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How to Identify the Araceae or Arum Family

Arisaema triphyllum flowers in a spadix enclosed in a sheath. Learn to identify the Araceae family with more morphology photos like this.
Arisaema triphyllum flowers in a spadix enclosed in a sheath, typical of the Araceae family.
Page Last Updated 2026-04-25

Introduction to the Araceae Family

When you learn how to identify the Araceae family, it’s important to understand that it is part of the Alismatales order of the basal monocot flowering plants. As a monocot, they typically have parallel veins in their leaves. However, as a basal monocot, they diverged early in evolution, so it’s unsurprising that their flowers are not typical at all for a monocot. However, the fact that they are very small and densely packed into a spadix enclosed by a showy spathe is a defining characteristic, making it very easy to recognize in the field.

The Araceae family has always been a favorite of mine, having grown up in the temperate rainforest of the British Columbia coast of western Canada, where skunk cabbage is often found in wet forests and swamps. Most people don’t like the smell, hence the name; however, I love the smell because I love swamps and stinky plants, and when I smell skunk cabbage, odds are there is a swamp for me to explore! I love it so much I even have it tattooed on my back!  

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to learn to identify the Araceae family, with no need to know any scientific jargon. Below is additional information on uses and morphology, as well as pictures to help identify family members and individual species found in North America. But for researchers or those wanting to learn a more in-depth version, check out the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems: Perennial herbs, shrubs, or vines; some are floating or emergent aquatic plants, and others grow as epiphytes (living on trees). They usually lack stems and instead frow from underground corms, rhizomes, or above-ground runners. Leaves are arranged spirally or alternately; most have stalks (petioles) and a base that sheaths or wraps around the plant. Leaves may be simple or compound, and they may contain natural holes. They often contain a milky or watery sap and tiny needle-like crystals in their leaves (visible with a hand lens).

Flowers: Unique tiny flowers densely packed in a cylindrical structure called a spadix is the most characteristic feature of this family, which makes them easy to identify. The spadix is almost always accompanied by a spathe, which is a large leaf-like bract that may be green or brightly colored and may persist or fall off. The flowers often emit strong, foul odors to attract pollinators.

Reproductive Features: Some species have bisexual flowers with male (stamens) and female (ovary, style, and stigma) parts in the same flower, while others have separate male and female flowers on the same spike (monoecious), usually with males on top and female flowers below. The flowers are very tiny, and it can be difficult to see the reproductive parts, but you can dive deeper in the Scientific Botanical Description below and using a hand lens.

Fruits: Fruits are usually fleshy berries with one to several seeds. Occasionally the berries merge into a single compound fruit.

Uses of the Araceae 

Many Araceae are toxic due to the presence of calcium oxalate crystals, which can irritate the digestive tract when ingested. Despite this, however, many are still important food sources in tropical regions, including Alocasia, Amorphophallus, Colocasia esculenta (taro), Monstera, and Xanthosoma sagittifolium. Some are used as traditional herbal medicines, or their roots are used for fiber or arrow poisons. Many genera are cultivated worldwide as ornamentals both outdoors and indoors as houseplants, including Dieffenbachia, Caladium, Philodendron, Zantedeschia, and many more. The Araceae also have unique plants such as Amorphophallus titanum, which has the most massive inflorescence in all the angiosperms, and Wolffia species, which have the smallest flowers.   

Morphology of Araceae in North America

Learn to identify the Araceae family with these morphology photos.

Some Araceae Species I have Covered So Far in North America

Aroideae Subfamily

Arisaema dracontium plant with flower showing an extra-long spadix

Arisaema dracontium—Green Dragon

Herbaceous perennial with one leaf that is compound and divided into two leaflets that are again palmately divided into 5 – 15 leaflets each. Flowers are in a very spadix that sticks out well beyond the sheath, several inches or more. This Araceae member is native to eastern North America, including northeastern Mexico.

Arisamea quinatum plants showing compound leaves with 5 leaflets; compound leaves are relatively common in the Araceae.

Arisaema quinatum or A. triphyllum ssp. quinatum—Southern Jack-in-the-Pulpit

Herbaceous perennial with 3 leaflets per leaf, but the lateral leaflets are deeply 2-lobed, making it look like it has 5 leaflets. Flowers in a spadix enclosed by a fleshy, hooded sheath. Endemic in the southeastern USA, from Texas east to North Carolina.

Arisaema triphyllum flowers in a spadix enclosed in a sheath, typical of the Araceae family.

Arisaema triphyllum – Jack-in-the-Pulpit

Herbaceous perennial with trifoliate compound leaves on long petioles from the ground up. Flowers are in a narrow spadix enclosed by a green or purplish hooded sheathe. Flowers are in a thin green spadix that is closed by a hooded sheath. Native to eastern North America.

Arum italicum plant with flowers

Arum italicum—Italian Arum

Herbaceous perennial with large showy sagittate leaves and a large white or yellow spathe that surrounds its white to reddish spadix. Native to the Mediterranean and cultivated in North America.

Calla palustris plants with flowers growing in a wet ditch

Calla palustris—Bog Arum

Aquatic perennial with rounded to cordate leaves 6 – 10 cm long and nearly as wide on a 10 – 20 cm petiole. Flowers in a greenish-yellow spadix and enclosed by a white sheath. Fruit is a cluster of red berries. Native to cool, temperate areas of the Northern Hemisphere. This was in Midland, MI, USA.

Colocasia esculenta invasive plants in Lake Catherine State Park, AR, USA.

Colocasia esculenta—Taro

Herbaceous perennial from an edible corm with large sagittate leaves on long thick petioles. Spreads vegetatively and rarely flowers. Native to tropical eastern Asia but introduced in North America, where it has become invasive. This was in Hot Springs, AR, USA

Peltandra virginica plant

Peltandra virginica – Green Arrow Arum

Mostly aquatic herbaceous perennials with large sagittate leaves and pale green to white inflorescences enclosed in a darker-colored spathe. Grows in wet, swampy areas. Native throughout eastern North America.

Lemnoideae Subfamily

Lemna minor, a floating aquatic plant

Lemna minor—Common Duckweed

A tiny floating aquatic plant with two (1-4) small leaves and a single free-floating root. Spreads vegetatively. Flowers are rare and inconspicuous.

Orontioideae Subfamily

Lysichiton americanus plant with spadix and spathe, which are both very typical of the Araceae family.

Lysichiton americanus—Skunk Cabbage

An herbaceous perennial of wet, soggy soils and swampy areas. It has large sessile ovate leaves and a large yellow spathe encasing a rough greenish-yellow spadix. Has a strong skunky odor. This lovely but often misjudged Araceae member is native to northwest North America.

Pothoideae Subfamily

Anthurium andraeanum plant showing spadix and a flattened colorful spathe

Anthurium andraeanum – Painter’s Palette

A tropical perennial plant that is often cultivated as a house plant. It has colorful orange to reddish spathes that do not enclose the yellow spadix. It is native to Colombia & Ecuador.

Scientific Botanical Description

Flowers of the Araceae

Plants may be hermaphrodite, monoecious, or rarely dioecious. The inflorescence is characteristic of the family, a terminal cylindrical or ovoid spadix on a peduncle with 3-900 sessile flowers very tightly packed together. When monoecious, the male flowers appear on the upper part of the spadix, with the female flowers below. The spadix is almost always subtended by a persistent or deciduous spathe that is often colored. In the Lemnoideae, the spadix is reduced to 1-4 flowers in a pouch. Flowers are small, sessile, actinomorphic, ebracteate, hypogynous, and often possess a strong odor that most find unpleasant. The perianth is usually only present in bisexual flowers, and no hypanthium is present. When present, the perianth is 2-whorled with 4-9 parts that may be free or basally joined.  

Androecium of the Araceae

The androecium has 4, 6, or 8 (1-32) members. Stamens may or may not all be fertile; staminodes are sometimes present. Stamens may be free or connate in Synandria. The arrangement is antitepalous in hermaphrodite flowers that possess a perianth. Anthers are often subsessile, and dehiscence is poricidal, longitudinal, or transverse. 

Gynoecium of the Araceae

The gynoecium has one ovary that is 1-3 (to many) locular, may be sessile or embedded in the spadix, and is syncarpous. The ovary is superior with 3(1-50) carpels, usually with as many locules as carpels. There is usually one short style (sometimes absent) with a hemispheric, capitate, or discoid stigma that is sometimes strongly lobed. Placentation is parietal, axile, basal, or apical. Ovules are one to many per carpel and are usually anatropous and bitegmic.  

Fruit of the Araceae

The fruits are one- to many-seeded indehiscent berries that may be distinct or, less often, connate in a syncarp. Rarely do the fruits form as dehiscent berries via a stylar plate. Seeds are variable in shape, almost always possess endosperm, are oily and sometimes starchy, and sometimes have a fleshy seed coat.

Habit & Leaf Form of the Araceae

Perennial herbs, shrubs, or vines that may be aquatic or terrestrial, emergent or floating, or epiphytic. They grow from vertical or horizontal rhizomes that may or may not be branched, starchy underground corms, or stolons found at or near the surface. Roots are often mycorrhizal and have no root hairs. They usually do not possess typical stems. Plants typically have calcium oxalate crystals or raphides, milky or watery latex, or rarely colored latex. Leaves may appear before or after the inflorescence, and cataphylls are often seen. Leaves are alternate, bifacial, spiral, distichous, or rarely solitary. A petiole is almost always present, and bases are sheathing with a membranous sheath. The leaf blade is simple or compound and sometimes is perforate or fenestrate. Leaf shape is elliptic to obovate or spatulate and sometimes sagittatecordate. Venation can be parallel, pinnate-netted, or palmate-netted. 

