 2 spp. native to N+SW+C Mexico; Iresine ~20 spp. native to all of Mexico, including many endemics; Pfaffia 1-3 neoendemic spp. native to S Mexico in Nay, Jal, Col, Mch, Gro, Oax, Chp, Cam, Tab, QR, and Yuc; Tidestromia 6 NAM & Caribbean endemic spp. native to BC, BCS, Son, Sin, Chi, Coa, NL, Tam, Dgo, Gto, Qro, SLP, Hgo, and Ags, inc. 1 narrow endemic of Coa. Polycnemoideae: Nitrophila 1 s.p native to NW Mexico. Salicornioideae: Allenrolfea 1 SW NAM endemic native to BCN, BCS, Son, Sin, Chi, NL, Tam, Dgo, and SLP (other 2 spp. endemic to Argentina); Arthroceras monospecific SW NAM endemic sp. native to BC, BCS, Son?, and Sin?; Salicornia 4?spp. native to most of Mexico except Pue, found in coastal & inland saline areas. Salsoideae: Salsola ~6 spp. intro to BC, BCS, Son, Sin, Chi, Coa, NL, Tam, Dgo, Zac, Gto, Qro, and Ags. Suaedoideae: Suaeda ~17 spp. native in all of Mexico including the Mexican Pacific Is.

Neotropical Amaranthaceae Genera Include:

Amaranthoideae: Achyranthes 2? spp. including 1 intro to CAM (exc. Costa Rica), Bahamas, Turks-Caicos, Aruba, Antilles (exc. Cayman Is.), SW Caribbean, Galapagos, and tropical SAM S to Peru, Bolivia, C+SE Brazil (exc. Ecuador) plus 1 native to Cuba, parts of the Antilles, Guatemeal, Colombia, and Venezuela; Amaranthus ~45 spp. native throughout N Caribbean, Antilles, SW Caribbean, Trinidad-Tobago, CAM, all of SAM, and Galapagos, and is intro to Bermuda and Juan Fernandez Is.; Celosia several spp. are native to Guatemala, El Salvador, Honduras, Nicaragua, Panama, Cuba, Hispaniola, Puerto Rico, Bahamas, Leeward & Windward Is., Venezuela, Colombia, Ecuador, Peru, and Brazil, and are intro in Belize, Cayman Is., Galapagos, Trinidad-Tobago, C Brazil, Bolivia, NE Argentina, and Paraguay; Chamissoa 3 Mexico & neoendemic spp. of Greater Antilles (exc. Cayman Is.), CAM S through tropical SAM to N Argentina exc. French Guiana and N Chile; Cyathula 3 spp. inc 2 native to CAM, Cuba, Hispaniola, Jamaica, Trinidad-Tobago, Colombia, Venezuela, Brazil, Ecuador, Peru, and Bolivia, and 1 intro to Puerto Rico, Leeward & Windward Is, Venezuelan Antilles, French Guiana, Suriname, Guyana; Hebanthodes monospecific endemic of Peru; Herbstia monospecific E SAM endemic of E+S Brazil, Paraguay, and NE Argentina; Lecosia 2 narrow endemic spp. of E Brazil (not in APG); Pleuropetalum 3 Mexico & neoendemic spp. native to CAM, Venezuela, Colombia, Ecuador, Peru, Bolivia, and Galapagos (1 narrow endemic of Galapagos). Betoideae: Beta 2 spp. intro to Honduras, Cuba, Cayman Islands, Dominican Republic, Puerto Rico, Ecuador, N & C Chile, Juan Fernández Is., Argentina, and Uruguay. Camphorosmoideae: Bassia 1-2 spp. intro to C. Chile and N. Argentina; Maireana 1 sp. intro in N Chile, Desventurados Is. Chenopodioideae: Atriplex ~20? spp. native to Honduras, Bermuda, Antilles (exc. Windward Is.), SW Caribbean, Venezuela, Colombia, Peru, Bolivia, Paraguay, Chile, Argentina, Uruguay, and S Brazil; Blitum 1 sp. intro to N Argentina; Chenopodiastrum 1 sp. intro to Honduras, Nicaragua, Panama, Bermuda, Bahamas, Turks-Caicos, Antilles (exc. Cayman Is, Windward Is.), Venezuela, Colombia, Ecuador, Peru, Bolivia, C+E Brazil, Chile, N Argentina, and Uruguay; Chenopodium ??spp. native to Honduras, Panama, Colombia, Ecuador, Peru, Chile, Argentina, Paraguay, and S Brazil, and intro C+E Brazil, Bermuda, Bahamas, Cuba, Dominican Republic, and Trinidad-Tobago; Dysphania 15 spp. native to CAM, Bermuda, all of SAM except Ecuador, French Guiana, Suriname, and Guyana where it is now intro, also intro in the Bahamas, Antilles, SW Caribbean, Trinidad-Tobago, Galapagos, and Juan Fernandez Is.; Holmbergia monospecific C SAM endemic of Bolivia, Paraguay, Uruguay, and N Argentina; Oxybasis 5 subcosmopolitan spp., including 3 native (2 endemic) of Peru, Bolivia, Chile, and NW+S Argentina and 2 spp. intro to Argentina; Spinacia 1 sp. intro to Hispaniola and Cayman Is. Gomphrenoideae: Alternanthera ~100? spp. native to Bermuda, Bahamas, Turks-Caicos, Aruba, Antilles, CAM, and all of SAM, including Galapagos, excluding S Chile; Froelichia 8 former Americas endemic spp. native to Guatemala, Honduras, Nicaragua, Cuba, Hispaniola, Jamaica, Colombia, Venezuela, Brazil, Ecuador, Galapagos, Peru, Bolivia, Paraguay, Uruguay, and N Argentina, including 4 narrow endemics of Galapagos (2), Para N Brazil, and Paraguay; Froelichiella monospecific endemic of C Brazil; Gomphrena ~90 spp. native to all of CAM and SAM, exc. S Chile, plus much of the Antilles region; Guilleminea 4 spp. native to Cuba, Hispaniola, Colombia, Ecuador, Peru, Bolivia, C Brazil, Paraguay, N Argentina, and Uruguay, inc. 2 endemics of Cuba and Uruguay; Iresine ~20 spp. Mexico + former neoendemic natives throughout all of CAM, the Bahamas, Antilles (exc. Aruba and Netherlands Antilles), and tropical SAM to N Argentina (exc. French Guiana and N Chile); Pedersenia 8 neoendemic spp. of Honduras S to Colombia, Venezuela, Ecuador, Peru, Paraguay, and N Brazil, plus Puerto Rico and the Windward Is.; Pfaffia 33 neoendemic spp. native to S Mexico, Haiti, Leeward & Windward Is., Venezuelan Antilles, Trinidad-Tobago, Costa Rica, and tropical SAM S to N Argentina (except N Chile); Pseudoplantago 2 SAM endemic spp. endemic to Venezuela (1) and S Brazil + NE Argentina (1); Quaternella 3 Brazil endemics exc. N Brazil; Tidestromia 1 S NAM & Caribbean endemic spp. native to the Dominican Republic; Xerosiphon 2 spp. endemic to most of Brazil except the south. Polycnemoideae: Nitrophila 2 SAM endemic spp. native to Argentina and N. Chile. Salicornioideae: Allenrolfea 2 Argentina endemic spp. native to all of Argentina; Heterostachys 2 neoendemic spp. native to Hispaniola, Colombia, Venezuela, Paraguay, and Argentina, including 1 narrow endemic of Buenos Aires, Mendoza & Rio Negro, Argentina, at the N limit of Patagonia; Mangleticornia monospecific endemic of Ecuador and Peru; Salicornia 5-6 spp. native in coastal and inland saline areas of the Bahamas, Turks and Caicos, Antilles (exc. Windward Is.), Colombia, Venezuela, Ecuador, Peru, Bolivia, Paraguay, S Brazil, Argentina, Chile, and Uruguay. Salsoideae: Salsola 2-3? spp. intro to Uruguay, C+S Chile, and Argentina. Suaedoideae: ~7-10 spp. native to the Bahamas, Turks and Caicos, Cuba, Hispaniola, Leeward Is., Peru, Bolivia, Chile, and Argentina, and extinct or extirpated Juan Fernandez Is.

Patagonia Amaranthaceae Genera Include:

Amaranthoideae: Amaranthus 4 spp. native to C+S Argentina, C+S Chile. Betoideae: Beta 1 sp. intro to C Chile and S Argentina. Camphorosmoideae: Bassia 1 sp. intro to C Chile and S Argentina. Chenopodioideae: Atriplex 1 sp. native throughout Patagonia and intro to Falkland Islands; Blitum 2 spp. inc. 1 intro and 1 endemic to Patagonia; Chenopodiastrum 1 sp. intro C+S Chile and S Argentina; Chenopodium 1 sp. native throughout Patagonia; Dysphania 1 sp. native to all of Patagonia; Oxybasis 4 subcosmopolitan spp., inc. 3 native to C+S Chile, S Argentina, and Falkland Islands, and 1 sp. intro to Patagonia. Gomphrenoideae: Alternanthera 1 sp. native just N of Patagonia in the mountains N of Bahia Blanca, S Argentina; Gomphrena 1-2 spp. native to S Argentina. Polycnemoideae: Nitrophila 1 SAM endemic sp. of S Argentina near N limit of Patagonia. Salicornioideae: Allenrolfea 1 Argentina endemic sp. native to S Argentina; Heterostachys 1 sp. is a narrow endemic of Buenos Aires, Mendoza & Rio Negro, Argentina, at the N limit of Patagonia; Salicornia 2? spp. native throughout all of Patagonia in salt marshes and lagoons. Salsoideae: Salsola 1? sp. intro throughout Patagonia. Suaedoideae: Suaeda 1 spp. native throughout Patagonia and the Falkland Islands.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as personal observations in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • Flora of North America (FNA) (1993+). https://floranorthamerica.org/Main_Page.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/
  • Stevens, P. F. (2001+). Angiosperm Phylogeny Website. Version 14, July 2017 [more or less continuously updated since]. http://www.mobot.org/MOBOT/research/APweb/
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout the fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


Identify the Anacardiaceae—Cashew/Sumac Family

Rhus typhina Staghorn Sumac plants with fruits (drupes). Learn how to identify the Anacardiaceae family with morphology photos.
Rhus typhina Staghorn Sumac plants with fruits (drupes).
Page Last Updated May 3, 2026.

Introduction to the Anacardiaceae Family

The Anacardiaceae family is part of the Sapindales order of core dicots. When you learn to identify the Anacardiaceae family, you will quickly learn its most well-known members in North America, the sumac (Rhus spp) and poison ivy (Toxicodendron spp). The sumac trees are often used ornamentally for their ease of growing and their pretty red drupes that persist all winter. Poison ivy is well known to anyone who has ever hiked in the forest since it causes very itchy contact dermatitis. 

There is an old saying, “Leaves of three, let it be,” since all of the Toxicodendron species that cause poison ivy rash have trifoliate leaflets. It is a good rule of thumb if you are unsure of your plant ID. But be aware that numerous innocuous species exist that also have trifoliate leaves, so do not assume it is always poison ivy. Poison ivy (shrub form) can also be mistaken for young oaks at times because the leaves often resemble those of the white oak.

Common Botanical Description

If you’re new to plant morphology, this is a perfect beginner’s description to learn to identify the Anacaradiaceae family, with no need to know any scientific jargon. For researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below.

Leaves and Stems: Mostly woody shrubs and trees or climbing vines with resin canals in their leaves, roots, and stems that often release a milky or resinous juice when damaged and that may be aromatic. Leaves are often arranged in a spiral pattern along the stem and may be simple (1 blade) or compound and made of multiple leaflets.

Flowers: These are usually quite small and occur in branched clusters called panicles. Most flowers have 5 sepals and 5 petals, but they may lack petals or they may all be more green and sepal-like. They often contain a ring-shaped nectary inside to attract pollinators.

Reproductive Features: These are quite diverse in the family, with bisexual flowers that contain both male and female parts or separate male and female flowers that may be on the same (monoecious) or separate plants (dioecious). There are usually 5-10 stamens and a superior ovary (sits above the point of petal attachment) with 1 or up to 6 styles (the tube that collects pollen) on top.

Fruits: The fruit is usually a fleshy drupe (think cherry but usually smaller) containing a fleshy outer part and a single hard pit. The fruits and sap of some species contain urushiol, the toxic oil responsible for the skin irritation of poison ivy (Toxicodendron species).

Uses of Anacardiaceae 

Some Anacardiaceae are edible plants, such as cashew nuts (Anacardium and the fleshy peduncle of the cashew apple), mango (Mangifera), Jamaica plum, hog-plum, imbu (a plum-like fruit from Spondias), and Amarula cream (from Sclerocarya typically made into a liqueur). Resins, oils, and lacquers are derived from Toxicodendron. Several non-native Anacardiaceae are cultivated in the Neotropics for their edible fruits, including Bouea macrophylla,, Harpephyllum caffrum, Mangifera indica, Schinus terebinthifolia, Sclerocarya birrea subspecies caffra, and Spondias dulcis. Many Anacardiacea species are notorious for their allergenic properties that often cause severe rashes, particularly Toxicodendron spp. (Poison Oak, Poison Ivies).

Morphology of Anacardiaceae in North America

Learn to identify the Anacardiaceae family with morphology photos

Some Anacardiaceae Species I have Covered So Far

Anacardioideae Subfamily

Mangifera indica - mango tree with fruits

Mangifera indica—Mango

A 10-20 m tall tree with long oblong leaves and large edible drupes on very long peduncles. Native to Asia but widely cultivated in southern North America, especially in Mexico.

Male flowers on a male Pistacia vera (pistacio) tree.

Pistacia vera—Pistacio

A small to medium-sized tree up to 10 m tall with deciduous, pinnate leaves 10–20 cm long. Flowers have no petals and male and female flowers occur on separate trees (dioecious). Fruits are a drupe containing the elongated pistacio “nut.” These flowers are male.

Rhus aromatica Fragrant Sumac plant with hairy drupes (fruits) that are common among the Anacardiaceae family.

Rhus aromatica—Fragrant Sumac

A somewhat lemon-scented shrub with trifoliate lobed leaves and hairy red drupes. Dioecious with small inconspicuous flowers. Native throughout the USA, Mexico, and southeastern Canada.

Rhus copallinum plant with flowers - note winged rachis and compound leaves; compound leaves are fairly common in the Anacardiacea family.

Rhus copallinum—Shining Sumac

A tall shrub with glossy odd-pinnate leaves that have a conspicuously winged rachis or petiolules (visible in the photo). Native to eastern North America, often cultivated.

Rhus glabra Smooth Sumac infructescence of drupes.

Rhus glabra—Smooth Sumac

A tall shrub with hairless branches and odd-pinnate leaves with 11 – 31 acuminate, serrated leaflets. Produces large dense panicles of red drupes like shown in the photo. Native to eastern North America with scattered populations in western North America.

Rhus microphylla Littleleaf Sumac tiny compound leaves and drupes, characteristic of the Anacardiaceae family..

Rhus microphylla—Littleleaf Sumac

A desert shrub that often flowers before its tiny, odd-pinnate, glossy green, leathery, hairy leaves emerge. Fruits are clusters of small hairy drupes. Native to the southwestern USA plus northern and central Mexico.

Rhus trilobata Skunkbush Sumac trifoliate leaves

Rhus trilobata—Skunkbush Sumac

Despite its pungent, bitter scent that gives it a bad rap, it is not nearly as unpleasant as a skunk. It is a shrub with trifoliate lobed leaves (shown in the pic) and flowers in catkins, and its fruits are hairy, sticky red drupes. Native to western North America, including western Mexico.

Rhus typhina Staghorn Sumac plants with fruits (drupes).

Rhus typhina—Staghorn Sumac

A large shrub with odd-pinnate leaves with 9 – 31 serrated leaflets; stems and petioles are densely red-hairy (unlike R. glabra). Native to northeastern USA and southeastern Canada.

Toxicodendron radians Eastern Poison Ivy. Plant with flowers growing on a tree.

Toxicodendron radicans—Eastern Poison Ivy

A climbing vine (sometimes a shrub) with woody stems & trifoliate shiny leaves that are usually not serrated. Flowers are small, greenish, and inconspicuous. The fruit is a gray-white drupe. Causes contact dermatitis. Native to the eastern USA and Canada.

Toxicodendron rydbergii, Western Poison Ivy. Shrubby plant with immature drupes.

Toxicodendron rydbergii—Western Poison Ivy

A shrub with opposite trifoliate leaves that are often asymmetrical. The fruits are white, yellow, or brown drupes that are ribbed (green like in the photo when still immature). Native throughout the USA and Canada but mostly found in the West.

Scientific Botanical Description of the Anacardiaceae

Habit & Leaf Form of the Anacardiaceae

Trees or shrubs, self-supporting or climbing, sometimes with a milky juice that may be resinous or laticiferous. Leaves are evergreen or deciduous and alternate spiral except in Bouea, where they are opposite, and may be aromatic when resinous but are otherwise odorless. They are simple or compound ternate, trifoliate, imparipinnate, or rarely paripinnate or bipinnate. Primary venation is pinnate or rarely palmate. Secondary venation is eucamptodromous, brochidodromous, craspedodromous, cladodromous, or rarely reticulodromous. If present, cladodromous venation is diagnostic for the Anacardiaceae family. Leaves are exstipulate. Lamina margins are entire. Domatia occur in 8 genera as pits, pockets, or hair tufts. Resin canals located in the inner fibrous bark and pith of the stems, roots, and leaves are characteristic of this family. Tannin sacs are very common.

Flowers of the Anacardiaceae

Plants can be hermaphroditic, monoecious, dioecious, gynodioecious, or polygamomonoecious. Pollination when heterantherous is entomophilous. Flowers are aggregated in racemose panicles. Flowers are small, regular, usually 5-merous, and either tetracyclic or pentacyclic. Free hypanthium is either absent or present but short. A hypogynous disk is present, intrastaminal and annular. The perianth has a distinct calyx and corolla or may be sepaline. The perianth has 3–5 or 6–10 parts in 1 or 2 whorls and is isomerous. Calyx has 3–5 parts in 1 whorl, is basally connate, often with cleft sepals, and lobes are shorter to longer than the tube. The calyx is regular and imbricate. The corolla, when present, has 3–5 parts in 1 whorl, is usually free but rarely can be connate basally, the lobes are longer than the tube, and it is imbricate.

Androecium of the Anacardiaceae

The androecium has 5–10(11–12) parts. Androecial members are free of the perianth and may be free of one another or sometimes coherent 1 adelphous (filaments basally connate). The members may be all equal or unequal and can be either 1 or 2 whorls. There may be 0-9 staminodes. There are 5–10 (1–12) stamens that are oppositisepalous, inserted at the base of the hypogynous disc, and are usually equal in number or twice as many as the petals. Anthers are usually dorsifixed or sometimes basifixed (Spondias), they are versatile, dehiscing via longitudinal slits, introrse, and tetrasporangiate.

Gynoecium of the Anacardiaceae

The gynoecium is usually 5-carpeled but can be 1–6 carpeled, and the pistil is 1–5 celled. The gynoecium is synovarious or synstylovarious, or rarely semicarpous. It is usually superior or sometimes partly inferior. The ovary is 1–5-locular. There is usually one style, but there may be 3–6, and in Buchanania, it has up to five styles from sterile carpels. There are 1-5 stigmas of the wet type, non-papillate, and Group IV type. Placentation, when unilocular, is parietal or basal, and when bi- or plurilocular, it is basal. There is 1 ovule per locule; it is pendulous or ascending, has either a ventral or dorsal raphe, is non-arillate, anatropous, unitegmic or bitegmic, and crassinucellate. 

Fruit of the Anacardiaceae

The fruit is usually a fleshy drupe, but sometimes it is an indehiscent non-fleshy dry fruit. The drupes have one stone, rarely open at maturity, and sometimes contain urushiol, a toxic irritant found in poison ivy and other species. Seeds are non-endospermic.

Taxonomy of the Anacardiaceae Family

The Anacardiaceae have 873 species in 80 genera of the Sapindales order of core eudicots. There are two accepted subfamilies with several dissident genera in their own groups or unplaced; here I have put them all as “unplaced in a subfamily.” As APG updates these, this record will be updated to reflect any changes.

  1. Anacardioideae – Trees or shrubs with black or colored resinous exudate and crystals are present in the xylem. Leaflets are not articulated, margins are usually entire, and the base of the petiole is often swollen. Flowers are 5(-7)-merous, the calyx is more or less connate basally, the gynoecium is typically 3-carpeled, and the stigma is dry and capitate or lobed. The fruit is a drupe that is layered with a crystalliferous endocarp.  
  2. Spondiadoideae – Deciduous trees or shrubs that typically cause contact dermatitis. Flower pedicels are often articulated, the gynoecium is 4-5(3) carpelled, and the stigma is only slightly expanded. Fruit is usually more than two-seeded (but sometimes only one), its pericarp may or may not have a lacuna, and the inner mesocarp is made of encircling fibers.
  3. Unplaced” includes the Sclerocarya complex of multiple genera, plus Buchanania and Campnosperma, which are grouped together, and some completely unplaced ones, including Attilaea, Haplospondias, Koordersiodendron, and Pentaspadon.

Genera of the Anacardiaceae:

Anacardioideae: Abrahamia (34), Actinocheita (1), Amphipterygium (5), Anacardium (13), Androtium (1), Apterokarpos (1?), Astronium (11), Baronia (1), Blepharocarya (2), Bonetiella (1), Bouea (3), Campylopetalum (1), Cardenasiodendron (1), Comocladia (27), Cotinus (7), Dobinea (2), Drimycarpus (4), Euroschinus (9), Faguetia (1), Fegimanra (3), Gluta (35), Haplorhus (1), Heeria (1), Holigarna (9), Laurophyllus (1), Lithraea (3), Loxopterygium (4), Loxostylis (1), Malosma (1), Mangifera (64), Mauria (15), Melanochyla (21), Melanococca (?), Metopium (4), Micronychia (10), Mosquitoxylum (1), Myracrodruon (2), Nothopegia (10), Ochoterenaea (1), Orthopterygium (1), Ozoroa (45), Pachycormus (1), Parishia (8), Pegia (2), Pistacia (12), Protorhus (3), Pseudosmodingium (5), Rhodosphaera (1), Rhus (51), Schinopsis (7), Schinus (40), Searsia (110), Semecarpus (87), Smodingium (1), Sorindeia (10), Swintonia (13), Thyrsodium (6), Toxicodendron (28), Trichoscypha (32).

Spondiadoideae: Allospondias (2), Antrocaryon (5), Choerospondias (1), Cyrtocarpa (4), Dracontomelon (9), Haematostaphis (1), Harpephyllum (1), Lannea (36), Operculicarya (9), Pleiogynium (3), Poupartia (7), Poupartiopsis (1), Pseudospondias (2), Sclerocarya (2), Solenocarpus (2), Spondias (18), Tapirira (9), Tumultivenia (1), Uniostium (1).

Unplaced in a subfamily: Attilaea (1), Buchanania (26), Campnosperma (14), Haplospondias (?), Koordersiodendron (1), Pentaspadon (6).

Key Differences From Similar Families

The Anacardiaceae differ from the similar Burseraceae by having usually alternate or sub-opposite leaves or leaflets compared to the almost always opposite leaflets of Burseraceae.

Distribution of Anacardiaceae

The Anacardiaceae family is mainly tropical to subtropical, with a few important genera found in temperate North America. They are widespread pantropically and also in the Mediterranean, eastern Asia, and the warm Americas.

Distribution of Anacardiaceae in the Americas

Canadian Genera Include:

Anacardioideae: Cotinus 1 sp. intro to ON; Rhus 5 spp. native to all S provinces except NL; Toxicodendron 3 spp. native in all S provinces except NL, and including YT. 

USA Genera Include:

Anacardioideae: Cotinus 2 spp. including 1 C+E USA endemic and 1 intro to UT, TX, OK, AR, IL, MO, AL, GA, TN, KY, OH, PA, NY, MD, DE, NJ, CT, MA, VT; Lithraea 1 sp. intro CA; Malosma monospecific S NAM endemic native CA; Mangifera 1 sp. intro to FL; Metopium 1 Gulf + Caribbean endemic sp. native to FL; Pistacia 3 spp, inc native in TX and 2 intro to CA, UT, OK, AL, GA, and VA; Rhus 16 spp. native and intro in the entire US, inc. HI; Schinus 5 former SAM endemic spp. intro to CA, TX, AL, FL, and HI; Searsia 1 sp. intro to CA and AZ; Sorindeia 1 sp. intro to FL; Toxicondrendon 5 spp. native in all the USA. Spondiadoideae: Spondias 1 sp. intro to FL. 

Mexico Genera Include:

Anacardioideae: Actinocheita monospecific Mesoamerican endemic of Honduras and Nay, Jal, Col, Mch, Gro, Oax, and Pue; Amphipterygium 5 Mexico & CAM endemic spp. native to most of Mexico except NW Mexico, including 2 endemics of SW Mexico (1), Dgo + Jal (1); Anacardium 1 sp. intro to Nay, Jal, Col, Mch, Gro, and Oax; Astronium 2 Mexico & neoendemic spp. native most of Mexico exc. central and NW, inc. 1 narrow endemic of Ver; Bonetiella monospecific endemic of NE Mexico; Comocladia 5 Mexico & CAM endemic spp. native in all of Mexico, inc. 3 endemic to Mexico; Cotinus 3 spp. native to Chi, Coa, NL, Tam, Dgo, Zac, Ags, Gto, Qro, and Hgo, including 2 endemics; Lithraea 1 sp. intro in Pue; Malosma monospecific S NAM endemic native BCN, BCS, Son, Sin, Chi, Coa, NL, Tam, SLP, Dgo, Zac, and Mexican Pacific Islands; Mangifera 1? sp. intro SW+SE+C Mexico; Metopium 1 Gulf & Caribbean endemic spp. native to Ver, Chp, Cam, Tab, Yuc, and QR; Mosquitoxylum monospecific N Neo endemic sp. native to S Mexico in Nay, Jal, Col, Mch, Gro, Oax, Chp, Ver, Tab, Cam, Yuc, and QR; Pachycormus monospecific endemic of BCN, BCS; Pistacia 2 sp. inc 1 native to most of Mexico except BC, BCS, Son, and Sin and 1 intro to N Mexico; Pseudosmodingium 5 spp. endemic to most of Mexico exc. SE Mexico; Rhus 15+ spp. native throughout Mexico including the Mexican Pacific Is.; Schinus 2 former SAM endemic spp. intro N and SW Mexico; Toxicodendron 4 spp. native to all of Mexico. Spondiadoideae: Spondias 3 spp. native to all of Mexico; Tapirira 2 Mexico & neoendemic spp. native SW Mexico, Pue, Ver, Cam, Tab, Yuc, and QR, including 1 endemic of Oax + Ver. Unplaced: Attilaea monospecific endemic of QR, Yuc, and Guatemala; Cyrtocarpa 3 Mexico & N neoendemic spp. native to most of Mexico exc. Ver, inc. 2 endemics.

Neotropical Genera Include:

Anacardioideae: Actinocheita monospecific Mesoamerican endemic to SC Mexico and Honduras; Amphipterygium 3 of 5 Mexico & CAM endemic spp. native to Guatemala, Honduras, Nicaragua, and NW Costa Rica; Anacardium 13 spp of former neoendemic genera native from Honduras S to Peru, Bolivia, Paraguay, S Brazil, and also in Cuba and intro to the rest of Antilles (exc. Netherlands Antilles & Aruba), Belize, and El Salvador; Apterokarpos (~Loxopterygium?) monospecific endemic to the Caatinga of NE Brazil?; Astronium 10 Mexico & neoendemic spp. native to CAM S through tropical SAM to Peru, Bolivia, NE Argentina, S Brazil, including 5 narrow endemics of Brazil (4) and Trinidad-Tobago; Cardenasiodendron monospecific endemic to Bolivia; Comocladia 24 Mexico & CAM endemic spp. native to Guatemala, Belize, Greater Antilles, and Leeward & Windward Is. with 15 single-island endemics of Jamaica (6), Hispaniola (5), Cuba (2), Dominican Republic, Haiti, and 1 extinct sp. of the Windward Is.; Haplorhus monospecific endemic to the dry inter- Andean valleys of Peru to N Chile; Lithraea 3 SAM endemic spp. of C+E+S Brazil, Bolivia, Paraguay, N Argentina, Uruguay, and C Chile, inc. 1 narrow endemic of C Chile; Loxopterygium 4 SAM endemic spp. of Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, NW Argentina, and NE Brazil; Mangifera 1 sp. intro to Guatemala, Belize, Honduras, El Salvador, Costa Rica, Galapagos, Antilles (excluding Netherlands Antilles, Aruba), Venezuela, Ecuador, Peru, Paraguay, and S Brazil; Mauria 15 neoendemic spp. from Costa Rica S to Colombia, Venezuela, Ecuador, Peru, Bolivia, and NW Argentina; Metopium 4 Gulf & Caribbean endemic spp. native to Guatemala, Belize, Honduras, Bahamas, Turks and Caicos, Antilles (except Leeward Is. & Venezuelan Antilles), and the SW Caribbean; Mosquitoxylum monospecific N neoendemic of Jamaica, S Mexico, Guatemala, Belize, Honduras, Nicaragua, Panama, and Ecuador; Myracrodruon 2 SAM endemic spp. of Brazil, Bolivia, Paraguay, and N Argentina; Ochoterenaea monospecific N SAM endemic to Andean Venezuela and Colombia, and Bolivia; Orthopterygium monospecific endemic to W Peru; Pistacia 1 sp. native to Guatemala and Honduras; Rhus 1-3? spp. native to Cuba, Bahamas, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica, and intro to Trinidad-Tobago; Schinopsis 7 SAM endemic spp. of Peru, Bolivia, C+E Brazil, Paraguay, and N Argentina; Schinus 40 former SAM endemic spp. native to Peru, Bolivia, and C+E+S Brazil and all south (absent Amazonia), and intro to Ecuador, Colombia, Bermuda, the Bahamas, Turks and Caicos, Cuba, the Dominican Republic, Puerto Rico, Trinidad-Tobago, Venezuelan Antilles, Guatemala, and El Salvador; Semecarpus 1 sp. intro to Trinidad-Tobago; Thyrsodium 6 N SAM endemic spp. of Colombia, Peru, Bolivia, Venezuela, French Guiana, Guyana, Suriname, and N+C+E Brazil; Toxicodendron 3 spp. including 2 native to Bermuda, the Bahamas, Guatemala, El Salvador, Honduras, Nicaragua, Panama, Colombia, Venezuela, Ecuador, Peru, and Bolivia and 1 sp. intro to Cuba. Spondiadoideae: Antrocaryon 1 sp. endemic to Colombia and Amazonian N+C Brazil (rest of genera are tropical Africa); Dracontomelon 1 sp. intro to Trinidad-Tobago; Spondias 12 spp. inc 11 natives of Antilles, CAM S through tropical SAM S to Peru, Bolivia, and C+SE Brazil, and 1 sp. intro to Guatemala, Belize, Nicaragua, Panama, the Greater Antilles, Colombia, Venezuela, Guyana, French Guiana, N+SE Brazil, Ecuador, and Peru, including 6 narrow endemics of E Brazil; Tapirira 8 Mexico & neoendemic spp. native from CAM S through tropical SAM to Peru, Bolivia, Paraguay, and S Brazil, including 3 narrow endemics of Costa Rica, Colombia, and French Guiana; Tumultivenia monospecific endemic of E + WC Brazil; Uniostium monospecific endemic of Colombia, Venezuela, Guyana, and the Netherlands Antilles. Unplaced: Attilaea monospecific endemic of SE Mexico and Guatemala; Campnosperma 2 neoendemic spp. from Honduras S to Colombia, Venezuela, Ecuador, Peru, and Amazonian N Brazil (most of the genus is in Madagascar, Asia, and Australasia); Cyrtocarpa 2 Mexico + neoendemic spp. native to Colombia, Venezuela, Guyana, N+C+E Brazil, and the Netherlands Antilles.

Patagonia Genera Include:

Anacardioideae: Lithraea 1 SAM endemic spp. native to Bio Bio to Los Lagos in C. Chile; Schinus 6 former SAM endemic spp. native to the entire region. 

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, along with my own observations in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Flora of North America (1993+). https://floranorthamerica.org/Main_Page.
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx Accessed 2020–current.
  • Neotropikey: Milliken, W., Klitgård, B. & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/ Retrieved Winter 2020-current.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Acanthaceae Family

Tetramerium nervosum inflorescence with showy bracts and flower. Learn how to identify the Acanthaceae family with morphology photos!
Tetramerium nervosum inflorescence with showy leaf-like hairy bracts and irregular flower.
Page Last Updated May 2, 2026

Introduction to the Acanthaceae Family

When you learn how to identify the Acanthaceae family, it helps to understand that it is part of the Lamiales order of core eudicots and, as such, is closely related to the Lamiaceae, or Mint family, with which it shares several characteristics. The Acanthaceae are widespread in tropical and subtropical regions, but the USA has quite a few, and they can even be found in temperate Canada. Acanthaceae are usually herbs or shrubs and typically have zygomorphic flowers in white, pink, blue, and shades of purple. Several species are used ornamentally and medicinally. In the field, if you are injured, you can crush the leaves of most Acanthaceae and apply them directly to the wounds to aid in healing. 

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to learn about the Acanthaceae family, with no need to know any scientific jargon. Below is additional information on uses and morphology, as well as pictures to help identify family members and individual species found in North America. But for researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below the images in addition to genus-level distribution data.

Leaves and Stems of the Acanthaceae: Most members are tropical herbs, shrubs, or climbing vines that twine in a clockwise direction. Leaves are mostly simple (not compound) and arranged in opposite pairs along the stem, which often has swollen joints (nodes). Many species have mineral crystals (cystoliths) visible in their leaves and are covered with small hairs.

Flowers of the Acanthaceae: Flowers may be small or very large and showy and are typically irregular in shape, often forming funnel-shaped flowers or two-lipped flowers where the petals are joined at the base into a tube. Flowers are almost always surrounded by large, showy, leaf-like structures called bracts, which are very characteristic of the family (excluding the Thunbergioideae subfamily, which lacks the showy bracts).

Reproductive Features of the Acanthaceae: Most flowers contain both male and female parts (hermaphrodite or bisexual), with 2-4 stamens (male parts) attached inside the petal tube, often accompanied by smaller, sterile “fake stamens” (staminodes). The superior ovary (attached above the point of petal attachment) has a long, thread-like tube (style) on top to catch pollen and a nectary at the base to attract pollinators.

Fruits of the Acanthaceae: Fruits are almost always a dry, 2-chambered capsule that usually splits explosively when dry to scatter its seeds via small, hooked stalks (retinacula) attached to the seeds that aid in their ejection from the fruit.

Uses of Acanthaceae 

The leaves of many Acanthaceae are used externally for wounds. Research has shown that the family has antifungal, anti-inflammatory, antipyretic, antioxidant, insecticidal, immunomodulatory, anti-platelet aggregation, and antiviral potential. More research is currently underway. Notable ornamentals include bear’s-breech (Acanthus mollis), clock vine (Thunbergia), shrimp plant (Justicia brandegeana), and caricature plant (Graptophyllum pictum).

Morphology of Acanthaceae in North America

Learn to identify the Acanthaceae family with Morphology photos of the Acanthaceae Family
Morphology of the Acanthaceae Family

Acanthaceae Species I have Covered So Far in North America

Acanthoideae Subfamily

Acanthus mollis flowers and bracts; these leafy bracts are very typical of the Acanthaceae family.