Taxonomy of Araceae

There are approximately 3,667 to 6,500 species in the Araceae family across 143 genera (per APG IV 2026) and currently eight accepted subfamilies, which are undergoing revisions and may change in the future. This family is part of the Alismatales order, which is considered part of the basal monocots, diverging early in monocot evolution: 

  1. Aroideae – Aroideae possess highly variable growth forms, making generalizations difficult. They have staminate flowers with connate stamens with a thick connective. Their pistillate flowers have staminodes. Neither flower type has a perianth. Stigmas, placentation, ovules, etc. are all variable. Their distribution is cosmopolitan.
  2. Gymnostachydoideae – Leaves are two-ranked and linear, margins are minutely toothed, and there is no distinction between the blade and petiole. Their inflorescences are branched. These are restricted to eastern Australia. 
  3. Lasioideae – These are often prickly, rooted aquatics. Their petioles are long, warty, aculeate, or brightly colored, and their spathe is often spirally twisted. Their inflorescence flowers basipetally, and they may or may not have a perianth. They have up to 12 stamens, and their anthers have oblique pore-like slits. They are found pantropically. 
  4. Lemnoideae – Small floating aquatic herbs with 0-5 unbranched hairless roots. They are made of thalloid stem-leaf units that possess only a primary vein without any vascular tissue. They have no perianth and possess only one stamen and one gynoecium. They have a cosmopolitan distribution.
  5. Monsteroideae – Herbs, climbers, and epiphytes. Their pollen is inaperturate, the style has abundant trichosclereids, ovules 1-4 (-many) per carpel are often basal or sometimes hemianatropous, and there are more than 10 seeds per fruit that are often embedded in mucilage. Distribution is pantropical. 
  6. Orontioideae – A variable group that has no vessels, possesses biforine raphides (exc. Lysichiton), leaf blades have a midrib (exc Orontium), have flowers with usually inferior ovaries (exc. Orontium), may or may not have styles. Their ovules are either hemianatropous or basal. Distribution is north temperate. 
  7. Pothoideae – The spathe does not enclose the spadix, may be erect to reflexed, and is persistent in the fruit. Placentation of ovules is basal or parietal. Distribution is pantropical, excluding Africa. 
  8. Zamioculcadoideae – Rhizomatous plants with leaves that are usually 1-3-compound or simple in Stylochaeton. Leaf fine venation is reticulate, and the leaves are pulvinate along the petiole or petiolules. They have staminate and pistillate flowers. Placentation of ovules is axile, and there is one ascending ovule per carpel. Distribution is restricted to Africa.  

Genera:

Aroideae: Adelonema (16), Aglaodorum (1), Aglaonema (27), Alocasia (104), Ambrosina (1), Amorphophallus (235), Anchomanes (6), Anubias (9), Apoballis (12), Aridarum (26), Ariopsis (3), Arisaema (205), Arisarum (6), Arophyton (7), Arum (59), Asterostigma (8), Bakoa (4), Biarum (21), Bognera (1), Bucephalandra (30), Caladium (20-38), Calla (1-2), Callopsis (1), Carlephyton (4), Cercestis (16), Chlorospatha (69), Colletogyne (1), Colobogynium (1), Colocasia (12-17), Croatiella (1), Cryptocoryne (68), Culcasia (30), Dieffenbachia (57-186), Dracunculus (2), Eminium (9), Englerarum (1), Fenestratarum (2), Filarum (1), Furtadoa (4), Galantharum (1), Gamogyne (1), Gearum (1), Gorgonidium (8), Hapaline (8), Helicodiceros (1), Hestia (1), Heteroaridarum (1), Homalomena (145), Hottarum (1), Idimanthus (1), Incarum (1), Jasarum (1), Lagenandra (15), Leucocasia (1), Lorenzia (1), Mangonia (2), Melioblastis (?), Montrichardia (2), Nephthytis (6), Ooia (10), Peltandra (2-4), Philodendron (489-674), Philonotion (3), Phymatarum (1), Pichinia (1), Pinellia (9), Piptospatha (16), Pistia (1), Protarum (1), Pseudohydrosme (2), Remusatia (4), Rhynchopyle (2), Sauromatum (10), Scaphispatha (2), Schismatoglottis (155), Schottariella (1), Schottarum (2), Spathantheum (1), Spathicarpa (3), Stauromatum (?), Steudnera (11), Synandrospadix (1), Syngonium (38-39), Taccarum (6), Thaumatophyllum (19-21), Theriophonum (5), Typhonium (77-82), Typhonodorum (1), Ulearum (2), Vietnamocasia (?), Xanthosoma (98-196), Zantedeschia (8), Zara (?), Zomicarpa (2), Zomicarpella (2).

Gymnostachydoideae: Gymnostachys (1).

Lasioideae: Dracontioides (2), Dracontium (29-30).  

Lemnoideae: Anaphyllopsis (3), Anaphyllum (2), Cyrtosperma (13), Lasia (3),Lasimorpha (1), Lemna (13-18), Podolasia (1), Pycnospatha (2), Spirodela (4), Urospatha (11), Wolffia (11), Wolffiella (10).

Monsteroideae: Alloschemone (2), Amydrium (5), Anadendrum (12), Epipremnum (15), Heteropsis (19-20), Holochlamys (1), Massovia (?), Monstera (49-69), Rhaphidophora (102), Rhodospatha (30-41), Scindapsus (38), Spathiphyllum (52-60), Stenospermation (52-81).

Orontioideae: Lysichiton (3), Lysichitum (1), Orontium (1), Symplocarpus (6).

Pothoideae: Anthurium (1139), Pedicellarum (1), Potha (?), Pothoidium (1), Pothos (79), Tapanawa (?).

Zamioculcadoideae: Gonatopus (5), Stylochaeton Lepr. (20), Stylochaeton Schott (23), Zamioculcas (1).

Key Differences From Similar Families

The Araceae are easily differentiated from similar families with their spadix of numerous small flowers with their unique subtending spathes, their leaves with either parallel or netted venation (unlike only parallel in most monocots), their raphide crystals, and their seeds with endosperm.     

Distribution of Araceae

The Araceae are most diverse in the neotropics of the Americas but are also found in the Old World tropics and temperate regions.

Distribution of Araceae in the Americas

Canadian Genera Include:

Aroideae: Arisaema 2 of 205 cosmopolitan spp native MB, ON, QC, NB, NS, PE; Arum 1 of 59 Eurasian spp intro BC; Peltandra 1 of 2 E NAM endemic spp native ON, QC; Pinellia 1 of 9 Asia spp ephemeral intro ON; Pistia 1 of 1 pantropical spp intro ON. Calloideae (now a tribe of Aroideae): Calla 1 of 1 north temperate spp native all of Canada except NT. Lemnoideae: Lemna 6 of 18 cosmopolitan spp native all of Canada exc NF where it was intro; Spirodela 1 of 4 cosmopolitan spp native BC, AB, SK, MB, ON, QC, NB, NS, PE, Wolffia 4 of 11 cosmopolitan spp inc 3 native BC, AB, SK, MB, ON, QC, NB and 1 intro ON. Orontioideae: Lysichiton 1 of 3 north temperate spp native BC; Symplocarpus 1 of 6 north temperate spp native ON, QC, NB, NS.

USA Genera Include:

Aroideae: Aglaonema 1 of 27 Asian spp intro FL; Alocasia 1 of 104 Australian/Asian spp intro FL, TX, HI; Arisaema 2 of 205 cosmopolitan spp native E USA from ND S to TX and all states E of that; Arum 2 of 59 Eurasian spp intro WA, OR, CA, MO, IL, VI, SC; Caladium 1 of 38 SAM spp intro LA, FL; Calla 1 of 1 north temperate spp native ND, MN, IA, WI, IL, IN, MI, OH, PA, MD, NJ, NY, CT, RI, MA, VT, NH, ME; Colocasia 1 of 12 Asian spp intro TX, LA, MS, AL, GA, FL, SC, NC, PA, and HI; Cryptocoryne 1 of 68 Asian spp intro TX, FL; Dracunculus 1 of 2 spp intro OR, CA, TN; Peltandra 2 of 2 E NAM spp native OR, CA, OK, TX plus E USA from MN S to LA and all states east, inc 1 endemic to SE USA; Philodendron 1 of 564 of tropical Americas spp intro FL and HI; Pinellia 1 of 9 E Asia spp intro CA, OH, WV, MD, NJ, PA, NY, CT; Pistia 1 of 1 pantropical spp native CA, AZ, CO, KA, TX, MO, LA, MS, GA, FL, SC, NC, OH, MD, DE, NY, NJ, CT; Syngonium 1 of 38 neotropical spp intro FL; Xanthosoma 2 of 98 neotropical spp intro TX, FL, and HI; Zantedeschia 2 of 8 S Africa spp intro CA, OR, PA, and HI. Lemnoideae: Lemna 9 of 18 cosmopolitan spp native throughout entire USA inc AK and HI; Spirodela 2 of 4 cosmopolitan spp native entire continental USA and HI; Wolffia 7 of 11 cosmopolitan spp native most of USA exc NV, AZ, NM, CO; Wolffiella 5 of 10 Cosmopolitan spp native WA, CA, TX, OK, MO, AR, LA, IL, IN, OH, KY, TN, MS, AL, GA, FL, SC, NC, VA, PA, NJ, MA. Monsteroideae: Epipremnum 1 of 15 Asian spp intro FL and HI. Orontioideae: Lysichiton 1 of 3 north temperate spp native AK, WA, OR, CA, ID, MT, WY; Orontium 1 of 1 E USA endemic spp native TX, LA, MS, AL, GA, FL, SC, NC, KY, TN, WV, VI, MD, DE, NJ, PA, NY, CT, RI, MA; Symplocarpus 2 of 6 north temperate spp native MN, IA, WI, IL, IN, OH, KY, TN, NC, VI, WV, MI, PA, MD, DE, NJ, NY, CT, RI, MA, VT, NH, ME.

Mexico Genera Include:

Aroideae: Aglaonema 1 of 27 Asian spp intro Nl, Sin, Jal, Son, Ver, Gro, Chi, Tab, Cam; Alocasia 5 of 104 Australian/Asian spp intro Sin E to Coa and all S Mexico; Arisaema 4 of 205 cosmopolitan spp native (and 1 intro) Sin, NL, Nay, Tam, SLP, Jal, Col, Mic, Mex, Pue, Gro, Ver, Oax, Chi; Arum 1 of 59 Eurasian spp intro NL, Jal, Mex; Dieffenbachia 2 of 57 neotropical spp native Sin to S NL and all through S Mexico; Dracontium 1 of 29 neo endemic spp a narrow endemic of SW Chi; Dracunculus 1 of 2 spp intro Chi, Cam; Leucocasia 1 of 1 S Asia spp intro Jal, Pue; Peltandra 1 of 2 E NAM spp intro Tlx; Philodendron 21 of 564 of tropical Americas spp native Sin to S NL and all through S Mexico; Pinellia 1 of 9 E Asia spp intro Mex; Pistia 1 of 1 pantropical spp native throughout Mexico; Syngonium 9 of 38 neotropical spp native BCS, Sin E to NL and all through S Mexico; Thaumatophyllum 3 of 19 N SAM spp intro Sin, Nay, Jal, Col, Ag, SLP, Gto, Cd Mex, Mex, Pue, Mic, Oax, Ver, Chi, Tab, Yuc, QR; Typhonium 1 of 77 Asia spp intro Oax, Pue, Ver, Chi, SLP; Xanthosoma 8 of 98 neotropical spp native most of Mexico exc Chi, BCN, Son; Zantedeschia 1 of 8 S Africa spp intro Mex. Lemnoideae: Lemna 7 of 18 cosmopolitan spp native throughout entire Mexico; Wolffia 2 of 11 cosmopolitan spp native Jal, Nay, Gto, Chi, Mex, Qro, Mor, Cd Mex; Wolffiella 4 of 10 Cosmopolitan spp native NE+C+SE Mexico; Monsteroideae: Monstera 13 of 49 neotropical spp native Sin E to NL and all through S Mexico; Rhodospatha 1 of 30 neotropical spp native Ver, Chi; Spathiphyllum 5 of 52 Neo & W Pacific spp inc 4 native, 1 intro to Nay E to Ver and all of S Mexico, inc 1 endemic to SE Mexico. Pothoideae: Anthurium 31 of 1139 neotropical spp native (and 1 intro) Sin to S NL and throughout S Mexico. Zamioculcadoideae: Zamioculcas 1 of 1 Africa spp intro QR.

Neotropical Genera Include:

Aroideae: Adelonema 16 of 16 spp Neo endemic spp native Costa RIca, Panama, Columbia, Peru, Bolivia, N Brazil, French Guiana, Guyana, Suriname; Aglaonema 1 of 27 Asian spp intro Cuba, Puerto Rico, Trinidad-Tobago, Venezuelan Antilles; Alocasia 5 of 104 Australian/Asian spp intro Cuba, Hispaniola, Jamaica, Puerto Rico, Leeward & Windward Is, Trinidad-Tobago, Venezuelan Antilles, NE BRazil, Venezuela, Paraguay, Galapagos; Amorphophallus 1 of 235 Old World Tropics spp intro Trinidad-Tobago; Arum 1 of 59 Eurasian spp intro NE Argentina; Asterostigma 8 of 8 W SAM endemic spp native NE+C+SE+S Brazil, NE Argentina; Bognera 1 of 1 sp endemic to N Brazil; Caladium 20 of 20 neotropical endemic spp native Honduras S to Peru, NW Argentina, Brazil, and intro Cubo, Hispaniola, Purto Rico, Leeward & Windward Is, Venezuelan Antilles, Trinidad-Tobago and pantropical; Chlorospatha 69 of 69 Neo endemic spp native Costa Rica, Panama, Columbia, Ecuador; Colocasia 2 of 17 Asian spp intro Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, Trinidad-Tobago, Venezuelan Antilles, Galapagos, Honduras, Nicaragua, Costa RIca, Columbia, Venezuela, Guyana, N+NE+S Brazil, Ecuador, Peru, Bolivia, Paraguay; Croatiella 1 of 1 sp endemic to Ecuador; Dieffenbachia 57 of 57 neotropical endemic spp native S Mexico S to Peru, Bolivia, NE Argentina (exc Uruguay) inc all of West Indies; Filarum 1 of 1 sp endemic to Peru; Gearum 1 of 1 sp endemic to N+C Brazil; Gorgonidium 8 of 8 SAM endemic spp native Peru, Bolivia, NW+NE Argentina; Idimanthus 1 of 1 sp endemic to SE Brazil; Incarum 1 of 1 W SAM endemic sp native to Ecuador, Peru, Bolivia; Jasarum 1 of 1 N SAM endemic sp native Venezuela, Guyana; Lorenzia 1 of 1 sp endemic to N Brazil; Mangonia 2 of 2 E SAM endemic spp native S Brazil, Uruguay; Montrichardia 2 of 2 Neo endemic spp native Guatemala, Belize, Honduras, Nicaragua S to Peru, Bolivia, N+SE Brazil, Trinidad-Tobago, Leeward & Windward Is; Peltandra 1 of 2 E NAM spp native Cuba; approx 564 of 564 Americas endemic spp native from Mexico S through to Peru, Bolivia, NE Argentina (exc Uruguay) and inc all of the West Indies; Philonotion 3 of 3 N SAM endemic spp native Colombia, Venezuela, Guyana, French Guiana, Suriname, N Brazil, Peru, Bolivia; Pistia 1 of 1 pantropical spp native throughout CAM, West Indies, and SAM S to Peru, Bolivia, NE Argentina; Scaphispatha 2 of 2 N SAM endemic spp native N+C+NE Brazil, Bolivia; Spathantheum 1 of 1 W SAM endemic spp native Peru, Bolivia, NW Argentina; Spathicarpa 3 of 3 SAM endemic spp native Brazil, Bolivia, Paraguay, NE Argentina, Uruguay; Synandrospadix 1 of 1 sp endemic to Peru, Bolivia, Paraguay, NW Argentina; Syngonium 38 of 38 Mexico + Neo endemic spp native Mexico So to Peru, Bolivia, C+SE Brazil, Greater Antilles, Trinidad-Tobago, Venezuelan Antilles, intro S Brazil, Bahamas, Netherland Antilles, Leeward & Windward Is, and Oceania; Taccarum 6 of 6 SAM endemic spp native Peru, Brazil, Paraguay, Bolivia, NE Argentina; Thaumatophyllum 19 of 19 SAM endemic spp native N SAM S to Peru, Bolivia, NE Argentina (exc Uruguay), intro Belize; Typhonium 1 of 77 Asia & Australia spp intro NE+S Brazil, Cuba, Trinidad-Tobago, Windward Is; Ulearum 2 of 2 SAM endemic spp native N Brazil, Ecuador, Peru; Xanthosoma 98 of 98 Americas endemic spp native from Mexico S through to Peru, NW Argentina, Paraguay, S Brazil (exc El Salvador), plus Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, Trinidad-Tobago, intro to Galapagos, Jamaica, Venezuelan Antilles, Cayman Is; Zantedeschia 1 of 8 S Africa spp intro Nicaragua, Costa Rica, Colombia, Ecuador, NE BRazil, Dominican Republic, Puerto Rico, Trinidad-Tobago; Zomicarpa 2 of 2 spp endemic to NE Brazil; Zomicarpella 2 of 2 N SAM endemic spp native Colombia, N Brazil, Peru. Lasioideae: Dracontioides 2 of 2 spp endemic to NE+SE Brazil; Dracontium 29 of 29 Neo endemic spp native S Mexico, Dominican Republic, Peurto Rico, Trinidad-Tobago, Windward Is, Nicaragua S to Peru, Bolivia, Paraguay, C+SE Brazil. Lemnoideae: Anaphyllopsis 3 of 2 N SAM endemic spp native Venezuela, French Guiana, Suriname, N Brazil; Lemna 4 of 18 cosmopolitan spp native throughout the entire Neotropical zone S to C Chile, Argentina; Spirodela 3 of 4 cosmopolitan spp inc 2 native and 1 intro throughout all of Neo zone exc Guyana, French Guiana, Galapagos, N Chile; Urospatha 11 of 11 Neo endemic spp native Guatemala, Honduras, Nicaragua S to Peru, Bolivia, Paraguay, C+SE Brazil; Wolffia 5 of 11 cosmopolitan spp inc 3 native (inc 1 endemic to Colombia) and 2 intro, found in Guatemala, El Salvador, Honduras, Greater & Lesser Antilles (exc Venezuelan Antilles), Colombia, Ecuador, Venezuela, Suriname, Brazil Bolivia, Paraguay, Uruguay, C Chile, NE+NW Argentina; Wolffiella 5 of 10 Cosmopolitan spp native Guatemala, El Salvador, Honduras, Costa Rica S to C Chile, N Argentina (exc N Chile), plus Greater Antilles, Leeward & Windward Is, Trinidad-Tobago. Monsteroideae: Alloschemone 2 of 2 N SAM endemic spp native N Brazil, Bolivia; Epipremnum 1 of 15 S Asia spp intro Bermuda, Cuba, Hispaniola, Puerto Rico, Trinidad-Tobago, Leeward & Windward Is, Venezuelan Antilles, Colombia, Ecuador, Suriname, E Brazil; Heteropsis 19 of 19 Neo endemic spp native Nicaragua S to Peru, Bolivia, N Brazil, plus NE+S+SE Brazil; Monstera 49 of 49 neotropical endemic spp native Mexico S through to Peru, Bolivia, C+S Brazil, Leeward & Windward Is, Netherlands Antilles, intro to Puerto Rico and Asia; Rhodospatha 30 of 30 neotropical spp native S Mexico S to Peru, Bolivia, N+SE Brazil (exc El Salvador), Trinidad-Tobago; Spathiphyllum ?? of 52 Neo, Malesia & W Pacific spp native S Mexico S to Peru, N+C+SE Brazil, Trinidad-Tobago and intro to Cuba, Puerto Rico, Dominican Republic, Venezuelan Antilles; Stenospermation 52 of 52 Neo endemic spp native Guatemala, Honduras, Nicaragua S to Peru, Bolivia, N Brazil, plus SE Brazil. Pothoideae: Anthurium 1139 of 1139 neotropical endemic spp native Mexico all S through to Peru, Bolivia, NE Argentina exc Uruguay plus all throughout the West Indies exc the Bahamas.