Acanthus mollis—Bear’s Breeches

A clump-forming perennial herb with tuberous roots, a basal rosette of deeply lobed leaves, and tall spikes of pinkish or purplish flowers. Native to the Mediterranean, often cultivated in North America for its leaves and flowers.

Carlowrightia arizonica flower and leaves

Carlowrightia arizonica – Arizona Wrightwort

Heavily branched subshrubs are 10–30 cm tall, or sometimes taller. Leaves are simple, opposite, and variable in size and shape but approximately lanceolate. Flowers are narrowly triangular with 2 lateral petal lobes, 2 upper lobes fused together with a yellow spot and purple streaks, and a keeled bottom lobe. Native to Texas, Arizona, California, and northern Mexico.

Dicliptera resupinata two-lipped flowers, a common floral form in the Acanthaceae family.

Dicliptera resupinata—Arizona Foldwing

Erect herb up to 60 cm tall with heavily branching pubescent stems; leaves are lanceolate, 2 – 6 cm long. Flowers are two-lipped, light purple with small dark purple lines near the center, and surrounded by two heart-shaped bracts. Native to southern AZ and NM, United States, and northwest Mexico.

Justicia pilosella plant with flower, Carlsbad National Park

Justicia pilosella—Hairy Tubetongue

Herbaceous perennial to 30 cm tall with opposite leaves that may be pubescent or glabrous but with ciliate margins and zygomorphic sessile flowers with a long, white, pubescent tubular corolla with 4 pink or purple lobes, 3 of which bend outwards strongly. Narrowly endemic to southern TX and southern NM in the USA, plus arid northeastern Mexico.

Ruellia blechnum Green Shrimp Plant showing flower and leafy green bracts so typical of the Acanthaceae

Ruellia blechum—Green Shrimp Plant

Short herbaceous erect or clambering perennial with opposite ovate leaves and a conspicuously pyramidal-shaped inflorescence with often pilose pyramidal bracts from Paco’s Nature Reserve, Sinaloa, Mexico; conspicuous inflorescence spike with pyramidal bracts. Native to Mexico, Central & South America.

Ruellia caroliniensis plant with flower Mammath Cave National Park

Ruellia caroliniensis – Carolina Wild Petunia

Unbranched herbaceous perennial to 1 m with opposite, oval leaves and sessile axillary clusters of 2 – 4 flowers with usually only 1 – 2 open at a time; leaves and flowers are crowded together at the top of the plant. Flowers are light purple to pinkish with a slender corolla tube and 5 petaloid lobes and long-pointed calyx lobes. Native throughout the eastern USA.

Ruellia humilis showy flowers

Ruellia humilis – Wild Petunia

Perennial herb up to 60 cm tall with tubular, bell-shaped flowers with a long, skinny white tube and five shallow rounded lavender to lilac-colored petals; singly or in clusters in upper axils, not crowded at the top of the plant. Native to the eastern and central USA.

Ruellia nudiflora plant with showy bractless flowers

Ruellia nudiflora – Violet Wild Petunia

An erect, 30 – 60 cm tall perennial with few branches and opposite gray-green leaves 5 – 12 cm long with wavy-toothed margins on short petioles. Terminal inflorescence of lavender to purple trumpet-shaped flowers (bractless, hence “nudiflora”) with inserted stamens. Native to the southern USA (AZ east to AL), Mexico, and Central America.

Ruellia strepens plant with flower in the Tallgrass Prairies

Ruellia strepens – Smooth Wild Petunia

An herbaceous perennial plant 0.5 – 1 m tall, sometimes branching with hairless or sparsely hairy stems; opposite leaves up to 13 cm long, lanceolate to ovate with smooth or slightly undulate margins and mostly hairless surfaces. Nearly sessile flowers in clusters of 1 – 3 in upper axils. Native to the south-central & southeast USA.

Tetramerium nervosum inflorescence with showy bracts and flower also typical of the Acanthaceae.

Tetramerium nervosum – Hairy Fournwort

Subshrub to 30 cm tall with opposite lanceolate leaves. Flowers in conspicuous pilose leafy bracts on hairy, 4-sided spike inflorescences; tubular with an upper lobe with a violet patch and a yellow base, 2 side lobes, and a keel-like lower lobe. Native to AZ and TX, USA, and south through Mexico and Central America to Venezuela.

Nelsonioideae Subfamily

Elytraria imbricata purle scaly stem plant with flowers from Sinaloa, Mexico

Elytraria imbricata – Purple Scaly Stem

Weedy subshrub with linear leaves crowded at the top of the plant that may be basal and stemless or up to 60 cm tall. Inflorescences with very appressed bracts, blue flowers. Native from the southern USA to northern Argentina, mostly in dry tropical forests.

Thunbergioideae Subfamily

Thunbergia grandiflora showy flower closeup

Thunbergia grandiflora – Blue Trumpet Vine

A twining evergreen vine 2 – 2.5 m long, with large, heart-shaped, bright green leaves and large, showy, lavender-blue, trumpet-shaped flowers up to 7 cm across with a yellow or white throat. Native throughout Southeast Asia. Often cultivated as an annual in North America.

Scientific Description of the Acanthaceae

This section is for researchers or others wanting a more in-depth scientific description to learn how to identify the Acanthaceae family.

Habit & Leaf Form of the Acanthaceae

Mostly tropical herbs, shrubs, or twining vines, with some epiphytes or rarely trees (with pneumatophores and sometimes stilt roots in Avicennia). Leaves are usually well developed but sometimes are much reduced, or occasionally plants are aphyllous switch plants. Leaves may be heterophyllous or isophyllous and are often swollen at the nodes. Branches are terete to angular in cross-section. Herbs are annual or perennial, with or without a basal aggregation of leaves. Most plants are self-supporting but may also be epiphytic or climbing as stem twiners, root climbers, or scramblers with twiners twining clockwise. Tree forms are always leptocaul. Plants may be hydrophytic, helophytic (including a few mangroves), mesophytic (many from damp tropical forests), or xerophytic. Leaves are arranged opposite distichous or decussate (rarely alternate or whorled), are simple, and may or may not be gland-dotted. The lamina is dissected or entire, pinnately veined, and cross-venulate. Leaves have no stipules. Lamina margins are entire, crenate, serrate, or dentate and may be flat, revolute, or involute. The leaf lamina is dorsiventral (sometimes incomplete) or bifacial (isobilateral in several genera), with or without epidermal salt glands, and the abaxial surface may be papillose. Stomata are almost always diacytic, but in Lepidagathis, they are paracytic; they are mainly confined to the abaxial surface. Diverse hairs are present, including eglandular, unicellular, uniseriate, glandular (may always be glandular in Nelsonioideae), and multicellular branched or simple. Cystoliths are very often present as streaks in the lamina.

Flowers of the Acanthaceae

Plants are always hermaphrodites. Pollination is entomophilous and may be conspicuously specialized. Flowers are aggregated in racemes, cymes, or verticils or often in dichasial cymes that become monochasial but are often condensed in the leaf axils; sometimes they are pseudanthial. Flowers are both bracteate and bracteolate, and they are often large and showy. In Thunbergioideae, bracts are absent, but bracteoles are present. In Nelsonioideae, bracts are present, but bracteoles are absent. Flowers are usually more or less zygomorphic, sometimes actinomorphic. Flowers are 4 or 5 merous and are tetracyclic. Free hypanthium is absent; hypogynous disk is present.

The perianth has a distinct calyx and corolla with 8(6–7) or 10 parts in 2 whorls and may be isomerous or anisomerous. The calyx has 4 (3) or 5 parts in 1 whorl, is connate, and is variously entire, lobulate, or blunt-lobed, with the lobes shorter or longer than the tube; it may be imbricate, valvate, contorted, or open in bud. When the calyx has 5 parts, the median member is free and located posteriorly. The corolla has 4 or 5 parts, or 3 parts when the upper lip is suppressed. It is 1 whorled and connate, at least basally. The corolla tube is sometimes adaxially deeply split, where the upper lip of the corolla is cut away almost to the base of the tube, making it two-lipped. The corolla is imbricate, ascending, cochlear, quincuncial, or contorted (left or right), or sometimes with open aestivation.  

Androecium of the Acanthaceae

The androecium has 2 or 4 (5) members that are adnate and usually inserted on the corolla tube. They may be equal or unequal, free of one another, or coherent 2 adelphous (partially connate in pairs), and 1 whorled. The androecium usually includes 1-3 staminodes in the same series as the fertile stamens, or may be made of exclusively fertile stamens (rarely Ruellia). The stamens are usually exserted, and are usually didynamous and may or may not be hairy or spurred. Stamens are always oppositisepalous. Anthers may be separate from one another or connivent; are dorsifixed (often with one lobe reduced or abortive) or adnate, dehiscing via longitudinal slits; are unilocular to bilocular, are tetrasporangiate, and may or may not be appendaged (often with a long connective).

Gynoecium of the Acanthaceae

The gynoecium is 2-carpelled, and the pistil is 2-celled. The gynoecium is synstylovarious to syncarpous and superior. The ovary is 2-locular and sessile. The gynoecium is median. There is one terminal style that is attenuate from the ovary and usually filiform and much longer than the ovary. There are two stigmas with the posterior often smaller: they are the dry type (wet in Thunbergioideae), non-papillate (papillate in Thunbergioideae) and the Group II type. Placentation is axile with 2-50 ovules per locule that are non-arillate or arillate (occasionally with a funicular aril), anatropous to campylotropous, and unitegmic.

Fruit of the Acanthaceae

Fruit is a non-fleshy, 2-chambered loculicidal capsule dehiscing somewhat explosively or occasionally an achene (Avicennia). In most species, the seeds are attached to a small hooked stalk that helps eject them from the capsule. Seeds are non-endospermic, borne on retinacula, may be conspicuously hairy, and may contain starch.

Taxonomy of Acanthaceae

The Acanthaceae family has 4605 species in 191 genera. It is part of the Lamiales order of the Core Eudicots. There are four subfamilies of the Acanthaceae that are recognized:

  1. Acanthoideae – herbs or sometimes shrubs, with petiole bundles arranged in a circle. Corolla often has the abaxial lobe outside others when in bud. Anthers are sagittate, or the thecae are displaced and not opposite, sometimes with one theca more or less reduced. Stigma is dry and typically bifid. Capsules are obovoid and explosive; seeds are flattened and borne on hook-like hardened funicles. 
  2. Avicennioideae – trees with pneumatophores, sometimes stilt roots. The leaf lamina is thick with salt glands on both sides, club-shaped hairs, and colleters. Flowers are 4(-6) merous and quincuncial, with nectar glands on the inside of the tube. Stamens are equal and alternate with the corolla. Stigma has two blunt lobes. Fruit is an achene with large seeds that are more or less viviparous
  3. Nelsonioideae – herbs with glandular hairs. The inflorescence may be terminal or axillary, bracts are spiral, and bracteoles are sometimes absent. Corolla with descending cochleate aestivation with the adaxial lobes outside the others. There are 2 stamens with variable anthers with thecae that may or may not be separate. The stigma is broadly lobed. 
  4. Thunbergioideae – twining vines, sometimes erect. Petiole bundles are arcuate or annular with wing bundles. Leaf lamina vernation is strongly curved. Inflorescence is axillary flowers or fasciculate. They have no bracts but have very large bracteoles that may or may not be connate. Anthers have lignified unicellular hairs, are sagittate, dehisce by pores or sometimes slits, and have an elongated connective. The stigma is small, wet, sub-bilobed to trumpet-shaped, and has broad and often unequal papillate lobes.

Genera:

Acanthoideae: Acanthopale (12), Acanthopsis (20), Acanthostelma (?), Acanthus (30), Achyrocalyx (?), Aechmanthera (?), Afrofittonia (1), Ambongia (1), Ancistranthus (1), Andrographis (27), Angkalanthus (1), Anisacanthus (11), Anisosepalum (3), Anisotes (29), Anthacanthus (?), Aphanosperma (1), Aphelandra (212), Aphelandrella (1?), Ascotheca (1), Asystasia (61), Asystasiella (?), Ballochia (3), Barleria (307), Barleriola (4), Blechum (1), Blepharis (128), Borneacanthus (6), Boutonia (1), Brachystephanus (21), Bravaisia (3), Brillantaisia (14), Brunoniella (6), Buceragenia (2? unplaced), Calacanthus (1), Calycacanthus (2), Camarotea (1), Carlowrightia (27), Celerina (1), Centrilla (?), Cephalacanthus (1), Chaetacanthus (1 unplaced?), Chalarothyrsus (1), Chamaeranthemum (4), Champluviera (2), Chileranthemum (3), Chlamydacanthus (3), Chlamydocardia (2), Chlamydostachya (1?), Chroesthes (4), Clinacanthus (3), Clistax (3), Codonacanthus (3), Cosmianthemum (14), Crabbea (14), Crossandra (54), Crossandrella (3), Cuenotia (1 not in APG), Cyclacanthus (2), Cylindrosolenium (1?), Cyphacanthus (1), Cystacanthus (1 unplaced?), Danguya (?), Dasytropis (1), Dianthera (41 when not inc. in Justicia), Dichazothece (1), Dicladanthera (2), Dicliptera (242), Dinteracanthus (5), Diotacanthus (?), Dischistocalyx (12), Duosperma (26), Dyschoriste (98), Ecbolium (22), Echinacanthus (4), Encephalosphaera (3?), Eranthemum (22), Eremomastax (1), Filetia (9), Fittonia (2), Forcipella (6), Geissomeria (3 unplaced?), Glossochilus (1), Golaea (?), Graphandra (1), Graptophyllum (15), Gymnostachyum (53), Gypsacanthus (1), Habracanthus (?), Haplanthodes (4), Harpochilus (3), Hemigraphis (31), Henrya (3), Herpetacanthus (21), Heteradelphia (2), Holographis (18), Hoverdenia (1), Hulemacanthus (2), Hygrophila (76), Hypoestes (139), Ionacanthus (?), Isoglossa (83), Isotheca (1), Jadunia (2), Juruasia (?), Justicia (959), Kalbreyeriella (4?), Kenyacanthus (1), Kolobochilus (?), Kosmosiphon (1), Kudoacanthus (1), Lankesteria (7), Lasiocladus (4), Leandriella (2), Lepidagathis (157), Leptosiphonium (10), Leptostachya (1), Liberatia (2 not in APG); Linariantha (1), Lindauea (?), Louteridium (11), Mackaya (5), Marcania (1), Megalochlamys (10), Megalostoma (?), Megaskepasma (1), Melittacanthus (1), Mellera (7), Metarungia (1), Mexacanthus (1), Meyenia (1), Mimulopsis (20), Mirandea (6), Monechma (14?), Monothecium (3), Morsacanthus (1), Neohallia (?), Neriacanthus (1), Neuracanthus (32), Odontonema (32), Oplonia (21), Oreacanthus (?), Orophochilus (?), Pachystachys (18), Pararuellia (11), Pelecostemon (?), Perenideboles (?), Pericalypta (1), Peristrophe (?), Petalidium (46), Phaulopsis (21), Phialacanthus (5), Phlogacanthus (42), Physacanthus (3), Podorungia (5), Poikilacanthus (13), Populina (2), Pranceacanthus (1), Pseuderanthemum (134), Pseudodicliptera (4), Psilanthele (1), Ptyssiglottis (38), Pulchranthus (4), Razisea (?), Rhinacanthus (24), Rhombochlamys (1?), Ritonia (3), Ruellia (379), Ruelliopsis (1), Rungia (87), Ruspolia (4), Ruttya (6), Saintpauliopsis (1), Salpixantha (1), Samuelssonia (1), Sanchezia (45), Sapphoa (2), Satanocrater (4), Schaueria (15), Sclerochiton (18), Sebastiano-schaueria (1), Siphonoglossa (?), Spathacanthus (4), Sphacanthus (2), Sphinctacanthus (2), Standleyacanthus (?), Stenandriopsis (20), Stenandrium (50), Stenosiphonium (?), Stenostephanus (95), Stenothyrsus (1), Streblacanthus (3), Streptosiphon (1), Strobilacanthus (?), Strobilanthes (466), Strobilanthopsis (1), Styasasia (?), Suessenguthia (8), Tacoanthus (?), Tessmanniacanthus (?), Tetramerium (31), Thysanostigma (2), Tremacanthus (?), Trichanthera (2), Trichocalyx (2), Trichosanchezia (1),Vindasia (1), Whitfieldia (13), Xantheranthemum (1), Xerothamnella (2), Yeatesia (3), and Zygoruellia (1).

Avicennioideae: Avicennia (8).

Nelsonioideae: Aymoreana (1), Elytraria (22), Nelsonia (3), and Staurogyne (155).

Thunbergioideae: Anomacanthus (1), Mendoncia (90), Pseudocalyx (6), and Thunbergia (150).

Key Differences From Similar Families

Acanthaceae are most often confused with Lamiaceae or Verbenaceae. Both the Lamiaceae and Verbenaceae usually have squarish stems as opposed to terete or angular, and their leaf nodes are not swollen. Acanthaceae has bracteoles not seen in the other two families, and their stamens are often hairy or spurred. Acanthaceae fruits are dry capsules that are often forcibly ejected as opposed to nutlets in Lamiaceae or drupes or berries in Verbenaceae. 

Distribution of Acanthaceae

Acanthanceae are mostly tropical and subtropical species, with a few temperate outliers. Native throughout Africa, Australia, Indonesia, and the Americas. 

Distribution of Acanthaceae in the Americas

Canadian Genera Include:

Acanthoideae: Dianthera 1 sp. native to ON and QC (s/t included in Justicia); Justicia 1 sp. native in QC and ON. 