Patagonia Genera Include:

Lemnoideae: Lemna 1 of 18 cosmopolitan spp native SE Argentina; Spirodela 1 of 4 cosmopolitan spp native SE Argentina, C Chile; Wolffiella 1 of 10 Cosmopolitan spp native SE Argentina.        

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, but also a lot comes from my own observations of North American species.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Flora of North America (FNA) (1993+). https://floranorthamerica.org/Main_Page.
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., KlitgÃ¥rd, B., & Baracat, A. eds. (2009 onwards). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/ Retrieved Winter 2020 – present.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

How to Identify the Araliaceae / Ginseng Family

Oplopanax horridus Devil's Club plant with flowers. Learn to identify the Araliaceae family.
Oplopanax horridus – Devil’s Club plant with flowers
Page Last Updated 2026-04-19

Introduction to the Araliaceae Family

The Araliaceae family is best known for its most popular plant, ginseng. But there are so many other lovely plants in this family. My personal favorite member of this family is Oplopanax horridusor Devil’s Club. It’s pretty typical of the family growing as a shrub with large leaves and small flowers in tall umbels followed by small red drupes.

The Araliaceae are part of the Apiales order of core dicots and are very closely related to the Apiaceae (carrot) family and have many overlapping characteristics. In general, however, the Apiaceae are usually herbs that produce dry schizocarps, while the Araliaceae are usually shrubs or trees and usually produce berry-like drupes. 

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to learn to identify the Araliaceae family, with no need to know any scientific jargon. Below this is additional information on uses and morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems: This family is mostly perennial trees and shrubs, but there is an occasional woody vine and herb as well. None release milky juices when damaged, but many have essential oils and are often resinous. Some have very thick stems, and most have medium to large leaves, including rare ones that are enormous (over 3 m long). Leaves are usually arranged in a spiral pattern on the stem and have sheaths that wrap around the stem. Leaf blades are simple or compound (made of leaflets); when simple, the margins are often divided in some way. Some species, like English Ivy (Hedera helix), have leaves that change shape as they mature (heterophylly).

Flowers: The flowers of this family are usually individually small and are often arranged in spikes or heads, or umbels similar to the carrot family, which can cause confusion. Sometimes their inflorescences (arrangements of flowers) are branched. The flowers are symmetrical with five petals that are often thick or fleshy and are usually free (not joined) and a calyx that is highly variable and sometimes even reduced to tiny teeth or a rim.

Reproductive Features: This varies in the family from bisexual flowers with both male (stamens) and female (ovary, style, stigma) parts in the same flower to separate male and female flowers on separate plants (dioecious). There are usually 5 stamens, but up to 100 in some species, that are free from the petals (unattached). The ovary is located below the point of petal attachment (inferior), and the flowers usually have a fleshy nectar-producing disk at the base of the styles (tubes that capture pollen and direct it to the ovary), which is a good identifying feature as it is very common in the family.

Fruits: Fruits are mostly fleshy berries or drupes (fleshy fruits with stony pits, like a cherry). But sometimes it is a dry fruit that spits into segments (schizocarp) similar to the Apiaceae (carrot) family.

Uses of Araliaceae 

Ornamentals include the angelica tree (Aralia spinosa) and ivy (Hedera spp.), as well as houseplants such as Hedera, Aralia, Polyscias, Schefflera, and Fatsia. Note that Hedera species have become a widespread invasive species in many areas and should only be grown with extreme caution. Chinese rice paper comes from the pith of Tetrapanax papyriferus.

Medicinal herbs include ginseng roots from Panax quinquefolius and devil’s club (Oplopanax horridus) root bark used for respiratory conditions.

Morphology of Araliaceae in North America

Learn how to identify the Araliaceae family with these morphology photos of the Araliaceae family.
Morphology of the Araliaceae Family

Some Araliaceae Species in North America

Aralioideae Subfamily

Aralia nudicaulis plant before flowering, showing compound leaves common in the Araliaceae family.

Aralia nudicaulis – Wild Sarsaparilla

Herbaceous perennial from underground stems. Large compound leaves have 5 (3 – 7) finely serrated leaflets, often purplish green. Small white flowers in rounded clusters 4 – 5 cm wide on scapes not much taller than the leaves. Flowers are followed by edible purple-black berries. Native to northern and eastern North America.

Aralia spinosa deciduous shrub getting its new spring leaves

Aralia spinosa – Devil’s Walking Stick

Aromatic spiny deciduous shrub or small tree 2 – 8 m tall with exceptionally large bipinnate leaves 70 – 120 cm long. Small white flowers in compound panicles are followed by purplish-black berries. Native to eastern North America.

Fatsia japonica plant with berry-like drupes (fruits), the most common fruit of the Araliaceae family.

Fatsia japonica – Paperplant

Evergreen shrub with stout, sparsely branched stems. Large 20 – 40 cm deeply palmately lobed leaves have 7 – 9 lobes, are spirally arranged, are leathery, and are on long petioles. Flowers are small, white, born in dense terminal compound umbels, followed by small black berries. Native to Japan and Korea, cultivated in North America.

Hedera helix - invasive English Ivy growing on a native tree; vines are uncommon in the Araliaceae family.

Hedera helix – English Ivy

A highly invasive, vigorous root-climbing vine with variably 3-5 lobed leaves, depending on the cultivar and if it has reached reproductive age. It seldom flowers but produces small umbels of greenish-yellow flowers followed by purple-black berries. It spreads mostly vegetatively and takes over entire areas when it is left unchecked. Click for more information!

Oplopanax horridus Devil's Club plant with flowers

Oplopanax horridus – Devil’s Club

Woody deciduous spiny perennial shrub with large spiny palmately lobed leaves, small yellow-green flowers in racemes followed by clusters of small red berry-like drupes. Endemic to North America, mostly the Pacific Northwest, with a small disjunct population in the Great Lakes. Click the link for more info!

Hydrocotyloideae Subfamily

Hydrocotyle ranunculoides plants in a Louisiana bayou lake; aquatics are rare in the Araliaceae family

Hydrocotyle ranunculoides – Floating Pennywort

A creeping, mat-forming aquatic perennial of slow-moving, shallow water or wet mud. It has thin stems that may be above or below the water and rounded to kidney-shaped leaves with about 3-7 shallow lobes on the margins and a deeply notched base that makes it appear almost peltate. Small flowers appear separately in clusters. Native to North, Central, & South America but has become invasive in other parts of the world.

Scientific Botanical Description of the Araliaceae Family

Habit & Leaf Form of the Araliaceae

Perennial trees of moderate size but occasionally very large with Peekeliopanax reaching 40 m high; arborescent, shrubs, woody epiphytes, lianas, or herbs (Panax, Stilbocarpa, some Aralia, etc.). They are non-laticiferous without colored juice, with or without essential oils, and resinous. Occasionally some are switch plants. Plants are green and photosynthesizing. Self-supporting, epiphytic, or climbing; when climbing, they may be stem twiners or root climbers. Almost always pachycaul with large leaves and thick stems, but sometimes they are leptocaul in Pseudopanax, where long and short shoots are seen.

Plants may or may not be conspicuously heterophyllous (sometimes, e.g., Hedera helix, where progression from lobed to entire leaves reflects irreversible shoot maturation). Leaves are usually medium to enormous (to over 3 m in Aralia) or rarely small (1–2 cm in Pseudopanax anomalum). Their attachment to the stem is nearly always alternate, mostly spiral, or rarely distichous, four-ranked, opposite (Cheirodendron, Polyscias), or whorled (Panax). Leaves are often leathery and usually are petiolate but also sometimes subsessile. They are usually more or less sheathing but may sometimes be non-sheathing; when sheathing, they have free margins. Leaves are gland-dotted or not and aromatic or odorless. Leaf shape is mostly simple (mostly) or compound ternate, pinnate, palmate, multiply compound, and sometimes peltate (some Harmsiopanax). Lamina, when simple, is usually dissected pinnatifid or palmatifid but may be entire. They are pinnately or palmately veined. Leaves may be stipulate or exstipulate. When stipules are present, they are intrapetiolar, often adnate to and scarcely distinguishable from the base of the petiole. Vegetative buds are scaly. Some taxa have spines (as in Oplopanax).

Flowers of the Araliaceae

Plants may be hermaphroditic, monoecious, andromonoecious, gynomonoecious, dioecious, or polygamomonoecious. Flowers are aggregated in spikes, heads, and sometimes umbels (compared to usual umbels in the Apiaceae). Inflorescences are terminal, axillary, leaf-opposed, or rarely epiphyllous. Some genera have racemes, usually with umbels or heads, often massed into compound inflorescences. Flowers are usually more or less 5-merous and cyclic and rarely calyptrate. The floral receptacle has neither an androphore nor a gynophore. Perianth has a distinct calyx and corolla or may be petaline, has 10(6–24) parts, and is usually two-whorled and isomerous or anisomerous. In Meryta, the flower is whorled, and the calyx is entirely lacking. Calyx, when present, has 3–5 (–12) parts; is one whorled, free or connate, entire, lobulate, blunt-lobed, or toothed (when sometimes reduced to small teeth or a rim); and is often open when in bud. Corolla has 5 (3–13) parts where partially divided or lobed segments sometimes complicate the interpretation of parts. It is one whorled and usually alternates with the calyx, with most genera having five sepals alternating with five petals (exceptions do occur). The corolla is free or partially connate at the base, is rarely calyptrate, may be valvate or imbricate (in Aralieae), is always regular, and is often fleshy. Petals are usually sessile, often with broad bases inserted around the whole circumference of the upper part of the ovary, or sometimes they are clawed, as in Mackinleyeae.  

Androecium of the Araliaceae

The androecium has 5(3–12) or 10–100 members. Androecial members are free of perianth and one another; all are equal, and all are fertile. Stamens vary in number, with 5 (3–12) or 10–100; are usually isomerous with the perianth; are inflexed in bud; and are filantherous with usually short and fleshy anthers. Stamens may alternate with the corolla members (usually when equal in number), or they may be opposite to them. Anthers are dorsifixed, dehisce via longitudinal slits, and introrse. Anthers are almost always tetrasporangiate (but occasionally appear bisporangiate by fusion during development) or multisporangiate in Octotheca and Dizygotheca.

Gynoecium of the Araliaceae

The gynoecium is 2–5(1–100) carpelled, and the pistil is 2–5(1–100) celled. The gynoecium is synovarious to syncarpous with the styles almost always forming a solid or sometimes hollow stylopodium. Or rarely, the gynoecium can appear monomerous in some Polyscias. It may be partly or fully inferior, or rarely superior (sometimes in Tetraplasandra). The carpel, when monomeric (rarely), is one-ovuled with a second abortive ovule. The ovary is 1–100-locular. Locules have no false septa. The epigynous disk is present with a nectariferous disk between the stylopodium and stamens. The number of styles varies widely from 1–100. When there are two or more, then they are free or partially joined and apical. Stigmas are usually present as a frequently double-stigmatic crest capping the stylopodium and are wet or dry type, papillate, and Group II and III types. Placentation, when bi/plurilocular (almost always), is axile to apical. When unilocular (rarely) parietal to apical. Ovules in the single cavity 1–2 (if two, the second is typically abortive) per locule, pendulous, epitropous, with ventral raphe, anatropous, unitegmic, and usually crassinucellate or otherwise tenuinucellate.  

Fruit of the Araliaceae

Fruit may be fleshy or non-fleshy, often an indehiscent berry or drupe with separable pyrenes or with one stone (there are as many pyrenes as locules). Sometimes it will be a schizocarp with 2-5(6-100) mericarps. Gynoecia may combine to form multiple fruits. Seeds are endospermic, ruminate (e.g., Hedera) or not, and oily.

Taxonomy of Araliaceae

There are 1450 species in 43 genera within the Apiales order of core Eudicots (dicots). Currently, there are two recognized subfamilies in the Araliaceae. 

  1. Aralioideae – The largest subfamily is made mostly of shrubs and trees, sometimes herbs or root or stem climbers. Leaves are usually pinnately to palmately compound, stipulate with connate, intrapetiolar, hooded, or sometimes cauline stipules. Fruit is almost always a drupe.  
  2. Hydrocotyloideae – Herbaceous perennials, sometimes annuals, and some aquatics; the stem has endodermis. The leaf lamina is orbicularpeltate or deeply twice-lobed palmately; the margin is crenate or serrate, and they are stipulate (cauline or petiolar). 

The Hydrocotyloideae historically was considered part of the Apiaceae. However, modern phylogenetics showed that it was polyphyletic in the Apiaceae and was moved to the Araliaceae instead.

Genera:

Aralioideae: Anakasia (1), Aralia (73), Astropanax (15), Astrotricha (20), Brassaiopsis (45), Cephalaralia (1), Cheirodendron (6), Chengiopanax (2), Crepinella (1), Cussonia (20), Dendropanax (91), Didymopanax (16), Eleutherococcus (29), Fatsia (3), Gamblea (4), Harmsiopanax (3), Hedera (18), Heptapleurum (63), Heteropanax (9), Kalopanax (1), Macropanax (18), Merrilliopanax (3), Meryta (29), Metapanax (2), Motherwellia (1), Neocussonia (16), Neopanax (5), Oplopanax (3), Oreopanax (147), Osmoxylon (61), Panax (13), Plerandra (33), Polyscias (173), Pseudopanax (7), Raukaua (6), Schefflera (309), Sciodaphyllum (21), Seemannaralia (1), Sinopanax (1), Tetrapanax (1), Trevesia (8), Woodburnia (1).

Hydrocotyloideae: Hydrocotyle (130), Trachymene (45).

Key Differences From Similar Families

The Araliaceae and Apiaceae are both similar to and closely related to each other, and often the two can be hard to differentiate. However, some more common differences include the fact that the Apiaceae are usually herbs vs. usually shrubs and trees in the Araliaceae and leaves that usually lack stipules in the Apiaceae vs. those that are usually present in the Araliaceae. Also, the fruit in Apiaceae is usually a dry schizocarp with two mericarps vs. usually a berry-like drupe in the Araliaceae (but sometimes a schizocarp).

Distribution of Araliaceae

Araliaceae is mostly a tropical family, but some are endemic to temperate climates as well. They are in Eurasia, Africa, Australia, New Zealand, New Caledonia, Pacific Islands, and the Americas, from Arctic Canada to temperate South America.

Distribution of Araliaceae in the Americas

Canadian Genera Include:

Aralioideae:Aralia 5 of 73 Americas & Asia spp native all of Canada exc NU; Eleutherococcus 1 of 29 E Asian spp intro ON; Hedera 1 of 18 mostly Eurasian spp intro BC, ON, very invasive where it has escaped;Kalopanax monospecific E Asia sp intro ON;Oplopanax 1 of 3 Americas & E Asia spp native BC, AB, YT, ON; Panax 2 of 13 E Asia & NAM spp native ON, QC, NB, NS, PE, most of the genus endemic to Asia. Hydrocotyloideae: Hydrocotyle 4 of 130 cosmopolitan spp native ON, QC, NB, NS, PE, NL (exc Labrador), intro BC.

USA Genera Include:

Aralia 8 of 73 Americas & Asia spp native and intro all of USA exc NV; Cheirodendron 5 of 6 C Pacific endemic spp endemic to HI; Eleutherococcus 1 of 29 E Asian spp intro UT, IN, KY, OH, WV, PA, NY, CT, MA; Hedera 3 of 18 mostly Eurasian spp intro most of USA EXC NV, MT S to NM, ND S to OK, MN, IA, WI, VT, NH, ME and inc HI, very invasive where it has escaped; Heptapleurum ? of 63 S & Tropical Asia spp intro FL and HI; Kalopanax 1 monospecific E Asia sp intro NY, CT, MD, VA, OH, IN;Oplopanax 1 of 3 Americas & E Asia spp native WA, OR, ID, WY, MI, NY, AK; Panax 2 of 13 E Asia spp native all E USA ND S to TX and all E exc ND, TX, FL, most of the genus endemic to E Asia; Polyscias 10 of 173 Old World Tropics & Pacific spp inc 9 spp native/endemic HI (sometimes as Munroiodendron, Reynoldsia, Tetraplasandra) and 1 sp intro FL;Schefflera 2 of 309 pantropical spp (exc Africa) native and intro FL, native HI; Tetrapanax monospecific S China sp intro AL, FL and HI. Hydrocotyloideae: Hydrocotyle 9 of 130 cosmopolitan spp native and intro most USA exc ID, MT, WY, CO, ND, SD, NE, IA and intro HI.     

Mexico Genera Include:

Aralioideae: Aralia ?? of 73 Americas & Asia spp native all of Mexico; Dendropanax ?? of 91 Americas & Tropical & E Asia spp native moist forests at lowland to mid-elevation areas most of Mexico exc BC, BCS, Son, Sin; Didymopanax 1 of 16 Mexico & Neo endemic spp native SW+SE Mexico, Ver; Oreopanax ?? of 147 Mexico & Neo endemic spp native throughout Mexico inc Mexican Pacific Is. Hydrocotyloideae: Hydrocotyle ?? of 130 cosmopolitan spp native throughout all of Mexico.