USA Genera Include:

Acanthoideae: Acanthus 1 sp. intro to CA; Andrographis 1 sp. intro to VA; Anisacanthus 4 spp. native to AZ, NM, and TX; Asystasia 1 sp. intro to FL and AL?; Barleria 2 spp. intro to FL; Carlowrightia 8 spp. native to CA, AZ, NM, and TX, and intro in FL; Dianthera 9 spp. including 8 native and 1 intro AZ, KS S to TX, and all E to NJ and S to FL from there plus IA, MI, NY, and VT (s/t included in Justicia); Dicliptera 4 spp. native to S half of USA from AZ E to NC and all S plus KS, MO, IL, IN, KY, and VA; Dyschoriste 6 spp. native to NM, AZ, TX, LA, OK, FL, AL, GA, and SC; Eranthemum 1 sp. intro to FL; Graptophyllum 1 sp. intro to FL; Hemigraphis 2 spp. intro to LA and FL; Henrya 1 sp. native to AZ; Hygrophila 6 spp. native and intro to TX, LA, AL, MS, GA, FL, and VA; Hypoestes 1 sp. intro to HI; Justicia 19 spp. native and intro in most of S half USA CA E to NJ and all S exc NV, UT, CO and inc IA, WI, MI, NY, and VT; Megaskepasma monospecific, formerly N SAM endemic, introduced in HI; Odontonema 2 spp. intro to AL and FL; Pseuderanthemum 1 sp. intro to FL and SC; Ruellia 22 spp. native in most of E half USA from ND S to TX and all E exc. ND, SD, CT, RI, MA, VT, NH, ME and including AZ and NM, but intro in AK and NY; Sanchezia 1 former neoendemic sp. intro to HI; Stenandrium 2 spp. native in NM, TX, FL, and GA; Strobilanthes 1 sp. intro LA and FL; Tetramerium 1 sub & tropical Americas endemic sp. native to NM, AZ, and TX; Yeatesia 2 SE NAM endemic spp. native to TX, LA, AL, MS, FL, and GA. Avicennioideae: Avicennia 2 of 8 pantropical spp native TX, MS, AL, GA, LA, FL, intro CA. Nelsonioideae: Elytraria 3 spp. native to NM, AZ, TX, FL, GA, and SC, including 1 narrow endemic of GA, FL, and SC; Nelsonia 1 sp. intro to FL. Thunbergioideae: Thunbergia 5 spp. intro in TX, FL, and HI. 

Mexico Genera Include:

Acanthoideae: Acanthus 1 sp. intro to Pue; Andrographis 1 sp. intro to Pue, Tlx, Mor, and Ver; Anisacanthus 7 spp. native throughout all of Mexico, including 3 endemic to Mexico; Aphanosperma sinaloensis monospecific endemic to BC, BCS, Son, Sin, Chi, Dgo, Zac, NL, and Tam; Aphelandra 3-7? spp. native throughout all of Mexico; Barleria 1? sp. native to most of Mexico except BC, BCS, Son, and Sin; Bravaisia 3 Mexico + N neoendemic spp. native to much of Mexico exc BC, BCS, Son, and Sin; Buceragenia 2 unplaced narrow endemics of C Mexico; Carlowrightia 26 spp. native through all of Mexico, including 15 endemics of Mexico, 7 of which are narrow endemics of Tam (3), Chi (2), Sin, and Yuc; Chalarothyrsus monospecific endemic of Nay, Jal, Col, Mch, Gro, Oax, and Pue; Chileranthemum 3 spp. native to most of Mexico except BC, BCS, Son, and Sin, and includes 1 narrow endemic of Jal + Gro; Dianthera ?? spp. native throughout all of Mexico, s/t included in Justicia now; Dicliptera ?? many spp. native to all of Mexico; Dyschoriste ~20 spp. native throughout all of Mexico; Gypsacanthus monospecific endemic of Nay, Jal, Col, Mch, Gro, Oax, and Pue; Henrya 3 Americas endemic spp. native in all of Mexico, including 2 endemic to S+W Mexico; Holographis 18 spp. endemic genera found throughout all of Mexico; Hoverdenia monospecific endemic to Chi, Coa, NL, SLP, Dgo, Zac, and Ver; Hygrophila 3? spp. native to Nay, Jal, Col, Mch, Gro, Oax, Chp, Ver, Yuc, QR, Cam, and Tab; Hypoestes 1 sp. intro to much of Mexico exc BC, BCS, Son, Sin, Cam, Tab, Chp, Yuc, and QR; Justicia 20+ spp. native throughout all of Mexico; Lepidagathis 1-2 spp. native to Nay, Jal, Col, Gro, Mch, Oax, Chp, Pue, Mor, Ver, Cam, Tab, Yuc, and QR; Louteridium 9 Mexico + neoendemic spp. native to most of Mexico except BC, BCS, Son, and Sin, including 6 narrow endemics of Mch, Oax, Jal + Mch, Ver + Chp, Gro, and Tam; Mexacanthus monospecific endemic of Nay, Jal, Col, Mch, Gro, and Oax; Mirandea 6 Mexico endemic spp. native to most of Mexico except Pue, BC, BCS, Son, and Sin; Odontonema 4-7 spp. native in all of Mexico; Pachystachys 1 neoendemic sp. native to Chp, Cam, Tab, QR, and Yuc, and intro to Nay, Jal, Col, Gro, Mch, and Oax; Poikilacanthus 5 Mexico + neoendemic spp. native to Nay, Jal, Col, Mch, Gro, Oax, Chp, Pue, Tab, Cam, Yuc, and QR, including 4 endemic to Mexico; Pseuderanthemum 5-8 spp. native throughout all of Mexico; Ruellia ~10?? spp. native throughout all of Mexico; Sanchezia 1 former neoendemic sp. intro to Chp, Tab, Cam, Yuc, and QR; Spathacanthus 3 Mexico + CAM endemic spp. native to Nay, Jal, Col, Mch, Gro, Pue, Ver, Chp, Tab, Cam, Yuc, and QR, including 1 narrow endemic of Ver; Stenandrium 8-10 spp. native throughout all of Mexico; Stenostephanus 15 Mexico + neoendemic spp. native to most of Mexico except BC, Son, and Sin; Streblacanthus 2 Mexico & N neoendemic spp. native to Nay, Jal, Col, Mch, Gro, Oax, and Ver; Tetramerium ~25 sub & tropical Americas spp. native in all of Mexico, including 21 endemics of Mexico; Yeatesia 3 SE NAM endemic spp. native to Chi, Coa, NL, Tam, Dgo, Zac, SLP, Hgo, and Mex?, including 1 endemic of NE Mexico. Avicennioideae: Avicennia 2 spp. native to most of Mexico except C. Mexico. Nelsonioideae: Elytraria 4 spp. native to all of Mexico, inc. Mexican Pacific Is., including 2 endemics to Mexico; Nelsonia 1 sp. intro to Nay, Jal, Col, Gro, Mch, and Oax; Staurogyne 1-2? spp. native to Nay, Jal, Col, Mch, Gro, and Oax; Thunbergioideae: Mendoncia 1-2 spp. native to Nay, Jal, Col, Mch, Gro, Oax, Chp, Ver, Tab, Cam, Yuc, and QR; Thunbergia 3? spp. intro to much of Mexico except BC, BCS, Son, and Sin.

Neotropical Genera Include:

Acanthoideae: Acanthus 1 sp. intro to Costa Rica; Ancistranthus monospecific endemic to Cuba; Andrographis 1 sp. intro to Bahamas, Cuba, Hispaniola, Leeward & Windward Is.; Anisacanthus ~9 spp. native to El Salvador, Honduras, Nicaragua, Costa Rica, E Brazil, inc 5 narrow endemics of Nicaragua, Costa Rica, and E Brazil (3); Aphelandra 212 former Mexico & neoendemic spp. native from Mexico S through CAM, Trinidad-Tobago, tropical SAM S to Peru, N Argentina exc. N Chile, Paraguay, and Uruguay, and intro in Cuba, Dominican Republic, and Windward Is.; Asystasia 2 spp intro 2 Cuba, Belize, El Salvador, Honduras, Nicaragua, Antilles (exc. Netherlands Antilles), Venezuela, and SE Brazil; Barleria 1? sp. native to CAM, Colombia, Venezuela, Guyana, intro Bahamas, Antilles (exc. Cayman Is.), SW Caribbean, Trinidad-Tobago, French Guiana, and S Brazil; Barleriola 4 Greater Antilles endemic spp. of Cuba, Dominican Republic, Haiti, and Puerto Rico; Blechum 1 sp? endemic to SE Brazil; Bravaisia 3 Mexico & N neoendemic spp. native to CAM, Cuba, Colombia, Venezuela, and Trinidad-Tobago and intro to Venezuelan Antilles and Windward Is.; Brillantaisia 1 sp. intro Jamaica; Carlowrightia 4 spp. native to CAM (except Panama), and includes 1 narrow endemic in Ecuador; Cephalacanthus monospecific endemic of Peru; Chamaeranthemum 4 neoendemic spp. of Costa Rica, Peru, and SE+S Brazil, including 3 narrow endemics of Costa RIca (2) and Rio de Janeiro SE Brazil; Chileranthemum 2 Mexico + CAM endemic spp. native to Guatemala and El Salvador; Clistax 3 spp. endemic to N+E+S Brazil; Crossandra 1 sp. intro to the Antilles (except Cayman Is. and Netherlands Antilles), El Salvador, and Nicaragua; Cuenotia monospecific endemic NE Brazil; Cyphacanthus monospecific endemic of Colombia; Dasytropis monospecific endemic to Cuba; Dianthera 41 spp. now s/t included in Justicia native from E Canada & USA S through Mexico, CAM, Antilles (exc. Cayman Is & Netherland Antilles), and S through tropical SAM S to N Argentina (exc. N Chile), and intro Bermuda; Dichazothece monospecific endemic SE Brazil; Dicliptera at least 20? spp. native to CAM, Bahamas, Turks-Caicos, Greater Antilles (except Cayman Is.), Leeward & Windward Is., Trinidad-Tobago, and tropical SAM S to N Chile and N Argentina; Dyschoriste ~45 spp. native to CAM (except Belize), Cuba, Hispaniola, Venezuela, Ecuador, Peru, Bolivia, Paraguay, C+S+SE Brazil, N Argentina, and Uruguay, and intro Leeward Is.; Encephalosphaera 3 N SAM endemic spp. of Colombia, Ecuador, Peru, and N Brazil (s/t included in Aphelandra); Eranthemum 1 sp. intro to El Salvador, Antilles (excluding Puerto Rico, Netherlands Antilles, and Cayman Is.), Trinidad-Tobago, and Suriname; Fittonia 2 former N SAM endemic spp. native to Colombia, Venezuela, Ecuador, Peru, Bolivia, and N Brazil and intro to El Salvador and Trinidad-Tobago; Graptophyllum 1 sp. intro to Cuba, Hispaniola, Puerto RIco, Jamaica, Leeward Is, Venezuelan Antilles, Trinidad-Tobago, Honduras, Nicaragua, Panama, and Venezuela; Harpochilus 2 narrow endemic spp. of NE Brazil; Henrya 1 sp. native to CAM (excluding Belize); Herpetacanthus 21 neoendemic spp. of Costa Rica, Honduras, Nicaragua, Panama, Ecuador, French Guiana, Suriname, Peru, Bolivia, and N+E Brazil; Hygrophila 3-10? spp. native to Greater Antilles (exc. Cayman Is.), Trinidad-Tobago, CAM (exc. Costa Rica), and tropical SAM S to N Argentina (exc. N Chile); Hypoestes 1 sp. intro in Guatemala, El Salvador, Honduras, Panama, Colombia, and Paraguay; Isotheca monospecific endemic to Venezuela and Trinidad-Tobago; Justicia ~500 spp. native and widespread in CAM, Greater Antilles (exc. Cayman Is.), Leeward & Windward Is., Trinidad-Tobago, and tropical SAM S to N Argentina (exc. N Chile); Kalbreyeriella 4 neoendemic spp. native to Costa Rica, Panama, Colombia, N Brazil, Ecuador, and Peru; Lankesteria 1 sp. intro to Trinidad-Tobago; Lepidagathis ~50-60 spp. native to CAM, Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, Trinidad-Tobago, tropical SAM S to Peru, NW Argentina, Paraguay, and S Brazil; Liberatia 2 SAM endemic spp. of SE+S Brazil (not in APG); Louteridium 5 Mexico & neoendemic spp. native to CAM (exc. El Salvador), including 1 narrow endemic of Belize; Megaskepasma monospecific, a former N SAM endemic of Suriname and Venezuela, now introduced in Costa Rica, Nicaragua, Panama, Trinidad-Tobago, and Comoros; Morsacanthus monospecific endemic of S Brazil; Neriacanthus monospecific endemic sp. of Jamaica; Odontonema 32 former Americas endemic spp native from Mexico S through CAM, Cuba, Haiti, Leeward & Windward Is., Trinidad-Tobago, Ecuador, Colombia, Venezuela, N+E Brazil, and Guyana, and intro Paraguay, the Dominican Republic, Puerto Rico, and Galapagos; Oplonia 16 neoendemic spp (rest of genus is Madagascar) native to the Bahamas, Turks and Caicos, Cuba, Hispaniola, Jamaica, Puerto Rico, Leeward Islands, Peru, Bolivia, and NW Argentina, including 13 narrow endemics of Cuba (8), Jamaica (3), Peru (2); Orophochilus monospecific endemic of Peru; Pachystachys 18 former neoendemic spp. native from S Mexico, Costa Rica, Panama, Cuba, Lesser Antilles (exc. Netherlands Antilles), and tropical SAM S to N Argentina (exc. Suriname, N Chile, and Uruguay), and intro Guatemala, El Salvador, Hispaniola, and Puerto Rico; Phaulopsis 1 sp. intro to Colombia and Jamaica; Phlogacanthus 1 sp intro to Windward Is.; Poikilacanthus 9 Mexico + neoendemic spp. native to Guatemala, Honduras, Nicaragua, Panama, Venezuela, Peru, Bolivia, E+S Brazil, Paraguay, Uruguay, and N Argentina, inc 6 narrow endemics of Guatemala (2), Bahia NE Brazil (1), São Paulo SE Brazil (1), Peru (1), Trujillo Venezuela (1); Pranceacanthus monospecific endemic of N+C Brazil and Bolivia; Pseuderanthemum ~60 spp. native to CAM, Colombia, Venezuela, Brazil, Ecuador, Peru, Bolivia, intro Cuba, Puerto Rico, Leeward & Windward Is., Venezuelan Antilles, and Trinidad-Tobago; Psilanthele monospecific endemic of Ecuador; Pulchranthus 4 N SAM endemic spp. of Colombia, Venezuela, N Brazil, Suriname, French Guiana, Ecuador, Peru, and Bolivia; Rhombochlamys monospecific endemic of Colombia; Ruellia ~300 spp. native to CAM, Bahamas, Turks-Caicos, Aruba, Antilles (exc. Netherlands Antilles), Trinidad-Tobago, SW Caribbean, Galapagos, and tropical SAM S to N Argentina (exc. N Chile); Ruspolia 1 sp. intro to Trinidad-Tobago; Salpixantha monospecific endemic of Jamaica; Samuelssonia  monospecific endemic of Haiti; Sanchezia 55 former neoendemic spp. native to Panama, Colombia, Venezuela, N Brazil, Ecuador, Peru, and Bolivia, and now intro to Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Antilles (exc. Cayman Is, Netherlands Antilles), and Trinidad-Tobago; Sapphoa 2 spp. endemic genera of Cuba; Schaueria 15 former endemic to C+E+S Brazil, but 1 sp now intro in Trinidad-Tobago; Sebastiano-schaueria monospecific endemic of SE Brazil; Spathacanthus 3 of 4 Mexico & CAM endemic spp. native to Guatemala, Honduras, and Costa Rica, including 1 narrow endemic of Costa Rica; Stenandrium 50 Americas endemic spp native from S USA to Nicaragua (exc. Belize), plus Bahamas, Turks-Caicos, Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is., and tropical SAM S to C Chile, N Argentina (exc. Guyana, Suriname, French Guiana, N Chile); Stenostephanus 95 Mexico + neoendemic spp. native from Mexico S through CAM (exc. Belize, El Salvador) to Colombia, Venezuela, N+SE Brazil, Ecuador, Peru, and Bolivia; Streblacanthus 3 Mexico & N neoendemic spp. native to Guatemala, Costa Rica, Panama, Peru, and N Brazil; Strobilanthes 1 sp. intro to Belize, El Salvador, Honduras, Nicaragua, Panama, Cuba, Hispaniola, Jamaica, Puerto Rico, Windward Is, Venezuelan Antilles, and Trinidad-Tobago; Suessenguthia 8 N SAM endemic spp. of N Brazil, Peru, and Bolivia; Tessmanniacanthus monospecific endemic Peru?; Tetramerium 9 sub & tropical Americas endemic spp. native to Guatemala, El Salvador, Honduras, Nicaragua, Panama, Venezuela, Colombia, Ecuador, Peru, and Bolivia, including 5 narrow endemics of Peru; Trichanthera 2 neoendemic spp. of Panama, Colombia, Venezuela, N Brazil, Guyana, Suriname, French Guiana, Ecuador, and Peru; Trichosanchezia monospecific endemic Peru; Xantheranthemum monospecific of endemic Peru. Avicennioideae: Avicennia 3 spp. native to CAM, Bahamas, Aruba, Turks and Caicos, Antilles, SW Caribbean, Trinidad-Tobago, Colombia, Venezuela, Guyana, Suriname, French Guiana, N, E, and S Brazil, Uruguay, Ecuador, and Peru. Nelsonioideae: Aymoreana monospecific narrow endemic of E Brazil; Elytraria 11 spp. native to CAM, Cuba, Haiti, Aruba, Trinidad-Tobago, and tropical SAM S to Peru, NW Argentina, C+SE Brazil, and intro to Galapagos and includes 9 narrow endemics of Cuba (6), Haiti, Peru, and Ecuador; Nelsonia 1 sp. intro to El Salvador, Honduras, Nicaragua, Panama, Puerto Rico, Colombia, Ecuador, French Guiana, Bolivia, and N+C+E Brazil; Staurogyne 28-30 spp. native from Nicaragua S to Colombia, Venezuela, Guyana, Suriname, French Guiana, Brazil, Bolivia, and Trinidad and Tobago. Thunbergioideae: Mendoncia ~74 spp. native to CAM (exc. El Salvador), Trinidad-Tobago, and tropical SAM S to Peru, Bolivia, Paraguay, and S Brazil; Thunbergia several spp. intro to Bahamas, Antilles (exc. Cayman Is., Netherlands Antilles), CAM (exc. Nicaragua), and tropical SAM S to N Argentina (exc. French Guiana, N Chile, and Uruguay).

Patagonia Genera Include:

Acanthoideae: Stenandrium 1 sp. native in Bio Bio in C Chile, also in Buenos Aires, Argentina just N of Patagonia.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as my own personal observations of species in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • Flora of North America. (1993+). https://floranorthamerica.org/Main_Page.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/ Retrieved Winter 2020 – present.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout the fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

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The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Araceae or Arum Family

Arisaema triphyllum flowers in a spadix enclosed in a sheath. Learn to identify the Araceae family with more morphology photos like this.
Arisaema triphyllum flowers in a spadix enclosed in a sheath, typical of the Araceae family.
Page Last Updated 2026-05-03

Introduction to the Araceae Family

When you learn how to identify the Araceae family, it’s important to understand that it is part of the Alismatales order of the basal monocot flowering plants. As a monocot, they typically have parallel veins in their leaves. However, as a basal monocot, they diverged early in evolution, so it’s unsurprising that their flowers are not typical at all for a monocot. However, the fact that they are very small and densely packed into a spadix enclosed by a showy spathe is a defining characteristic, making it very easy to recognize in the field.

The Araceae family has always been a favorite of mine, having grown up in the temperate rainforest of the British Columbia coast of western Canada, where skunk cabbage is often found in wet forests and swamps. Most people don’t like the smell, hence the name; however, I love the smell because I love swamps and stinky plants, and when I smell skunk cabbage, odds are there is a swamp for me to explore! I love it so much I even have it tattooed on my back!  

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to learn to identify the Araceae family, with no need to know any scientific jargon. Below is additional information on uses and morphology, as well as pictures to help identify family members and individual species found in North America. But for researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems: Perennial herbs, shrubs, or vines; some are aquatic plants, and others grow on trees (epiphytes). They usually lack stems and instead frow from underground corms, rhizomes, or above-ground runners. Leaves are arranged spirally or alternately; most have stalks (petioles) and a base that sheaths or wraps around the plant. Leaves may be simple or compound, and they may contain natural holes. They often contain a milky or watery sap.

Flowers: Unique tiny flowers densely packed in a cylindrical structure called a spadix is the most characteristic feature of this family, which makes them easy to identify. The spadix is almost always accompanied by a spathe, which is a large leaf-like bract that may be green or brightly colored and may persist or fall off. The flowers often emit strong, foul odors to attract pollinators.

Reproductive Features: Some species have bisexual flowers with male (stamens) and female (ovary, style, and stigma) parts in the same flower, while others have separate male and female flowers on the same spike (monoecious), usually with males on top and female flowers below. But the flowers are very tiny, and a hand lens would be needed to see any detail.

Fruits: Fruits are usually fleshy berries with one to several seeds. Occasionally the berries merge into a single compound fruit.

Uses of the Araceae 

Many Araceae are toxic due to the presence of calcium oxalate crystals, which can irritate the digestive tract when ingested. Despite this, however, many are still important food sources in tropical regions, including Alocasia, Amorphophallus, Colocasia esculenta (taro), Monstera, and Xanthosoma sagittifolium. Some are used as traditional herbal medicines, or their roots are used for fiber or arrow poisons. Many genera are cultivated worldwide as ornamentals both outdoors and indoors as houseplants, including Dieffenbachia, Caladium, Philodendron, Zantedeschia, and many more. The Araceae also have unique plants such as Amorphophallus titanum, which has the most massive inflorescence in all the angiosperms, and Wolffia species, which have the smallest flowers.   

Morphology of Araceae in North America

Learn to identify the Araceae family with these morphology photos.

Some Araceae Species I have Covered So Far in North America

Aroideae Subfamily

Arisaema dracontium plant with flower showing an extra-long spadix

Arisaema dracontium—Green Dragon

Herbaceous perennial with one leaf that is compound and divided into two leaflets that are again palmately divided into 5 – 15 leaflets each. Flowers are in a very spadix that sticks out well beyond the sheath, several inches or more. This Araceae member is native to eastern North America, including northeastern Mexico.

Arisamea quinatum plants showing compound leaves with 5 leaflets; compound leaves are relatively common in the Araceae.

Arisaema quinatum or A. triphyllum ssp. quinatum—Southern Jack-in-the-Pulpit

Herbaceous perennial with 3 leaflets per leaf, but the lateral leaflets are deeply 2-lobed, making it look like it has 5 leaflets. Flowers in a spadix enclosed by a fleshy, hooded sheath. Endemic in the southeastern USA, from Texas east to North Carolina.

Arisaema triphyllum flowers in a spadix enclosed in a sheath, typical of the Araceae family.

Arisaema triphyllum – Jack-in-the-Pulpit

Herbaceous perennial with trifoliate compound leaves on long petioles from the ground up. Flowers are in a narrow spadix enclosed by a green or purplish hooded sheathe. Flowers are in a thin green spadix that is closed by a hooded sheath. Native to eastern North America.

Arum italicum plant with flowers

Arum italicum—Italian Arum

Herbaceous perennial with large showy sagittate leaves and a large white or yellow spathe that surrounds its white to reddish spadix. Native to the Mediterranean and cultivated in North America.

Calla palustris plants with flowers growing in a wet ditch

Calla palustris—Bog Arum

Aquatic perennial with rounded to cordate leaves 6 – 10 cm long and nearly as wide on a 10 – 20 cm petiole. Flowers in a greenish-yellow spadix and enclosed by a white sheath. Fruit is a cluster of red berries. Native to cool, temperate areas of the Northern Hemisphere. This was in Midland, MI, USA.

Colocasia esculenta invasive plants in Lake Catherine State Park, AR, USA.

Colocasia esculenta—Taro

Herbaceous perennial from an edible corm with large sagittate leaves on long thick petioles. Spreads vegetatively and rarely flowers. Native to tropical eastern Asia but introduced in North America, where it has become invasive. This was in Hot Springs, AR, USA

Peltandra virginica plant

Peltandra virginica – Green Arrow Arum

Mostly aquatic herbaceous perennials with large sagittate leaves and pale green to white inflorescences enclosed in a darker-colored spathe. Grows in wet, swampy areas. Native throughout eastern North America.

Lemnoideae Subfamily

Lemna minor, a floating aquatic plant

Lemna minor—Common Duckweed

A tiny floating aquatic plant with two (1-4) small leaves and a single free-floating root. Spreads vegetatively. Flowers are rare and inconspicuous.

Orontioideae Subfamily

Lysichiton americanus plant with spadix and spathe, which are both very typical of the Araceae family.

Lysichiton americanus—Skunk Cabbage

An herbaceous perennial of wet, soggy soils and swampy areas. It has large sessile ovate leaves and a large yellow spathe encasing a rough greenish-yellow spadix. Has a strong skunky odor. This lovely but often misjudged Araceae member is native to northwest North America.

Pothoideae Subfamily

Anthurium andraeanum plant showing spadix and a flattened colorful spathe

Anthurium andraeanum – Painter’s Palette

A tropical perennial plant that is often cultivated as a house plant. It has colorful orange to reddish spathes that do not enclose the yellow spadix. It is native to Colombia & Ecuador.

Scientific Botanical Description of the Arecaceae

Flowers of the Araceae

Plants may be hermaphrodite, monoecious, or rarely dioecious. The inflorescence is characteristic of the family, a terminal cylindrical or ovoid spadix on a peduncle with 3-900 sessile flowers very tightly packed together. When monoecious, the male flowers appear on the upper part of the spadix, with the female flowers below. The spadix is almost always subtended by a persistent or deciduous spathe that is often colored. In the Lemnoideae, the spadix is reduced to 1-4 flowers in a pouch. Flowers are small, sessile, actinomorphic, ebracteate, hypogynous, and often possess a strong odor that most find unpleasant. The perianth is usually only present in bisexual flowers, and no hypanthium is present. When present, the perianth is 2-whorled with 4-9 parts that may be free or basally joined.  

Androecium of the Araceae

The androecium has 4, 6, or 8 (1-32) members. Stamens may or may not all be fertile; staminodes are sometimes present. Stamens may be free or connate in Synandria. The arrangement is antitepalous in hermaphrodite flowers that possess a perianth. Anthers are often subsessile, and dehiscence is poricidal, longitudinal, or transverse. 

Gynoecium of the Araceae

The gynoecium has one ovary that is 1-3 (to many) locular, may be sessile or embedded in the spadix, and is syncarpous. The ovary is superior with 3(1-50) carpels, usually with as many locules as carpels. There is usually one short style (sometimes absent) with a hemispheric, capitate, or discoid stigma that is sometimes strongly lobed. Placentation is parietal, axile, basal, or apical. Ovules are one to many per carpel and are usually anatropous and bitegmic.  

Fruit of the Araceae

The fruits are one- to many-seeded indehiscent berries that may be distinct or, less often, connate in a syncarp. Rarely do the fruits form as dehiscent berries via a stylar plate. Seeds are variable in shape, almost always possess endosperm, are oily and sometimes starchy, and sometimes have a fleshy seed coat.

Habit & Leaf Form of the Araceae

Perennial herbs, shrubs, or vines that may be aquatic or terrestrial, emergent or floating, or epiphytic. They grow from vertical or horizontal rhizomes that may or may not be branched, starchy underground corms, or stolons found at or near the surface. Roots are often mycorrhizal and have no root hairs. They usually do not possess typical stems. Plants typically have calcium oxalate crystals or raphides, milky or watery latex, or rarely colored latex. Leaves may appear before or after the inflorescence, and cataphylls are often seen. Leaves are alternate, bifacial, spiral, distichous, or rarely solitary. A petiole is almost always present, and bases are sheathing with a membranous sheath. The leaf blade is simple or compound and sometimes is perforate or fenestrate. Leaf shape is elliptic to obovate or spatulate and sometimes sagittatecordate. Venation can be parallel, pinnate-netted, or palmate-netted. 

Taxonomy of Araceae

There are approximately 3,667 to 6,500 species in the Araceae family across 143 genera (per APG IV 2026) and currently eight accepted subfamilies, which are undergoing revisions and may change in the future. This family is part of the Alismatales order, which is considered part of the basal monocots, diverging early in monocot evolution: 

  1. Aroideae – These have highly variable growth forms, making generalizations difficult. They have staminate flowers with connate stamens with a thick connective. Their pistillate flowers have staminodes. Neither flower type has a perianth. Stigmas, placentation, ovules, etc. are all variable. Their distribution is cosmopolitan.
  2. Gymnostachydoideae – Leaves are two-ranked and linear, margins are minutely toothed, and there is no distinction between the blade and petiole. Their inflorescences are branched. These are restricted to eastern Australia. 
  3. Lasioideae – These are often prickly, rooted aquatics. Their petioles are long, warty, aculeate, or brightly colored, and their spathe is often spirally twisted. Their inflorescence flowers basipetally, and they may or may not have a perianth. They have up to 12 stamens, and their anthers have oblique pore-like slits. They are found pantropically. 
  4. Lemnoideae – Small floating aquatic herbs with 0-5 unbranched hairless roots. They are made of thalloid stem-leaf units that possess only a primary vein without any vascular tissue. They have no perianth and possess only one stamen and one gynoecium. They have a cosmopolitan distribution.
  5. Monsteroideae – Herbs, climbers, and epiphytes. Their pollen is inaperturate, the style has abundant trichosclereids, ovules 1-4 (-many) per carpel are often basal or sometimes hemianatropous, and there are more than 10 seeds per fruit that are often embedded in mucilage. Distribution is pantropical. 
  6. Orontioideae – A variable group that has no vessels, possesses biforine raphides (exc. Lysichiton), leaf blades have a midrib (exc Orontium), have flowers with usually inferior ovaries (exc. Orontium), may or may not have styles. Their ovules are either hemianatropous or basal. Distribution is north temperate. 
  7. Pothoideae – The spathe does not enclose the spadix, may be erect to reflexed, and is persistent in the fruit. Placentation of ovules is basal or parietal. Distribution is pantropical, excluding Africa. 
  8. Zamioculcadoideae – Rhizomatous plants with leaves that are usually 1-3-compound or simple in Stylochaeton. Leaf fine venation is reticulate, and the leaves are pulvinate along the petiole or petiolules. They have staminate and pistillate flowers. Placentation of ovules is axile, and there is one ascending ovule per carpel. Distribution is restricted to Africa.  

Genera of the Arecaceae:

Aroideae: Adelonema (16), Aglaodorum (1), Aglaonema (26), Alocasia (91), Ambrosina (1), Amorphophallus (246), Anchomanes (6), Anubias (8), Apoballis (14), Aridarum (5), Ariopsis (3), Arisaema (224), Arisarum (3), Arophyton (7), Arum (26), Asterostigma (8), Bakoa (1), Bidayuha (1), Boakoaella (?), Biarum (24), Bognera (1), Boycea (1), Bucephalandra (32), Burttianthus (9), Caladium (19), Calla (1), Callopsis (1), Carlephyton (4?), Cercestis (11), Chlorospatha (70), Colletogyne (1), Colobogynium (1), Colocasia (15), Cryptocoryne (78), Culcasia (27), Dieffenbachia (60), Dracunculus (2), Eminium (8), Fenestratarum (2), Filarum (1), Furtadoa (?), Galantharum (1), Gamogyne (6), Gearum (1), Gorgonidium (8), Gosong (1), Hapaline (9), Hayarum (1), Helicodiceros (1), Hera (1), Hestia (1?), Heteroaridarum (3), Homalomena (177), Hottarum (1), Idimanthus (1), Incarum (1 Not listed in APG), Jasarum (1), Josefia (1), Kiewia (3), Lagenandra (24), Lazarum (16), Leucocasia (1), Lorenzia (1), Mangonia (2), Montrichardia (2), Nabalu (1), Naiadia (1), Nephthytis (5), Ooia (12), Peltandra (2), Philodendron (626), Philonotion (3 or syn. of Schismatoglottis), Phymatarum (1), Pichinia (1), Pinellia (11), Piptospatha (3), Pistia (1), Protarum (1), Pseudodracontium (?), Pseudohydrosme (3), Purseglovia (?), Remusatia (4), Rhynchopyle (7), Sauromatum (11), Scaphispatha (2), Schismatoglottis (97), Schottariella (1), Schottarum (3), Spathantheum (1), Spathicarpa (3), Steudnera (11), Synandrospadix (1), Syngonium (41), Taccarum (6), Tawaia (1), Theriophonum (6), Toga (6), Typhonium (79), Typhonodorum (1), Ulearum (2), Vesta (1), Vietnamocasia (1), Xanthosoma (199), Zantedeschia (8), Zomicarpa (2), Zomicarpella (2).

Gymnostachydoideae: Gymnostachys (1).

Lasioideae: Dracontioides (2) and Dracontium (30).  

Lemnoideae: Anaphyllopsis (3), Anaphyllum (2), Cyrtosperma (15), Landoltia (?), Lasia (2), Lasimorpha (1), Lemna (13), Podolasia (1), Pycnospatha (2), Spirodela (4), Urospatha (13), Wolffia (11), and Wolffiella (10).

Monsteroideae: Alloschemone (2), Amydrium (5), Anadendrum (15), Epipremnum (15), Heteropsis (19), Monstera (73), Rhaphidophora (105), Rhodospatha (45), Scindapsus (36), Spathiphyllum (76 inc. Holochlamys), and Stenospermation (58).

Orontioideae: Lysichiton (2), Orontium (1), and Symplocarpus (6).

Pothoideae: Anthurium (1459), Pothoidium (1), and Pothos (66).

Zamioculcadoideae: Gonatopus (5), Stylochaeton (22 s/t as Stylochiton), and Zamioculcas (1).

Key Differences From Similar Families

The Araceae are easily differentiated from similar families with their spadix of numerous small flowers with their unique subtending spathes, their leaves with either parallel or netted venation (unlike only parallel in most monocots), their raphide crystals, and their seeds with endosperm.     

Distribution of Araceae

The Araceae are most diverse in the neotropics of the Americas but are also found in the Old World tropics and temperate regions.

Distribution of Araceae in the Americas

Canadian Genera Include:

Aroideae: Arisaema 2 spp. native to MB, ON, QC, NB, NS, and PE; Arum 1 sp. intro to BC; Calla 1 pan-north temperate sp native to all of Canada except NT; Peltandra 1 E NAM (+Cuba) endemic sp. native to ON and QC; Pinellia 1 sp. ephemeral intro ON; Pistia 1 pantropical sp. intro to ON. Lemnoideae: Lemna 6 of 18 cosmopolitan spp. native to all of Canada except NF, where it was introduced; Spirodela 1 cosmopolitan sp. native to BC, AB, SK, MB, ON, QC, NB, NS, and PE; Wolffia 4 cosmopolitan spp., including 3 native to BC, AB, SK, MB, ON, QC, and NB and 1 intro in ON. Orontioideae: Lysichiton 1 sp. native to BC; Symplocarpus 1 E NAM endemic sp. native to ON, QC, NB, and NS.