Neotropical Genera Include:

Aralioideae: Aralia ?? of 73 Americas & Asia spp native CAM, Cuba, Hispaniola, Colombia, Venezuela, Peru, Bolivia, Paraguay, N Argentina, E+S Brazil, most in seasonally dry forests below 500m; Crepinella monospecific SAM endemic sp of Colombia, Venezuela, Guyana, N+C Brazil, Ecuador, Peru; Dendropanax ?? of 91 Americas & Tropical & E Asia spp native moist forests at low to mid-elevation areas up to around 1500m in CAM, Greater Antilles, Leeward Is, Venezuelan Antilles, Trinidad-Tobago, tropical SAM S to Peru, Bolivia, NE Argentina (exc Suriname, French Guiana, Uruguay); Didymopanax 16 of 16 spp Mexico & Neo endemic spp native CAM, Cuba, Hispaniola, Puerto Rico, Leeward Is, Trinidad-Tobago, tropical SAM S to NE Argentina (exc NW Argentina, N Chile), inc 12 narrow endemics of Brazil; Fatsia 1 of 3 Japan, Korea & Taiwan spp intro Juan Fernandez Is; Heptapleurum ?? of 63 S & Tropical Asia spp intro Bermuda, Bahamas, Hispaniola, Jamaica, Puerto Rico, Leeward & Windward Is; Oreopanax 147 of 147 Mexico & Neo endemic spp, mostly in mountains in the tropics above 1500m in CAM, Antilles (exc Cayman Is, Aruba, Netherlands Antilles), Trinidad-Tobago, Colombia, Venezuela, Guyana, Suriname?, French Guiana, N+E+S Brazil, Ecuador, Peru, Bolivia, NW Argentina; Plerandra 1 of 33 S Pacific Is spp intro Trinidad-Tobago; Polyscias ?? of 173 Old World Tropics & Pacific spp intro El Salvador, Bahamas, Hispaniola, Leeward Is, Puerto Rico, Trinidad-Tobago, Venezuelan Antilles; Raukaua 2 of 6 temperate S Hemisphere spp endemic to N+C Chile + S Argentina (1), C Chile (1); Schefflera ?? of 309 pantropical spp (exc Africa) native in moist habitats at mid to high elevations up to 3000 m in Costa Rica, Panama, tropical SAM S to Peru, Bolivia, C+S Brazil, Hispaniola, Jamaica, Leeward & Windward Is, intro Bahamas, Bermuda; Sciodaphyllum 21 of 21 Neo endemic spp of Jamaica, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia, Venezuela; Tetrapanax  monospecific S China sp intro S Brazil. Hydrocotyloideae: Hydrocotyle ?? of 130 cosmopolitan spp widespread in moist habitats of CAM, Bermuda, Bahamas, Cuba, Hispaniola, Jamaica, Puerto Rico, Leeward & Windward Is, Galapagos, and all of SAM, with high diversity in the Andes.

Patagonia Genera Include:

Aralioideae: Raukaua 2 of 6 temperate S Hemisphere spp endemic to N+C Chile + S Argentina (1), C Chile (1). Hydrocotyloideae: Hydrocotyle ? of 130 cosmopolitan spp native Patagonia inc Falkland Is.

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. This is where all of the family descriptions come from. Below should be most of my references for this, as well as my own personal observations of plants in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., KlitgÃ¥rd, B., & Baracat, A. eds. (2009 onwards). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/ Retrieved Winter 2020–current.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout fall of 2020.
  • Wikipedia: The free encyclopedia. (2004, July 22). FL: Wikimedia Foundation, Inc. Retrieved throughout 2019-current, from https://www.wikipedia.org
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

Plant Families of North America

Welcome to my Plant Families of North America page! I have been absent awhile, but I am back, and this is my focus, teaching how to identify plant families. Over the next few weeks, I will be updating what I have so far. I will be adding new layman’s descriptions in addition to botanical descriptions, and I will be adding more morphology photos, including dissections of flowers where available. Please come back often and see what’s blooming!

A quick overview of basic taxonomy and how I have ordered this site. Plants are given a scientific name composed of two parts, the genus they belong to and the particular species they are. The genus can be thought of like plant cousins, whereas at the species level they are more like siblings. Beyond genus you get the family, which is all of the cousins from all the related families. Above that you get an order, which is like a group of similar families.

Below I have organized the families in their orders so that you can see similarities between related families. When you click on a particular family, you will get the characteristics and morphology pictures that represent that family, along with some of the species I have covered in North America from that family.

Note that most technical words have a hover-over description available, or go to the dictionary of botanical terms to learn more. In no time, you’ll be able to identify plants like a scientist!

Alismatales Order – Monocot Clade

Apiales Order – Eudicot Clade

Caryophyllales Order – Eudicot Clade

Gentianales Order – Eudicot Clade

Lamiales Order – Eudicot Clade

Magnoliales Order – Magnoliids Clade

Piperales Order – Magnoliids Clade

Sapindales Order – Eudicot Clade

Learn how to identify the Anacardiaceae family
Learn how to identify the Anacardiaceae family

Other References

Some other great resources for information on North American species:

iNaturalist https://www.inaturalist.org/

Canadensys Plant Search https://data.canadensys.net/vascan/search

Flora of North America https://eflora.org

iNaturalist Plant Search https://www.inaturalist.org/home

USDA Plants Database https://plants.sc.egov.usda.gov/home

Currently Seeking Funding To Continue This Non-Profit, Ad-Free Work

If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you! (COMING SOON)


How to Identify the Annonaceae or Custard Apple Family

Asimina triloba, American Pawpaw fruit, in flower. Learn how to identify the Annonaceae Family
Asimina triloba, American Pawpaw fruit, in flower
Page Last Updated 2026-04-25

Introduction to the Annonaceae Family

The Annonaceae is an interesting family to learn to identify. They are mostly tropical trees and shrubs with unique flowers and fleshy, often edible and delicious fruits. The most well-known member of the Annonaceae in North America is the pawpaw fruit, or Asimina triloba. It is the most northern member of this family and produces delicious fruits that taste similar to bananas. Learn how to identify the Annonaceae family with morphology photos like the flowers shown in the photo above of Asimina triloba, which are typical for the Annonaceae family, with their 6 petals and 3 sepals and their androecium in a ball in the center.

The Annonaceae family is part of the Magnoliales order in the Magnoliids clade of angiosperms, the third largest clade after dicots (eudicots) and monocots. Occasionally this clade is referred to as a ‘peripheral angiosperm’ because it is neither a dicot nor a monocot, where the vast majority of flowering plants are classified. Instead, this clade is characterized by features of both dicots and monocots, including trimerous flowers (monocot) and branching veins (dicot), as well as pollen with one pore.  

Common Botanical Description

If you’re new to plant morphology, this guide is a perfect beginner’s description to learning to identify the Annonaceae family, with no need to know any scientific jargon. Below this section is additional information on uses and morphology photos to help you identify the family, followed by pictures of individual species found in North America. But for researchers or those wanting to learn a more in-depth version, refer to the Scientific Botanical Description below the images for highly detailed scientific descriptions and genus-level distribution data.

Leaves and Stems: The Annonaceae family is made of woody trees, shrubs, or vines that may be evergreen or deciduous. They are known for their fibrous and aromatic barks containing essential oils and resins.

Leaves are simple with smooth (entire) edges and are arranged alternately along the stem. The leaves do not wrap around or sheath the stem at their base. And while the leaves are generally odorless, some may be aromatic or dotted with tiny glands.

Flowers: The flowers are arranged singly or in branched clusters, are usually symmetrical, and their parts are usually found in multiples of three, similar to most monocots. However, they generally have a distinct outer layer of sepals and an inner layer of petals, which differentiates them from monocots that have no sepals. This is why they are considered “peripheral” angiosperms since they are neither a monocot or a dicot. These parts often total 9 and are arranged in 3 rings (whorls). The petals are often thick and fleshy and sometimes contain visible nectar glands.

Reproductive Features: The male parts are very distinct, with between 25 and 100 stamens that are packed together in a spiral that forms a characteristic dense ball or flat-topped mass in the center of the flower, which is a good characteristic to help identify the family. The ovary is superior, sitting above the point of where the petals attach.

Fruits: Fruits are usually fleshy aggregates of multiple berries that fuse together to form a larger, secondary fruit that is often edible. The seeds inside are sometimes brightly colored.

Uses of Annonaceae 

Many have edible fruits, but they should be eaten with caution, as they contain varying levels of annonacin, which has been implicated in neurodegenerative diseases. Several important tropical commercial fruits come from Annona species (atemoya, cherimoya, custard apple, ilama, sugar apple, sweetsop, and soursop) and Artabotrys.

Medicinal uses include its use as an analgesic and astringent and to treat various conditions, including snakebite, diarrhea, dysentery, arthritis pain, rheumatism, convulsions, neuralgia, and weight loss.

Morphology of Annonaceae in North America

So far in North America, I have only photographed Asimina triloba, since it is the most northern species of this primarily tropical family, but here are some pictures of different morphological aspects of that species.

Learn how to identify the Annonaceae family with morphology photos.

Annonaceae Species I have Covered So Far in North America

So far in North America, I have only photographed Asimina triloba, the morphology photos above show pictures of that species. When I cover more species, I will add more pictures here.

Asimina triloba, American Pawpaw fruit, in flower; the most northern member of the Annonaceae family.

Asimina triloba—American Pawpaw

A large shrub or small tree growing to 11 m with simple, alternate-spirally arranged obovatelanceolate leaves 25 – 30 cm long with a cuneate base, acute tip, and entire margin. Leaves smell similar to green bell peppers if bruised. Flowers are red-purple or maroon, 3 – 5 cm wide, and borne singly on stout, hairy axillary stalks (peduncles), appearing with or before the leaves in the spring. The fruit is a large yellowish to brown berry 5 – 15 cm long and weighing up to 510 g that is edible and sweet. Native to eastern North America. Click here to read my blog on Asimina triloba

Scientific Botanical Description of the Annonaceae

Flowers of the Annonaceae

Plants are usually hermaphrodite or sometimes dioecious and rarely monoecious. The flowers are usually 3-merous. They are axillary and may be singular or found in racemose, compound inflorescences. The receptacle may be elevated, enlarged, or flat. The outer whorls are inserted below the ovaries. Flowers are regular and cyclic or partially acyclic. Sometimes the androecium is spiraled (acyclic). Free hypanthium is absent, and a hypogynous disk is present. The perianth has a distinct calyx and corolla that may be petaline or made of tepals; however, sometimes it can be difficult to determine when the outer 1-2 whorls are sepaloid. The perianth has 9 (10–12) parts, which may be basally joined or free, is usually 3-whorled, is isomerous, and may be similar or different in the whorls. The calyx has 3 or 6 parts, typically 2 whorled and free (sometimes basally connate), and the lobes are longer than the tube. The corolla has 3 or 6 parts, is 1–2 whorled, is free (sometimes basally connate), and is imbricate or valvate. They may have 6 petals in 2 unequal whorls of 3 with larger outer whorls and fleshier inner whorls that might share the same nectar glands or 6 to 15 petals with impressed veins on the inner face (Annona muricata)

Androecium of the Annonaceae

The androecium contains 25–100 (usually many) members. Androecial members usually mature centripetally, are free of the perianth and each other, are all equal, are usually spiraled, and are rarely 3 or 6 whorled. The androecium is usually made entirely of fertile stamens but may include staminodes (e.g. in Uvaria). Staminodes, when present, are external to the fertile stamens and are non-petaloid. Stamens are inserted below the ovary and arranged spirally, forming a characteristic ball or flat-topped mass of short, stout filaments and linear to oblong anthers that face upwards. Anthers are adnate, non-versatile, dehisce via longitudinal slits or longitudinal valves, are extrorse, are tetrasporangiate, and are appendaged via expansion of the connective.