USA Genera Include:

Aroideae: Aglaonema 1 sp. intro to FL; Alocasia 1 sp. intro to FL, TX, and HI; Arisaema 2 spp. native to the E USA from ND S to TX and all states E of that; Arum 2 spp. intro to WA, OR, CA, MO, IL, VI, and SC; Caladium 1 sp. intro to LA and FL; Calla 1 pan-north temperate sp native ND, MN, IA, WI, IL, IN, MI, OH, PA, MD, NJ, NY, CT, RI, MA, VT, NH, and ME; Colocasia 1 sp. intro to TX, LA, MS, AL, GA, FL, SC, NC, PA, and HI; Cryptocoryne 1 sp. intro to TX and FL; Dracunculus 1 sp. intro to OR, CA, and TN; Epipremnum 1 sp. intro to FL, HI, Virgin Is., and Puerto Rico; Peltandra 2 E NAM (+Cuba) spp. native KS S to TX plus E USA from MN S to LA and all states east, including 1 endemic to SE USA, also intro in OR and CA; Philodendron 1 sp. intro to FL and HI; Pinellia 1 sp. intro to CA, OH, WV, MD, NJ, PA, NY, and CT; Pistia monospecific pantropical sp. native to CA, AZ, CO, KA, TX, MO, LA, MS, GA, FL, SC, NC, OH, MD, DE, NY, NJ, and CT; Syngonium 1 sp. intro to FL; Xanthosoma 2 spp. intro to TX, FL, and HI; Zantedeschia 2 spp. intro to CA, OR, PA, and HI. Lemnoideae: Lemna 9 spp. native throughout the entire USA, including AK and HI; Spirodela 2 cosmopolitan spp. native throughout the entire continental USA and HI; Wolffia 7 cosmopolitan spp. native to most of the USA except NV, AZ, NM, and CO; Wolffiella 5 cosmopolitan spp. native to WA, CA, TX, OK, MO, AR, LA, IL, IN, OH, KY, TN, MS, AL, GA, FL, SC, NC, VA, PA, NJ, and MA. Monsteroideae: Epipremnum 1 sp. intro to FL and HI. Orontioideae: Lysichiton 1 sp. native to AK, WA, OR, CA, ID, MT, and WY; Orontium monospecific SE USA endemic sp. native from TX E to FL, N to KY, MA, and RI; Symplocarpus 1 E NAM endemic sp. native to MN, IA, WI, IL, IN, OH, KY, TN, NC, VI, WV, MI, PA, MD, DE, NJ, NY, CT, RI, MA, VT, NH, and ME.

Mexico Genera Include:

Aroideae: Aglaonema 1 sp. intro to NL, Sin, Jal, Son, Ver, Gro, Chi, Tab, and Cam; Alocasia 5 spp. intro from Sin E to Coa and all S Mexico; Arisaema 5 spp. native and including 1 intro in Sin, NL, Nay, Tam, SLP, Jal, Col, Mic, Mex, Pue, Gro, Ver, Oax, and Chi; Arum 1 sp. intro to NL, Jal, and Mex; Dieffenbachia 2 spp. native from Sin to S NL and all through S Mexico; Dracunculus 1 sp. intro to Chi, Cam; Leucocasia 1 sp. intro to Jal and Pue; Peltandra 1 sp. intro Tlx; Philodendron ~44-46 spp. native and some intro throughout all of Mexico; Pinellia 1 sp. intro Mex; Pistia monospecific pantropical sp. native throughout Mexico; Syngonium 9 neo spp. native BCS, Sin E to NL, and all through S Mexico; Typhonium 1 sp. intro to Oax, Pue, Ver, Chi, and SLP; Xanthosoma 8 neotropical spp. native most of Mexico exccept north-central and northeast; Zantedeschia 1 sp. intro to Mex. Lasioideae: Dracontium 1 sp. a narrow endemic of SW Chi. Lemnoideae: Lemna 7 spp. native throughout all of Mexico; Spirodela # sp. native throughout all of Mexico; Wolffia 2 cosmopolitan spp. native in Jal, Nay, Gto, Chi, Mex, Qro, Mor, and Cd Mex; Wolffiella 4 cosmopolitan spp. native to NE+C+SE Mexico. Monsteroideae: Monstera 13 neoendemic spp. native Sin E to NL and all through S Mexico, but primarily in the tropical south; Rhodospatha 1 neo sp. native to Ver and Chi; Spathiphyllum 5 spp. including 4 native and 1 intro to Nay E to Ver and all of S Mexico, including 1 endemic to SE Mexico. Pothoideae: Anthurium 31 spp. native (and 1 intro) from Sin to S NL and throughout S Mexico. Zamioculcadoideae: Zamioculcas 1 sp. intro QR.

Neotropical Genera Include:

Aroideae: Adelonema 16 neoendemic spp. native to Costa Rica, Panama, Colombia, Peru, Bolivia, N Brazil, French Guiana, Guyana, and Suriname; Aglaonema 1 sp. intro to Cuba, Puerto Rico, Trinidad-Tobago, and Venezuelan Antilles; Alocasia 5 spp. intro to Cuba, Hispaniola, Jamaica, Puerto Rico, Leeward & Windward Is., Trinidad-Tobago, Venezuelan Antilles, NE Brazil, Venezuela, Paraguay, and Galapagos; Amorphophallus 1 sp. intro Trinidad-Tobago; Arum 1 sp. intro to NE Argentina; Asterostigma 8 W SAM endemic spp. native to most of brazil and in NE Argentina; Bognera monospecific endemic to N Brazil; Caladium 19 neoendemic spp. native from Honduras S to Peru, NW Argentina, Brazil, and to Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is., Venezuelan Antilles, and Trinidad-Tobago; Chlorospatha 70 neoendemic spp. native to Costa Rica, Panama, Colombia, and Ecuador; Colocasia 2 spp. introduced to Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, Trinidad-Tobago, Venezuelan Antilles, Galapagos, Honduras, Nicaragua, Costa Rica, Colombia, Venezuela, Guyana, Brazil, Ecuador, Peru, Bolivia, and Paraguay; Dieffenbachia 60 neoendemic spp. native from S Mexico S through to Peru, Bolivia, NE Argentina (except Uruguay), and including all of the West Indies; Epipremnum 1 sp. intro to Bermuda, Honduras, Costa Rica, Colombia, Ecuador, Suriname, Trinidad-Tobago, Venezuelan Antilles, and Leeward and Windward Is.; Filarum monospecific endemic to Peru; Gearum monospecific endemic to N+C Brazil; Gorgonidium 8 SAM endemic spp. native to Peru, Bolivia, and N Argentina; Idimanthus monospecific narrow endemic to SE Brazil; Incarum 1 W SAM endemic sp. native to Ecuador, Peru, and Bolivia (NOT listed in APG); Jasarum 1 N SAM endemic sp. native to Venezuela and Guyana; Lorenzia monospecific endemic to N Brazil; Mangonia 2 E SAM endemic spp. native to S Brazil and Uruguay; Montrichardia 2 neoendemic spp. native to Guatemala, Belize, Honduras, Nicaragua S to Peru, Bolivia, N+SE Brazil, Trinidad-Tobago, and the Leeward & Windward Is.; Peltandra 1 E NAM sp. native to Cuba; Philodendron ~600 neoendemic spp. native throughout CAM, the West Indies, and N SAM S to Bolivia, NE Argentina, and S Brazil (excluding Uruguay); Philonotion 3 N SAM endemic spp. native to Colombia, Venezuela, Guyana, French Guiana, Suriname, N Brazil, Peru, and Bolivia (listed as syn. of Schismatoglottis in APG??); Pistia monospecific pantropical sp. native throughout CAM, the West Indies, and SAM S to Peru, Bolivia, and NE Argentina; Scaphispatha 2 N SAM endemic spp. native to most of Brazil and Bolivia; Spathantheum monospecific W. SAM endemic sp. of Peru, Bolivia, and NW Argentina; Spathicarpa 3 E SAM endemic spp. native to Brazil, Bolivia, Paraguay, NE Argentina, and Uruguay; Synandrospadix monospecific endemic to Peru, Bolivia, Paraguay, and NW Argentina; Syngonium 41 Mexico + neoendemic spp. native from Mexico S to Peru, Bolivia, C+SE Brazil, Greater Antilles, Trinidad-Tobago, Venezuelan Antilles, and intro S Brazil, Bahamas, Netherland Antilles, Leeward & Windward Is.; Taccarum 6 SAM endemic spp. of Peru, Brazil, Paraguay, Bolivia, and NE Argentina; Typhonium 1 sp. intro to NE+S Brazil, Cuba, Trinidad-Tobago, and Windward Is.; Ulearum 2 N SAM endemic spp. of N Brazil, Ecuador, and Peru; Xanthosoma ~140 spp. native from Mexico S through to Peru, NW Argentina, Paraguay, S Brazil (exc El Salvador), plus Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is., Trinidad-Tobago, and is intro to Galapagos, Jamaica, Venezuelan Antilles, and Cayman Is.; Zantedeschia 1 sp. intro to Nicaragua, Costa Rica, Colombia, Ecuador, NE Brazil, Dominican Republic, Puerto Rico, and Trinidad-Tobago; Zomicarpa 2 narrow endemic spp. of NE Brazil; Zomicarpella 2 N SAM endemic spp. of Colombia, N Brazil, and Peru. Lasioideae: Dracontioides 2 spp. endemic to NE+SE Brazil; Dracontium 29 neoendemic spp. native to the Dominican Republic, Puerto Rico, Trinidad-Tobago, Windward Is., and from Nicaragua S to Peru, Bolivia, Paraguay, and C+SE Brazil. Lemnoideae: Anaphyllopsis 3 N SAM endemic spp. native Venezuela, French Guiana, Suriname, and N Brazil; Lemna 4 spp. native throughout the entire Neotropical zone S to C Chile, Argentina; Spirodela 3 spp. including 2 native and 1 intro throughout all of Neo zone except Guyana, French Guiana, Galapagos, and N Chile; Urospatha 13 neoendemic spp. native to Guatemala, Honduras, Nicaragua S to Peru, Bolivia, Paraguay, and C+SE Brazil; Wolffia 5 cosmopolitan spp., including 3 native (includes 1 endemic to Colombia) and 2 intro found in Guatemala, El Salvador, Honduras, Greater & Lesser Antilles (except Venezuelan Antilles), Colombia, Ecuador, Venezuela, Suriname, Brazil, Bolivia, Paraguay, Uruguay, C Chile, and NE+NW Argentina; Wolffiella 5 cosmopolitan spp. native to Guatemala, El Salvador, Honduras, Costa Rica S to C Chile, N Argentina (exc N Chile), plus Greater Antilles, Leeward & Windward Is, and Trinidad-Tobago. Monsteroideae: Alloschemone 2 N SAM endemic spp. native to N Brazil and Bolivia; Epipremnum 1 sp. intro to Bermuda, Cuba, Hispaniola, Puerto Rico, Trinidad-Tobago, Leeward & Windward Is., Venezuelan Antilles, Colombia, Ecuador, Suriname, and E Brazil; Heteropsis 19 neoendemic spp. native from Nicaragua S to Peru, Bolivia, N Brazil, plus NE+S+SE Brazil; Monstera 73 former neoendemic spp. native from S Mexico S through to Peru, Bolivia, C+S Brazil, Leeward & Windward Is., Netherlands Antilles, and intro to Puerto Rico; Rhodospatha 45 neoendemic spp. native from S Mexico S through CAM (except El Salvador) and N SAM to Peru, Bolivia, N+SE Brazil, and Trinidad-Tobago; Spathiphyllum ~60 spp. native from S Mexico S through CAM and N SAM to Peru, N+C+SE Brazil, Trinidad-Tobago, and intro to Cuba, Puerto Rico, Dominican Republic, and Venezuelan Antilles; Stenospermation 58 neoendemic spp. native to Guatemala, Honduras, and Nicaragua S to Peru, Bolivia, N Brazil, plus disjunct in SE Brazil. Pothoideae: Anthurium 1459 neoendemic spp. native from Mexico all the way south through to Peru, Bolivia, and NE Argentina (except Uruguay) plus all throughout the West Indies (excluding the Bahamas).

Patagonia Genera Include:

Lemnoideae: Lemna 1 sp. native to SE Argentina; Spirodela 1 cosmopolitan sp. native to SE Argentina and C. Chile; Wolffiella 1 cosmopolitan sp. native to SE Argentina.        

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, but also a lot comes from my own observations of North American species.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Flora of North America (FNA) (1993+). https://floranorthamerica.org/Main_Page.
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009 onwards). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/ Retrieved Winter 2020 – present.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


How to Identify the Araliaceae / Ginseng Family

Oplopanax horridus Devil's Club plant with flowers. Learn to identify the Araliaceae family.
Oplopanax horridus – Devil’s Club plant with flowers
Page Last Updated May 14, 2026

Introduction to the Araliaceae Family

The Araliaceae family is best known for its most popular plant, ginseng. But there are so many other lovely plants in this family. My personal favorite member of this family is Oplopanax horridusor Devil’s Club. It’s pretty typical of the family growing as a shrub with large leaves and small flowers in tall umbels followed by small red drupes.

The Araliaceae are part of the Apiales order of core dicots and are very closely related to the Apiaceae (carrot) family and have many overlapping characteristics. In general, however, the Apiaceae are usually herbs that produce dry schizocarps, while the Araliaceae are usually shrubs or trees and usually produce berry-like drupes. 

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to learn to identify the Araliaceae family, with no need to know any scientific jargon. Below this section are morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems: This family is mostly perennial trees and shrubs, but there is an occasional woody vine and herb as well. None release milky juices when damaged. Most have medium to large leaves, including ones over 3 m long. Leaves are usually arranged in a spiral pattern around the stem and have sheaths that wrap around their base. Leaf blades are simple or compound (made of leaflets); when simple, the margins are often divided in some way, often lobed and maple leaf-like.

Flowers: The flowers are usually individually small but arranged in spikes or heads, sometimes branched, and occasionally in umbels similar to the Apiaceae family, which can cause confusion. The flowers are symmetrical with five petals that are often thick or fleshy with a highly variable calyx that may be reduced to a rim of teeth.

Reproductive Features: This varies in the family from bisexual flowers with both male (stamens) and female (ovary, style, stigma) parts in the same flower to separate male and female flowers on separate plants (dioecious). There are usually 5 stamens, but up to 100 in some species. Flowers usually have a fleshy nectar-producing disk at the base of the styles (tubes that capture pollen).

Fruits: Fruits are mostly fleshy berries or drupes (fleshy fruits with stony pits, like a cherry). But sometimes it is a dry fruit that spits into segments (schizocarp) similar to the Apiaceae (carrot) family.

Uses of Araliaceae 

Ornamentals include the angelica tree (Aralia spinosa) and ivy (Hedera spp.), as well as houseplants such as Hedera, Aralia, Polyscias, Schefflera, and Fatsia. Note that Hedera species have become a widespread invasive species in many areas and should only be grown with extreme caution. Chinese rice paper comes from the pith of Tetrapanax papyriferus.

Medicinal herbs include ginseng roots from Panax quinquefolius and devil’s club (Oplopanax horridus) root bark used for respiratory conditions.

Morphology of Araliaceae in North America

Learn how to identify the Araliaceae family with these morphology photos of the Araliaceae family.
Morphology of the Araliaceae Family

Some Araliaceae Species in North America

Aralioideae Subfamily

Aralia nudicaulis plant before flowering, showing compound leaves common in the Araliaceae family.

Aralia nudicaulis – Wild Sarsaparilla

Herbaceous perennial from underground stems. Large compound leaves have 5 (3 – 7) finely serrated leaflets, often purplish green. Small white flowers in rounded clusters 4 – 5 cm wide on scapes not much taller than the leaves. Flowers are followed by edible purple-black berries. Native to northern and eastern North America.

Aralia spinosa deciduous shrub getting its new spring leaves

Aralia spinosa – Devil’s Walking Stick

Aromatic spiny deciduous shrub or small tree 2 – 8 m tall with exceptionally large bipinnate leaves 70 – 120 cm long. Small white flowers in compound panicles are followed by purplish-black berries. Native to eastern North America.

Fatsia japonica plant with berry-like drupes (fruits), the most common fruit of the Araliaceae family.

Fatsia japonica – Paperplant

Evergreen shrub with stout, sparsely branched stems. Large 20 – 40 cm deeply palmately lobed leaves have 7 – 9 lobes, are spirally arranged, are leathery, and are on long petioles. Flowers are small, white, born in dense terminal compound umbels, followed by small black berries. Native to Japan and Korea, cultivated in North America.

Hedera helix - invasive English Ivy growing on a native tree; vines are uncommon in the Araliaceae family.

Hedera helix – English Ivy

A highly invasive, vigorous root-climbing vine with variably 3-5 lobed leaves, depending on the cultivar and if it has reached reproductive age. It seldom flowers but produces small umbels of greenish-yellow flowers followed by purple-black berries. It spreads mostly vegetatively and takes over entire areas when it is left unchecked. Click for more information!

Oplopanax horridus Devil's Club plant with flowers

Oplopanax horridus – Devil’s Club

Woody deciduous spiny perennial shrub with large spiny palmately lobed leaves, small yellow-green flowers in racemes followed by clusters of small red berry-like drupes. Endemic to North America, mostly the Pacific Northwest, with a small disjunct population in the Great Lakes. Click the link for more info!

Hydrocotyloideae Subfamily

Hydrocotyle ranunculoides plants in a Louisiana bayou lake; aquatics are rare in the Araliaceae family

Hydrocotyle ranunculoides – Floating Pennywort

A creeping, mat-forming aquatic perennial of slow-moving, shallow water or wet mud. It has thin stems that may be above or below the water and rounded to kidney-shaped leaves with about 3-7 shallow lobes on the margins and a deeply notched base that makes it appear almost peltate. Small flowers appear separately in clusters. Native to North, Central, & South America but has become invasive in other parts of the world.

Scientific Botanical Description of the Araliaceae Family

Habit & Leaf Form of the Araliaceae

The Araliaceae are a family of shrubs, moderate-sized trees (occasionally very large in some Polyscias), woody epiphytes, vines, and occasionally herbs (Panax, some Aralia, and the Hydrocotyloideae); occasionally some are switch plants. They are self-supporting, epiphytic, or climbing; when climbing, they may be stem twiners or root climbers. Almost always pachycaul with large leaves and thick stems, but sometimes they are leptocaul in Pseudopanax, where long and short shoots are seen. They are non-laticiferous without colored juice, with or without essential oils, and resinous.

Plants may or may not be conspicuously heterophyllous (sometimes, e.g., Hedera helix). Leaves are usually medium-sized but can be enormous (over 3 m in Aralia) or rarely small (1–2 cm in Pseudopanax anomalum). Their attachment to the stem is nearly always alternate, mostly spiral, or rarely distichous, four-ranked, opposite (Cheirodendron, Polyscias), or whorled (Panax). Leaves are often leathery, usually petiolate, but sometimes subsessile. They are usually more or less sheathing but may sometimes be non-sheathing; when sheathing, they have free margins. Leaves may be gland-dotted or not, and they may be aromatic or odorless. Leaf shapes are mostly simple or sometimes compound, including ternate, pinnate, palmate, multiply compound, and sometimes peltate (as in Hydrocotyloideae and some Harmsiopanax). When simple, the lamina is usually dissected pinnatifid or palmatifid but may be entire. They are pinnately or palmately veined. Leaves may or may not have stipules; when present, they are intrapetiolar, often adnate to and hard to distinguish from the petiole base. Some taxa have spines (as in Oplopanax).

Flowers of the Araliaceae

Plants may be hermaphroditic, monoecious, andromonoecious, gynomonoecious, dioecious, or polygamomonoecious. Flowers are aggregated in spikes, heads, and sometimes umbels (comparable to the Apiaceae) and sometimes in large compound inflorescences. Inflorescences are terminal, axillary, leaf-opposed, or rarely epiphyllous. Flowers are usually more or less 5-merous and cyclic and rarely calyptrate. The floral receptacle has neither an androphore nor a gynophore. Perianth has a distinct calyx and corolla or may be petaline, has 10(6–24) parts, and is usually two-whorled and isomerous or anisomerous, but in Meryta, the flower is whorled.

Calyx, when present, has 3–5 (–12) parts; is one whorled, free or connate, entire, lobulate, blunt-lobed, or toothed (when sometimes reduced to small teeth or a rim); and is often open when in bud. Corolla has 5 (3–13) parts where partially divided or lobed segments sometimes complicate the interpretation of parts. It is one whorled and usually alternates with the calyx, with most genera having five sepals alternating with five petals (exceptions do occur). The corolla is free or partially connate at the base, is rarely calyptrate, may be valvate or imbricate (in Aralieae), is always regular, and is often fleshy. Petals are usually sessile, often with broad bases inserted around the whole circumference of the upper part of the ovary.  

Androecium of the Araliaceae

The androecium has 5 (3–12) or 10–100 members made of exclusively fertile stamens that are free of the perianth and one another and all of equal size. Stamens are usually isomerous with the perianth, are inflexed in bud, with filaments and have usually short and fleshy anthers. Stamens may alternate with the corolla members (usually when equal in number), or they may be opposite to them. Anthers are dorsifixed, dehisce via longitudinal slits, and introrse. Anthers are almost always tetrasporangiate (but occasionally appear bisporangiate by fusion during development) or multisporangiate in some Plerandra.

Gynoecium of the Araliaceae

The gynoecium is 2–5(1–100) carpelled, and the pistil is 2–5(1–100) celled. The gynoecium is synovarious to syncarpous with the styles almost always forming a solid or sometimes hollow stylopodium. Or rarely, the gynoecium can appear monomerous in some Polyscias. It may be partly or fully inferior, or rarely superior (sometimes in Tetraplasandra). The ovary is 1–100 locular, and locules have no false septa. The epigynous disk is present with a nectariferous disk between the stylopodium and stamens. The number of styles varies widely from 1–100. When there are two or more, then they are free or partially joined and apical. Stigmas are usually present as a double-stigmatic crest capping the stylopodium and are wet or dry type, papillate, and of Group II and III types. Placentation, when bi/plurilocular (almost always), is axile to apical. When unilocular (rarely) parietal to apical. Ovules in the single cavity (1–2, if two, the second is typically abortive) per locule, pendulous, epitropous, with ventral raphe, anatropous, unitegmic, and usually crassinucellate or otherwise tenuinucellate.  

Fruit of the Araliaceae

Fruit may be fleshy or non-fleshy, often an indehiscent berry or drupe (most Aralioideae) with separable pyrenes or with one stone (there are as many pyrenes as locules). Sometimes it will be a schizocarp with 2-5(6-100) mericarps (mostly Hydrocotyloideae). Gynoecia may combine to form multiple fruits. Seeds are endospermic and oily and may be ruminate (e.g., Hedera).

Taxonomy of Araliaceae

There are 1450 species in 43 genera within the Apiales order of core Eudicots (dicots). Currently, there are two recognized subfamilies in the Araliaceae. 

  1. Aralioideae – The largest subfamily is made mostly of shrubs and trees, sometimes herbs or root or stem climbers. Leaves are usually pinnately to palmately compound, stipulate with connate, intrapetiolar, hooded, or sometimes cauline stipules. Fruit is almost always a drupe.  
  2. Hydrocotyloideae – Herbaceous perennials, sometimes annuals, and some aquatics, but are often found near bodies of water. The stem has endodermis; it is often prostrate, creeping, or mat-forming. The leaf lamina is orbicularpeltate or deeply twice-lobed palmately; the margin is crenate or serrate, and they are stipulate (cauline or petiolar). Flowers are small, and sepals are often tiny or absent.

The Hydrocotyloideae historically was considered part of the Apiaceae. However, modern phylogenetics showed that it was polyphyletic in the Apiaceae and was moved to the Araliaceae instead. Also, there have been a lot of generic changes, with some genera increasing massively, others decreasing, some appearing, and some disappearing, so while this is as up-to-date (May 2026) as the information I have, the family composition is likely to change some in the near future.

Genera:

Aralioideae: Anakasia (1), Aralia (73), Arthrophyllum (2 = Phylloctenium?), Astropanax (15), Astrotricha (20), Brassaiopsis (45), Cephalaralia (1), Cheirodendron (6), Crepinella (33), Cuphocarpus (5?), Cussonia (20), Dendropanax (97), Didymopanax (38), Eleutherococcus (29), Fatsia (3), Gamblea (4), Gastonia (??), Harmsiopanax (3), Hedera (19), Heptapleurum (322), Heteropanax (9), Hunaniopanax (1-2?), Kalopanax? (1), Macropanax (18), Megalopanax? (?), Merrilliopanax (3), Meryta (28), Motherwellia (1), Munroidendron (?), Neocussonia (16), Oplopanax (3?), Opopanax (4), Oreopanax (148), Osmoxylon (61), Panax (15), Pentapanax (?), Plerandra (33), Polyscias (181), Pseudopanax (7), Raukaua (6), Schefflera (13), Sciodaphyllum (146), Seemannaralia (1), Sinopanax (1), Tetrapanax (1), Tetraplasandra (? or syn. of Polyscias), Trevesia (8), Woodburnia (1).

Hydrocotyloideae: Hydrocotyle (182), Trachymene (59).

Key Differences From Similar Families

The Araliaceae and Apiaceae are both similar to and closely related to each other, and often the two can be hard to differentiate. However, some more common differences include the fact that the Apiaceae are usually herbs vs. usually shrubs and trees in the Araliaceae and leaves that usually lack stipules in the Apiaceae vs. those that are usually present in the Araliaceae. Also, the fruit in Apiaceae is usually a dry schizocarp with two mericarps vs. usually a berry-like drupe in the Araliaceae (but sometimes a schizocarp).

Distribution of Araliaceae

Araliaceae is mostly a tropical family, but some are endemic to temperate climates as well. They are in Eurasia, Africa, Australia, New Zealand, New Caledonia, the Pacific Islands, and the Americas, from Arctic Canada to temperate South America.

Distribution of Araliaceae in the Americas

Canadian Genera Include:

Aralioideae: Aralia 5 spp. native to all of Canada except NU; Eleutherococcus 1 sp. intro to ON; Hedera 1-2 spp. intro BC and ON, very invasive in coastal BC; Kalopanax 1 sp. intro to ON; Oplopanax 1 sp. native to BC, AB, YT, and ON; Panax 2 spp. native to ON, QC, NB, NS, and PE. Hydrocotyloideae: Hydrocotyle 4 native to ON, QC, NB, NS, PE, and NL (excluding Labrador) and intro to BC.

USA Genera Include:

Aralia 8 spp. are native and intro to all of the USA exccept NV; Cheirodendron 5 of 6 C Pacific endemic spp. endemic to HI; Eleutherococcus 1 sp. intro to UT, IN, KY, OH, WV, PA, NY, CT, and MA; Hedera 3 spp intro and invasive to most of the USA except NV, MT S to NM, ND S to OK, MN, IA, WI, VT, NH, ME, and inc. HI; Heptapleurum ? spp. intro FL and HI; Kalopanax 1 sp. intro to NY, CT, MD, VA, OH, and IN; Oplopanax 1 sp. native to WA, OR, ID, WY, MI, NY, and AK; Panax 2 spp. native to all E USA from ND S to TX and all E except ND, TX, and FL; Polyscias 10 spp including 9 spp. native/endemic to HI and 1 sp. intro in FL; Tetrapanax monospecific intro AL, FL, and HI. Hydrocotyloideae: Hydrocotyle 9 spp. native and intro to most of the USA except ID, MT, WY, CO, ND, SD, NE, and IA and it is intro in HI.     

Mexico Genera Include:

Aralioideae: Aralia ~5-9 spp. native throughout all of Mexico; Dendropanax ~3-6 spp. native to most of Mexico except BC, BCS, Son, and Sin; Didymopanax 1 sp. native to SW+SE Mexico, Ver; Oreopanax 13? Mexico + neoendemic spp. native throughout all of Mexico, including the Mexican Pacific Is. Hydrocotyloideae: Hydrocotyle 5-10 spp. native throughout all of Mexico.

Neotropical Genera Include:

Aralioideae: Aralia ~ 8 spp. native to CAM, Cuba, Hispaniola, Colombia, Venezuela, Peru, Bolivia, Paraguay, N Argentina, E+S Brazil; Crepinella 33 NW SAM endemic spp. of Colombia, Venezuela, Guyana, N+C Brazil, Ecuador, and Peru; Dendropanax ~75 spp. native to CAM, Greater Antilles, Leeward Is, Venezuelan Antilles, Trinidad-Tobago, tropical SAM S to Peru, Bolivia, NE Argentina (except for Suriname, French Guiana, and Uruguay); Didymopanax 38 Mexico & neoendemic spp. native to CAM, Cuba, Hispaniola, Puerto Rico, Leeward Is, Trinidad-Tobago, tropical SAM S to NE Argentina (exc NW Argentina, N Chile), including at least 12 narrow endemics of Brazil; Fatsia 1 sp. intro Juan Fernandez Is; Heptapleurum 1? sp. intro to Bermuda, Bahamas, Hispaniola, Jamaica, Puerto Rico, Leeward & Windward Is; Oreopanax 148 Mexico + neoendemic spp. of CAM, Antilles (exc Cayman Is, Aruba, Netherlands Antilles), Trinidad-Tobago, Colombia, Venezuela, Guyana, Suriname?, French Guiana, N+E+S Brazil, Ecuador, Peru, Bolivia, NW Argentina; Plerandra 1 sp. intro to Trinidad-Tobago; Polyscias ? spp. intro to El Salvador, Bahamas, Hispaniola, Leeward Is., Puerto Rico, Trinidad-Tobago, and Venezuelan Antilles; Raukaua 1 sp. endemic to N+C Chile; Sciodaphyllum 146 neoendemic spp. of Jamaica, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia, and Venezuela; Tetrapanax monospecific intro to S Brazil. Hydrocotyloideae: Hydrocotyle ?? spp. native and widespread in moist habitats of CAM, Bermuda, the Bahamas, Cuba, Hispaniola, Jamaica, Puerto Rico, the Leeward & Windward Is., Galapagos, and all of SAM, with high diversity in the Andes.

Patagonia Genera Include:

Aralioideae: Raukaua 2 spp. endemic to N+C Chile (1) + S Argentina (1). Hydrocotyloideae: Hydrocotyle ? spp. native throughout Patagonia and the Falkland Is.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as my own personal observations of plants in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B., & Baracat, A. eds. (2009 onwards). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/ Retrieved Winter 2020–current.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

My Current Plant Family Education Fundraiser

I am currently seeking funding to expand my website and SEO capabilities as I keep adding new families, and I am also looking to invest in a new macro lens, as I will soon be adding floral dissections to the families as they become available to me. You can donate to help support native plant education using the GoFundMe link, also at the bottom of the page.

Copyright Information

The information and the photos on this site are free to use for educational purposes, with proper attribution. For other uses, please contact me first.

You can cite this site as follows: Willis, Lyrae (2020+). Lyrae’s Nature Blog – Plant Families of North America. https://lyraenatureblog.com/. Accessed [Enter Date].


Plant Families of North America

Welcome to my Plant Families of North America page! I have been absent awhile, but I am back, and this is my focus, teaching how to identify plant families. Over the next few weeks, I will be updating what I have so far. I will be adding new layman’s descriptions in addition to botanical descriptions, and I will be adding more morphology photos, including dissections of flowers where available. Please come back often and see what’s blooming!

A quick overview of basic taxonomy and how I have ordered this site. Plants are given a scientific name composed of two parts, the genus they belong to and the particular species they are. The genus can be thought of like plant cousins, whereas at the species level they are more like siblings. Beyond genus you get the family, which is all of the cousins from all the related families. Above that you get an order, which is like a group of similar families.

Below I have organized the families in their orders so that you can see similarities between related families. When you click on a particular family, you will get the characteristics and morphology pictures that represent that family, along with some of the species I have covered in North America from that family.

Note that most technical words have a hover-over description available, or go to the dictionary of botanical terms to learn more. In no time, you’ll be able to identify plants like a scientist!

Alismatales Order – Monocot Clade

Apiales Order – Eudicot Clade

Caryophyllales Order – Eudicot Clade

Gentianales Order – Eudicot Clade

Lamiales Order – Eudicot Clade

Magnoliales Order – Magnoliids Clade

Piperales Order – Magnoliids Clade

Sapindales Order – Eudicot Clade

Learn how to identify the Anacardiaceae family
Learn how to identify the Anacardiaceae family

Other References

Some other great resources for information on North American species:

iNaturalist https://www.inaturalist.org/

Canadensys Plant Search https://data.canadensys.net/vascan/search

Flora of North America https://eflora.org

iNaturalist Plant Search https://www.inaturalist.org/home

USDA Plants Database https://plants.sc.egov.usda.gov/home

Currently Seeking Funding To Continue This Non-Profit, Ad-Free Work

If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you! (COMING SOON)


How to Identify the Annonaceae or Custard Apple Family

Asimina triloba, American Pawpaw fruit, in flower. Learn how to identify the Annonaceae Family
Asimina triloba, American Pawpaw fruit, in flower
Page Last Updated August 6, 2026

Introduction to the Annonaceae Family

The Annonaceae is an interesting family to learn to identify. They are mostly tropical trees and shrubs with unique flowers and fleshy, often edible and delicious fruits. The most well-known member of the Annonaceae in North America is the pawpaw fruit, or Asimina triloba. It is the most northern member of this family and produces delicious fruits that taste similar to bananas. Learn how to identify the Annonaceae family with morphology photos like the flowers shown in the photo above of Asimina triloba, which are typical for the Annonaceae family, with their 6 petals and 3 sepals and their androecium in a ball in the center.

The Annonaceae family is part of the Magnoliales order in the Magnoliids clade of angiosperms, the third largest clade after dicots (eudicots) and monocots. Occasionally this clade is referred to as a ‘peripheral angiosperm’ because it is neither a dicot nor a monocot, where the vast majority of flowering plants are classified. Instead, this clade is characterized by features of both dicots and monocots, including trimerous flowers (monocot) and branching veins (dicot), as well as pollen with one pore.  

Beginner’s Guide to the Custard Apple (Annonaceae) Family

If you’re new to plant morphology, this guide is a perfect beginner’s description to learning to identify the Annonaceae family, with no need to know any scientific jargon. Below this section is additional information on uses and morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems of the Custard Apple Family

The Annonaceae family is made of woody trees, shrubs, or vines that may be evergreen or deciduous. They are known for their fibrous and aromatic barks containing essential oils and resins.

Leaves are simple with smooth (entire) edges and are arranged alternately along the stem. The leaves do not wrap around or sheath the stem at their base. And while the leaves are generally odorless, some may be aromatic or dotted with tiny glands. Leaves never have tiny appendages at their base called stipules, which helps distinguish them from the Magnoliaceae.

Flowers of the Custard Apple Family

The flowers are arranged singly or in branched clusters. They are usually symmetrical, and their parts are usually found in multiples of three, similar to most monocots. However, they generally have a distinct outer layer of sepals and 2 inner layers of petals that are often thick and fleshy and sometimes contain visible nectar glands.

Reproductive Features of the Custard Apple Family

The male parts are very distinct, usually with many stamens that are packed together in a characteristic dense ball or flat-topped mass in the center of the flower, which is a good character to help identify the family. The ovary is superior, sitting above the point of where the petals attach.

Fruits of the Custard Apple Family

Fruits are usually fleshy aggregates of multiple berries that fuse together to form a larger, secondary fruit that is often edible. The seeds inside are sometimes encased in brightly colored fleshy coatings (aril or sarcotesta) that attract birds and animals.

Morphology of Annonaceae in North America

So far in North America, I have only photographed Asimina triloba, since it is the most northern species of this primarily tropical family, but here are some pictures of different morphological aspects of that species.

Learn how to identify the Annonaceae family with morphology photos.

Annonaceae Species I have Covered So Far in North America

So far in North America, I have only photographed Asimina triloba, the morphology photos above show pictures of that species. When I cover more species, I will add more pictures here.