Gynoecium of the Annonaceae

The gynoecium is 10–100(+) carpeled. When syncarpous, the pistil is 1- or 2- to 15+-celled. It is usually apocarpous (carpels spiraled or cyclic) or synstylovarious to syncarpous (e.g., Monodora); the ovary is superior. The carpel is 1–10-ovuled. Placentation of free carpels is basal. The ovary, when syncarpous, is 1 or 2–15+ locular. Stigmas are wet type and papillate and Group III type. Placentation, when unilocular, is parietal or basal; when plurilocular, it is basal. Ovules in the single cavity when unilocular are 1–50, with 10–50 per locule. Ovules are ascending and apotropous, have a ventral raphe, may be arillate or not, and are anatropous, bitegmic, and crassinucellate.  

Fruit of the Annonaceae

The fruit is a fleshy aggregate that is often made of berries. Fruiting carpels may coalesce into a secondary syncarp or they may not. The fruiting carpel is indehiscent and baccate. Seeds are endospermic with 1 to many per pistil and are often brightly colored. The endosperm is ruminate, oily, and has amyloid.

Habit & Leaf Form of the Annonaceae

Trees, shrubs, or lianas that are deciduous or evergreen and produce essential oils and may be resinous. The inner bark is typically fibrous and aromatic, and the pith is septate to diaphragmed. Plants may be self-supporting or climbing. When climbing, they may be scrambling or using stem or petiole twining. Branching is distichous or spiral.

Leaves are typically arranged alternately, are non-sheathing, simple, petiolate, and may or may not be gland-dotted or aromatic. Lamina is entire, pinnately veined (sometimes palmately), and cross-venulate. Leaves are exstipulate. Lamina margins are entire. Domatia occurs in three genera as pockets or occasionally as hair tufts.

Taxonomy of Annonaceae

The Annonaceae family has 2503 species in 108 genera. They are part of the Magnoliales order in the Magnoliids clade of the peripheral (not a true dicot or monocot) angiosperms.

The family is divided into 4 subfamilies as follows:

  1. Anaxagoreoideae is a group of 25 species in a single genus found in tropical America and the tropics in and around Indonesia (absent from tropical Africa). They have 2-ranked trunk leaves, sessile stigmas, and dry follicles that are explosively dehiscent.
  2. Ambavioideae is a more widespread group found throughout the tropics and into temperate eastern North America. This subfamily is characterized by a truncate and dilated anther connective, an intine that does not extrude through the aperture, and ovules with a middle integument.
  3. Annonoideae is another mostly tropical subfamily that is rarely found in temperate areas. It is also characterized by an intine that does not extrude through the aperture but can be differentiated based on having a lamelliform ruminate endosperm.
  4. Malmeoideae is restricted to the lowland tropics and is characterized by a glass-like endosperm with ruminations spiniform.

Genera:

Anaxagoreoideae: Anaxagorea (25).

Ambavioideae: Cananga (4), Cleistopholis (3), Cyathocalyx (9), Drepananthus (27), Lettowianthus (1), Mezzettia (4), Tetrameranthus (8).

Annonoideae: Afroguatteria (2), Annona (171), Anonidium (4), Artabotrys (108), Asimina (9), Asteranthe (2), Bocagea (2), Boutiquea (1), Cardiopetalum (3), Cleistochlamys (1), Cymbopetalum (27), Dasymaschalon (27), Dennettia (??), Desmos (18), Diclinanona (3), Dielsiothamnus (1), Disepalum (9), Duckeanthus (1), Duguetia (95), Fissistigma (58), Friesodielsia (38), Froesiodendron (3), Fusaea (3), Goniothalamus (134), Guatteria (184), Hexalobus (5), Hornschuchia (10), Isolona (20), Letestudoxa (3), Lukea (??), Mischogyne (5), Monocyclanthus (1), Monodora (15), Neostenanthera (5), Ophrypetalum (1), Porcelia (7), Pseudartabotrys (1), Pyramidanthe (9), Sanrafaelia (1), Sphaerocoryne (7), Toussaintia (4), Trigynaea (9), Uvaria (168), Uvariastrum (5), Uvariodendron (15), Uvariopsis (10?), and Xylopia (109).

Malmeoideae: Alphonsea (36), Annickia (11), Bocageopsis (4), Brieya (2), Cremastosperma (31), Dendrokingstonia (3), Desmopsis (39), Ephedranthus (7), Fenerivia (5), Greenwayodendron (6), Huberantha (34), Klarobelia (13), Leoheo (1), Maasia (6), Malmea (6), Marsypopetalum (5), Meiocarpidium (1), Meiogyne (32), Miliusa (59), Mitrephora (49), Mkilua (1), Monanthotaxis (77), Monocarpia (4), Monoön (72), Mosannona (14), Mwasumbia (1), Neo-uvaria (7), Onychopetalum (2), Orophea (60), Oxandra (28), Phaeanthus (8), Phoenicanthus (2), Piptostigma (13), Platymitra (2), Polyalthia (92), Polyalthiopsis (1), Polyceratocarpus (10), Popowia (32), Pseudephedranthus (2), Pseudomalmea (4), Pseudoxandra (24), Pseuduvaria (58), Ruizodendron (1), Sageraea (9), Sapranthus (7), Sirdavidia (1), Stelechocarpus (3), Tridimeris (2), Trivalvaria (8), Unonopsis (48), Wangia (2), and uodendron (1).

Key Differences From Similar Families

The Annonaceae is easily distinguished from other Magnoliales by its fibrous and aromatic bark. Myristicaceae can be differentiated by their red latex. Magnoliaceae can be differentiated by their large deciduous stipules.  

Distribution of Annonaceae

Mostly subtropical and tropical, rarely temperate. Widespread, especially in the Old World. In the Americas, it is found all over the Neotropics and north as far as southern Canada. It is particularly common in lowland forested areas. 

Distribution of Annonaceae in the Americas

Canada Genera:

Asimina 1 of 9 E NAM endemic spp native ON.    

USA Genera:

Annona 3 of 171 Americas + Africa spp native FL; Artabotrys 1 of 108 Old World Tropics spp intro HI; Asimina 9 of 9 E NAM endemic spp native E USA from ND S to TX and all states E exc ND, SD, MN, CT, RI, VT, NH, ME, inc 8 spp endemic to E USA; Deeringothamnus monospecific endemic of FL; Polyalthia 1 of 92 Australasian spp intro FL. 

Mexico Genera:

Anaxagorea 1 of 25 Americas + tropical Asia spp native SW mexico, Ver; Annona (inc Rollinia) 14 of 171 Americas + Africa spp inc 12 native all through Mexico and 1 sp intro Sin E to Tam and S to Oax, Chp, QR, Yuc; Cananga 1 of 2 tropical Asia spp intro SW Mexico, Ver; Cymbopetalum 2 of 27 Mexico + Neo endemic spp native Chp, Ver, Oax?; Desmopsis 8 of 23 Mexico + N Neo endemic spp native Gro, Mic, Chp, Oax, Ver inc 6 narrow endemics of Ver (2), Mic (1), Gro (1), Oax + Ver (2); Guatteria 2 ?? of 184 Neo endemic spp spp native SW+SE+S Mexico, Ver; Mosannona 1 of 14 Mexico + Neo endemic spp native Cam, Chp, QR, Ver, Yuc; Oxandra 4 of 28 Mexico + Neo endemic spp native SW+C+SE Mexico, Ver, inc 1 narrow endemic of Chp; Sapranthus 3 of 7 Mexico + Neo endemic spp native Gro, Oax, QR, Sin, Chp, Ver, inc 1 narrow endemic of Chp; Stenanona 8 of 16 Mexico + N Neo endemic spp native rainforests of SW+SE Mexico, Ver inc 7 narrow endemics of Chp (3), Ver + Oax (2), Tab + Chp (1), Ver + Oax + Chp (1); Tridimeris 2 of 2 Mexican endemic spp native E+C Mexico inc 1 narrow endemic of Chp; Unonopsis ?? of 48 Neo endemic spp native SW+SE Mexico; Uvaria 1 of 168 Old World Tropics spp intro Jal; Xylopia 1 of 169 pantropical spp native S Chi.