Asimina triloba, American Pawpaw fruit, in flower; the most northern member of the Annonaceae family.

Asimina triloba—American Pawpaw

A large shrub or small tree growing to 11 m with simple, alternate-spirally arranged obovatelanceolate leaves 25 – 30 cm long with a cuneate base, acute tip, and entire margin. Leaves smell similar to green bell peppers if bruised. Flowers are red-purple or maroon, 3 – 5 cm wide, and borne singly on stout, hairy axillary stalks (peduncles), appearing with or before the leaves in the spring. The fruit is a large yellowish to brown berry 5 – 15 cm long and weighing up to 510 g that is edible and sweet. Native to eastern North America. Click here to read my blog on Asimina triloba

Uses of Annonaceae 

Many have edible fruits, but non-commercial varieties should be eaten with caution, as they contain varying levels of annonacin, which has been implicated in neurodegenerative diseases. Still, several important tropical commercial fruits come from Annona species (atemoya, cherimoya, custard apple, ilama, sugar apple, sweetsop, and soursop) and Artabotrys.

Medicinal uses include its use as an analgesic and astringent and to treat various conditions, including snakebite, diarrhea, dysentery, arthritis pain, rheumatism, convulsions, neuralgia, and weight loss.

Ecosystem and Wildlife Values of Annonaceae 

The Annonaceae family provides critical, large, nutrient-rich aggregate fruits that serve as major food resources for tropical frugivores like primates, bats, toucans, tapirs, and peccaries, while in temperate zones, Asimina triloba fruits feed raccoons, foxes, opossums, and black bears.

Many species have fleshy flowers that smell of rotting meat, relying on beetles and small flies for pollination, who are rewarded with food or brood sites, while they also serve as host plants for many Lepidoptera, particularly swallowtail butterflies, that rely on the leaves for food.

In tropical forests, especially in Africa and the neotropics, Annonaceae species are critical parts of the forest structure, often dominating the understory.

Taxonomy of Annonaceae

The Annonaceae family has between 2,400 and 2,500 species in 107-110 genera, depending on the classification. They are part of the Magnoliales order in the Magnoliids clade, which is an early-diverging lineage of mesangiosperms, not a true dicot or monocot.

The family is divided into 4 subfamilies as follows:

  1. Anaxagoreoideae is a group of 25 species in a single genus found in tropical America and the tropics in Southeast Asia. They have 2-ranked trunk leaves, sessile stigmas, and dry follicles that are explosively dehiscent, which is a key diagnostic feature.
  2. Ambavioideae is a more widespread group found throughout the tropics and subtropics. This subfamily is characterized by a truncate and dilated anther connective, an intine that does not extrude through the aperture, and ovules with a middle integument.
  3. Annonoideae is a widespread but mostly tropical subfamily, but it does extend into temperate eastern North America. It is characterized by trimerous whorls, often highly modified stamens, and specialized aggregate fruits.
  4. Malmeoideae is restricted to the lowland tropics and is characterized by pollen with a single aperture and specific molecular markers, as well as often having a glass-like endosperm with spiniform ruminations.

Genera of the Annonaceae Family:

Anaxagoreoideae: Anaxagorea (25).

Ambavioideae: Ambavia (2), Cananga (2), Cleistopholis (4), Cyathocalyx (7-12), Drepananthus (28), Lettowianthus (1), Mezzettia (5), and Tetrameranthus (8).

Annonoideae: Afroguatteria (3), Annona (171), Anonidium (5), Artabotrys (111), Asimina (11), Asteranthe (2), Bocagea (4), Boutiquea (1?), Cardiopetalum (3), Cleistochlamys (1), Cymbopetalum (27), Dasymaschalon (31), Desmos (19), Diclinanona (3), Dielsiothamnus (1), Disepalum (10), Duckeanthus (1), Duguetia (97), Fissistigma (59), Friesodielsia (53), Froesiodendron (3), Fusaea (2), Goniothalamus (139), Guatteria (175+), Hexalobus (5), Hornschuchia (12), Isolona (20), Letestudoxa (3), Lukea (2), Mischogyne (5), Monocyclanthus (1), Monodora (14), Neostenanthera (5), Ophrypetalum (1), Porcelia (7), Pseudartabotrys (1), Pyramidanthe (12), Sanrafaelia (1), Sphaerocoryne (8), Toussaintia (4), Trigynaea (9), Uvaria (170), Uvariastrum (5), Uvariodendron (18), Uvariopsis (18), and Xylopia (197).

Malmeoideae: Alphonsea (39), Annickia (11), Bocageopsis (4), Brieya (2), Cremastosperma (31), Dendrokingstonia (3), Desmopsis (46), Ephedranthus (7), Fenerivia (11), Greenwayodendron (6), Huberantha (35), Klarobelia (14), Leoheo (1), Maasia (6), Malmea (7), Marsypopetalum (5), Meiocarpidium (1), Meiogyne (39), Miliusa (67), Mitrephora (53), Mkilua (1), Monanthotaxis (94), Monocarpia (4), Monoön (80), Mosannona (14), Mwasumbia (1), Neo-uvaria (7), Onychopetalum (2), Orophea (61), Oxandra (29), Phaeanthus (9), Phoenicanthus (2), Piptostigma (13), Platymitra (2), Polyalthia (98), Polyalthiopsis (5), Polyceratocarpus (11), Popowia (29), Pseudephedranthus (2), Pseudomalmea (4), Pseudoxandra (24), Pseuduvaria (60), Ruizodendron (1), Sageraea (9), Sapranthus (10), Sirdavidia (1), Stelechocarpus (3), Tridimeris (10), Trivalvaria (13), Unonopsis (48), Wangia (2), and Wuodendron (1).

Key Differences From Similar Families

Myristicaceae members can look similar, but they are easily distinguished by their clear to blood-red latex, which is visible when the bark is slashed. Plants are also dioecious with small, apetalous flowers compared to usually bisexual Annonaceae flowers with distinct sepals and petals.

Magnoliaceae can be differentiated by their large deciduous stipules that leave a distinct ring-like scar surrounding each node, while the Annonaceae have no stipules at all.  

Scientific Botanical Description of the Annonaceae

Habit & Leaf Form of the Annonaceae

The Annonaceae are always trees, shrubs, or lianas that are deciduous or evergreen and produce essential oils and may be resinous. The inner bark is typically fibrous and aromatic, and the pith is septate to diaphragmed. Plants may be self-supporting or climbing via scrambling or hooked, woody pedicels.

Leaves are typically arranged alternately in a spiral and are non-sheathing, simple, and petiolate and may or may not be gland-dotted or aromatic. The lamina and its margins are entire, and the lamina is always pinnately veined and cross-venulate. Leaves have no stipules. Domatia occurs in 10+ genera as pockets or occasionally as hair tufts.

Flowers of the Annonaceae

Annonaceae plants are usually hermaphrodite or sometimes dioecious and rarely monoecious. They are often axillary but may be terminal, leaf-opposed, or cauliflorous. They are typically in monochasial cymes but may be reduced to a solitary flower or in fasciculate clusters.

The receptacle may be elevated, enlarged, or flat. Flowers are regular and cyclic or partially acyclic. Free hypanthium is absent, and a hypogynous disk is present.

The flowers are strictly 3-merous, arranged in 3 distinct whorls, with 3 parts in each whorl (9 parts, rarely more). The perianth has a distinct calyx and corolla that may be petaline; however, sometimes it can be difficult to determine when the outer 1-2 whorls are sepaloid.

The calyx has 3 parts, typically free (sometimes basally connate), and the lobes are longer than the tube but smaller and thinner than the petals.

The corolla typically has 6 free parts (sometimes basally connate) in 2 distinct whorls. Petals are usually thick, fleshy, or coriaceous, with valvate or imbricate aestivation. Inner petals may be smaller, clawed, or connivent over the reproductive organs and frequently bear nectar-secreting glands at their bases. Rarely is the inner petal whorl missing.  

Androecium of the Annonaceae

The androecium contains 6–100+ members that mature centripetally. Members are usually fertile stamens, but sterile staminodes occur in several lineages and are usually located outside or inside the fertile stamens; staminodes are non-petaloid and are often reduced, glandular, or shield-like.

Stamens are free of the perianth and each other, are all equal, and are usually densely packed and spirally arranged in multiple whorls on the receptacle, often forming a compact protective ball or glat-topped shield around the gynoecium.

Filaments are characteristically short, stout, and thick. Anthers are adnate, non-versatile, linear to oblong, tetrasporangiate, and extrorse and dehisce via longitudinal slits away from the gynoecium.

Gynoecium of the Annonaceae

The gynoecium is superior and 1 (rare) to 100(+) carpelled. It is usually apocarpous (carpels spiraled or cyclic) or synstylovarious to syncarpous (e.g., Monodora).

The carpel is 1–many-ovuled. Placentation in apocarpous taxa of individual carpels is ventral or basal when reduced to a single ovule; free carpels are basal. When syncarpous, placentation is parietal.

Stigmas are of the wet type and papillate, secreting a sticky, mucilaginous exudate. In many apocarpous species, individual stigmas coalesce into an aggregate head.

Ovules are anatropous, bitegmic, and crassinucellate. They are ascending with a ventral raphe and may be arillate at the base, developing a distinct sarcotesta or aril.  

Fruit of the Annonaceae

The fruit of the Annonaceae is a fleshy aggregate that is often made of berries; sometimes fruiting carpels coalesce into a secondary syncarp. The fruiting carpel is indehiscent or pseudo-dehiscent.

Seeds are endospermic with 1 to many per pistil and may be encased in fleshy white, yellow, orange, or red arils or sarcotesta, which attract frugivorous birds and animals. The endosperm is ruminate, oily, and has amyloid.

Distribution of Annonaceae

The Annonaceae are primarily a subtropical and tropical family, especially common in lowland forests, and is very rarely temperate. It is widespread, especially in the Old World. In the Americas, it is found all over the Neotropics and north as far as southern Canada.  

Distribution of Annonaceae in the Americas

Canada Annonaceae Genera

Annonoideae Subfamily in Canada

Asimina 1 E NAM endemic spp. native to southern ON, the northernmost Annonaceae in the world.    

USA Annonaceae Genera

Annonoideae Subfamily in the USA

Annona 3 spp. native to FL; Artabotrys 1 sp. introduced in HI; Asimina 11 E NAM endemic spp. native to the E USA from ND S to TX and all states E excluding ND, SD, MN, CT, RI, VT, NH, and ME; Deeringothamnus monospecific endemic of FL; Polyalthia 1 sp. intro to FL. 

Mexico Annonaceae Genera

Anaxagoreoideae Subfamily in Mexico

Anaxagorea 1 sp. native to SW Mexico, Ver.

Ambavioideae Subfamily in Mexico

Cananga 1 sp. intro to SW Mexico, Ver.

Annonoideae Subfamily in Mexico

Annona 14 spp., including 12 native all through Mexico and 1 sp. introduced to Sin, east to Tam, and south to Oax, Chp, QR, and Yuc; Cymbopetalum 2 spp. native to Chp, Ver, and Oax?; Guatteria 2 spp. are native to SW+SE+S Mexico, Ver; Xylopia 1 sp. native to S Chi.

Malmeoideae Subfamily in Mexico

Desmopsis 20 spp. inc. 19 endemic to rainforests of SW+SE Mexico and Ver; Mosannona 1 sp. native to Cam, Chp, QR, Ver, Yuc; Oxandra 4 spp. native to SW+C+SE Mexico and Ver, including 1 narrow endemic of Chp; Sapranthus 3 ~neoendemic spp. native Gro, Oax, QR, Sin, Chp, and Ver, including 1 narrow endemic of Chp; Tridimeris 10 Mexican endemic spp. native to E+C Mexico including 1 narrow endemic of Chp; Unonopsis 2 neoendemic spp. native to SW+SE Mexico, including 1 endemic to Oax; Uvaria 1 of 168 Old World Tropics spp intro Jal;

Neotropical Annonaceae Genera

Anaxagoreoideae Subfamily in the Neotropics

Anaxagorea 22? spp. native CAM (exc. El Salvador), Trinidad-Tobago, tropical SAM S to Peru, Bolivia, and C+SE Brazil, including 5 narrow endemics of Costa Rica & Panama, S Venezuela (2), N Peru, and SE Brazil.

Ambavioideae Subfamily in the Neotropics

Cananga 1 sp. intro Guatemala, Nicaragua, Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is., and Trinidad-Tobago; Tetrameranthus 8 N SAM endemic spp. of N Brazil, Colombia, Venezuela, French Guiana, Ecuador, and Peru.

Annonoideae Subfamily in the Neotropics

Annona ~160 spp., mostly endemics also widely cultivated throughout CAM, Bahamas, Antilles, SW Caribbean, tropical SAM S to N Argentina (excluding Chile); Artabotrys 1 sp. intro to Bahamas, Leeward Is., Windward Is, and Trinidad-Tobago; Bocagea 4 spp. endemic to E Brazil; Cardiopetalum 3 N SAM endemic spp. of Suriname, French Guiana, N+C+E Brazil, Peru, and Bolivia; Cymbopetalum 27 Mexico + Neoendemic spp. native from S Mexico S through CAM and tropical SAM S to Peru, Bolivia, C+E Brazil; Desmos 1 sp. intro to C+E Brazil and Trinidad-Tobago; Diclinanona 3 N SAM endemic spp. native N+C Brazil, Colombia, Peru, and Venezuela; Duckeanthus monospecific narrow endemic of N Brazil; Duguetia 91 spp. native from Nicaragua S through tropical SAM S to Peru, Bolivia, Paraguay, S Brazil (+4 spp. endemic to W Africa); Ephedranthus 7 SAM endemic spp. native to tropical SAM S to Peru, Bolivia, Paraguay, SE Brazil (exc. Ecuador); Froesiodendron 3 N SAM endemic spp. native to Colombia, Peru, and N Brazil; Fusaea 2 N SAM endemic spp. native to N+NE+C Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela, and Bolivia; Guatteria 186 neoendemic spp. native from S Mexico S through CAM (excluding El Salvador), Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, tropical SAM S to Peru, Bolivia, C+S Brazil; Hornschuchia 12 narrow endemic spp. of E. Brazil; Monodora 1 sp. intro Trinidad-Tobago; Porcelia 7 neoendemic spp. of Costa Rica, Panama, Colombia, Venezuela, N+E+S Brazil, Ecuador, Peru, and Bolivia; Trigynaea 9 N SAM endemic spp. native to Colombia, Venezuela, Guyana, Suriname, N+E Brazil, Ecuador, Peru, and Bolivia; Uvaria 1 sp. intro to Trinidad-Tobago; Xylopia 40 spp. native to CAM, Cuba, Jamaica, Trinidad-Tobago, tropical SAM S to Peru, Bolivia, NE Argentina. 

Malmeoideae Subfamily in the Neotropics

Bocageopsis 4 SAM endemic spp. native to Colombia, Venezuela, Guyana, Suriname, French Guiana, N+C+S Brazil, Peru, and Bolivia; Cremastosperma 34 neoendemic spp. native to Costa Rica, Panama, Colombia, Venezuela, N+C Brazil, Ecuador, Peru, and Bolivia, with most diversity in the narrow tropical zone W of the Andes; Desmopsis 27 spp. native from S Mexico S through all of CAM to Colombia, including narrow endemics in Cuba (1), Costa Rica (4), Colombia (3), Honduras (3), and Panama (3). Klarobelia 14 neoendemic spp. native from Costa Rica S to Colombia, Venezuela, N Brazil, Ecuador, Peru, Bolivia, including several endemics to Ecuador; Malmea 7 neoendemic spp. native to Panama, Colombia, Guyana, Suriname, N+NE Brazil, and Peru; Monoön 1 sp. intro Trinidad-Tobago; Mosannona 14 Mexico + neoendemic spp. native to Guatemala, Belize, Honduras, Costa Rica, Panama, Colombia, N Brazil, Guyana, Suriname, Ecuador, Peru, and Bolivia, including 6 narrow endemics of Costa Rica, Barro Colorado Is. Panama, Guatemala, Suriname and Guyana, and Ecuador; Onychopetalum 2 SAM endemic spp. of Venezuela, N+C Brazil, Peru, and Bolivia; Oxandra 28 Mexico + neoendemic spp. native from S Mexico, Guatemala, Belize, Nicaragua, Costa Rica, Panama, Greater Antilles (excluding Cayman Is), Leeward + Windward Is, SW Caribbean, tropical SAM S to Peru, Bolivia, C+SE Brazil, including 9 narrow endemics of Colombia (3), Suriname, Guyana, Brazil (4); Polyalthia 1 sp. intro to Trinidad-Tobago; Pseudephedranthus 2 N SAM endemic spp. of Venezuela, Guyana, Suriname, and N Brazil; Pseudomalmea 4 N SAM endemic spp. of Colombia, Venezuela, N Brazil, Ecuador, Peru, and Bolivia; Pseudoxandra 24 tropical SAM endemic spp. native to Colombia, Venezuela, Guyana, French Guiana, N+C Brazil, Peru, and Bolivia; Ruizodendron monospecific neoendemic of Colombia, N Brazil, Ecuador, Peru, Bolivia, and disjunct Honduras?; Sapranthus 9 Mexico + N neoendemic spp. native to CAM and Colombia, including 1 narrow endemic of Cesar, Colombia. Unonopsis 47 neoendemic spp. are native from S Mexico, CAM (excluding El Salvador), and tropical SAM S to Peru, Bolivia, and C+S Brazil;

Patagonia Annonaceae Genera

Absent 

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. Below should be most of my references for this along with my own personal observations in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Cruz-Chacon, Ivan de la Marisol Castro-Moreno, Lorena Mercedes Luna-Cazares, and Alma Rosa Gonzalez-Esquinca (2016). La Familia Annonaceae Juss. en México. Lacandonia, year 10, vol. 10, num. 2: 71-82, December 2016.
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., Klitgård, B. & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/ Retrieved Winter 2020-current.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout the fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

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