Neotropical Genera:

Anaxagorea 22 of 25 Americas + tropical Asia spp native CAM (exc El Salvador), Trinidad-Tobago, tropical SAM S to Peru, Bolivia, C+SE Brazil, inc 5 narrow endemics of Costa Rica & Panama (1), S Venezuela (2), N Peru (1), SE Brazil (1); Annona ?? of 171 spp mostly Americas and some tropical Africa spp native and widely distributed and cultivated for fruit, throughout CAM, Bahamas, Antilles, SW Caribbean, tropical SAM S to N Argentina (exc N Chile); Bocagea 2 of 2 spp endemic to E Brazil; Bocageopsis 4 of 4 SAM endemic spp native Colombia, Venezuela, Guyana, Suriname, French Guiana, N+C+S Brazil, Peru, Bolivia; Cananga 1 of 2 tropical Asia spp intro Guatemala, Nicaragua, Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, Trinidad-Tobago; Cardiopetalum 3 of 3 N SAM endemic spp of Suriname, French Guiana, N+C+E Brazil, Peru, Bolivia; Cremastosperma 34 of 34 Neo endemic spp native Costa Rica, Panama, Colombia, Venezuela, N+C Brazil, Ecuador, Peru, Bolivia, found in lowland or pre-montane forests, most diversity in the narrow tropical zone W of the Andes; Cymbopetalum 27 of 27 Mexico + Neo endemic spp native from S Mexico S through CAM and tropical SAM S to Peru, Bolivia, C+E Brazil; Desmopsis 17 of 23 S Mexico + N Neo endemic spp native CAM, Cuba, Colombia; Desmos 1 of 18 Australasian spp intro C+E Brazil, Trinidad-Tobago; Diclinanona 3 of 3 N SAM endemic spp native N+C Brazil, Colombia, Peru, Venezuela; Duckeanthus monospecific narrow endemic of N Brazil; Duguetia 91 of 95 Neo + Africa spp native Nicaragua S through tropical SAM S to Peru, Bolivia, Paraguay, S Brazil, 4 spp endemic to W Africa; Ephedranthus 7 of 7 SAM endemic spp native tropical SAM S to Peru, Bolivia, Paraguay, SE Brazil (exc Ecuador); Froesiodendron 3 of 3 N SAM endemic spp native Colombia, Peru, N Brazil; Fusaea 3 of 3 N SAM endemic spp native N+NE+C Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela, Bolivia; Guatteria 184 of 184 Neo endemic spp native from S Mexico S through CAM (exc El Salvador), Cuba, Hispaniola, Puerto Rico, Leeward & Windward Is, tropical SAM S to Peru, Bolivia, C+S Brazil, a dominant genus in SAM mature forests; Hornschuchia 10 of 10 narrow endemic spp of E Brazil; Klarobelia 13 of 13 Neo endemic spp native from Costa Rica S to Colombia, Venezuela, N Brazil, Ecuador, Peru, Bolivia, inc several endemics to Ecuador; Malmea 6 of 6 Neo endemic spp native Panama, Colombia, Guyana, Suriname, N+NE Brazil, Peru; Monodora 1 of 15 African spp intro Trinidad-Tobago; Monoon 1 of 72 Australasian spp intro Trinidad-Tobago; Mosannona 14 of 14 Mexico + Neo endemic spp native Guatemala, Belize, Honduras, Costa Rica, Panama, Colombia, N Brazil, Guyana, Suriname, Ecuador, Peru, Bolivia, common in rainforests around Andes, inc 6 narrow endemics of Costa Rica (1), Barro Colorado Is Panama (1), Guatemala (1), Suriname + Guyana (1), Ecuador (1); Onychopetalum 2 of 2 Neo endemic spp of Venezuela, N+C Brazil, Peru, Bolivia; Oxandra 28 of 28 Mexico + Neo endemic spp native S Mexico, Guatemala, Belize, Nicaragua, Costa Rica, Panama, Greater Antilles (exc Cayman Is), Leeward + Windward Is, SW Caribbean, tropical SAM S to Peru, Bolivia, C+SE Brazil, inc 9 narrow endemics of Colombia (3), Suriname (1), Guyana (1), Brazil (4); Polyalthia 1 of 92 SE Asia + Australasia spp intro Trinidad-Tobago; Porcelia 7 of 7 Neo endemic spp native Panama, Colombia, Venezuela, N+E+S Brazil, Ecuador, Peru Bolivia; Pseudephedranthus 2 of 2 N SAM endemic spp of Venezuela, Guyana, Suriname, N Brazil; Pseudomalmea 4 of 4 N SAM endemic spp of Colombia, Venezuela, N Brazil, Ecuador, Peru, Bolivia; Pseudoxandra 24 of 24 SAM endemic spp native Colombia, Venezuela, Guyana, French Guiana, N+C Brazil, Peru, Bolivia; Ruizodendron monospecific N SAM endemic of Colombia, N Brazil, Ecuador, Peru, Bolivia; Sapranthus 6 of 7 Mexico + N Neo endemic spp native CAM, Colombia, inc 1 narrow endemic of Cesar Colombia; Stenanona 9 of 16 Mexico + N Neo endemic spp native rainforests of CAM (exc El Salvador), Colombia, inc 6 narrow endemics of Honduras (2), Costa Rica (1), Panama (1), Colombia (2), the rest are all narrow endemics of Mexico; Tetrameranthus 8 of 8 N SAM endemic spp of N Brazil, Colombia, Venezuela, French Guiana, Ecuador, Peru; Trigynaea 9 of 9 N SAM endemic spp native Colombia, Venezuela, Guyana, Suriname, N+E Brazil, Ecuador, Peru, Bolivia; Unonopsis 48 of 48 Neo endemic spp native S Mexico, CAM (exc El Salvador), tropical SAM S to Peru, Bolivia, C+S Brazil; Uvaria 1 of 168 Old World Tropics spp intro Trinidad-Tobago; Xylopia 40 of 169 pantropical spp native CAM, Cuba, Jamaica, Trinidad-Tobago, tropical SAM S to Peru, Bolivia, NE Argentina. 

Patagonia Genera:

Absent 

Additional Information and References

  • Visit Lyrae’s Dictionary of Botanical Terms to learn the terminology of botanists. Note that if you hover over most of the words in the articles, you can also get definitions from them there.
  • Willis, Lyrae (Unpublished). Plant Families of North America. Below should be most of my references for this along with my own personal observations in North America.
  • Canadensys: Acadia University, Université de Montréal Biodiversity Centre, University of Toronto Mississauga, University of British Columbia. http://data.canadensys.net/explorer (accessed 2020 – current)
  • Cruz-Chacon, Ivan de la Marisol Castro-Moreno, Lorena Mercedes Luna-Cazares, and Alma Rosa Gonzalez-Esquinca (2016). La Familia Annonaceae Juss. en México. Lacandonia, year 10, vol. 10, num. 2: 71-82, December 2016.
  • Delta: Watson, L., and Dallwitz, M.J. (1992+). The Families of Flowering Plants: descriptions, illustrations, identification, and information retrieval. Version: 2nd May 2020. delta-intkey.com. Accessed spring through fall of 2020.
  • GBIF.org (2020), GBIF Home Page. Available from: https://www.gbif.org
  • Naturalista: CONABIO http://www.naturalista.mx (Accessed 2020–current).
  • Neotropikey: Milliken, W., KlitgÃ¥rd, B. & Baracat, A. eds. (2009+). Neotropikey: Interactive key and information resources for flowering plants of the Neotropics. www.kew.org/neotropikey.com (accessed 2020 – current).
  • Patagonia Wildflowers: Wildflower Identification Site. https://patagoniawildflowers.org/ Accessed throughout the fall of 2020.
  • POWO (2019). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet: http://www.plantsoftheworldonline.org/ Retrieved Winter 2020-current.
  • USDA, NRCS. 2020. The PLANTS Database (http://plants.usda.gov, 2 June 2020). National Plant Data Team, Greensboro, NC, USA; accessed throughout the fall of 2020.
  • WFO (2022): World Flora Online. Published on the Internet: http://www.worldfloraonline.org. Accessed Spring 2022 – current

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Cypripedium acaule Pink Lady's Slipper Orchid

Cypripedium acaule the Pink Lady Slipper or Moccasin Flower - photo from Allatoona Creek, Georgia by Lyrae Willis
Cypripedium acaule the Pink Lady Slipper or Moccasin Flower – photo from Allatoona Creek, Georgia by Lyrae Willis

Pink Lady Slipper or Moccasin Flower Cypripedium acaule

Introduction

These gorgeous orchids are always a lovely treat when you find them growing in the forest. Fortunately, they are not excessively rare, though they are not all that common either. Cypripedium acaule is part of the Cypripedioideae subfamily in the Orchidaceae family. It goes by two common names the Pink Lady Slipper Orchid or the Moccasin Flower after their slipper or moccasin-shaped flowers that are unique to their genus. Orchids everywhere are declining in population due to habitat loss, climate change, and highly specific pollination with poor overall pollination rates. If you do find these please never pick the beautiful flowers. Instead, leave them there to hopefully be pollinated and produce the next generation. Take out your phone and take pictures of the beautiful flowers instead. You can put them on your computer desktop or print them and frame them for your wall and enjoy them for many years.

Description of Cypripedium acaule

Stem & Leaves

Pink Lady Slippers are erect herbaceous perennials 15 – 60 cm tall with solitary flowers on a leafless peduncle (flower stalk).

Two leaves grow opposite from each other directly from the rhizome. They are ovate or broadly elliptic to oblong in shape and measure 9 – 30 cm long and 2.5 – 15 cm wide. Rarely plants will only have one leaf.

Young leaves of Cypripedium acaule; photo from Allatoona Creek, Georgia, USA
Young leaves of Cypripedium acaule; photo from Allatoona Creek, Georgia, USA
Flower closeup of Cypripedium acaule showing the vertical slit in the pouch; Allatoona Creek, GA, USA
Flower closeup of Cypripedium acaule showing the vertical slit in the pouch; Allatoona Creek, GA, USA

Flowers & Fruits

The showy solitary flowers bloom from April to July. They have tepals (sepals and petals) that are yellowish-brown, reddish-brown, pinkish, or even green. They are deflexed or spreading and may be slightly twisted, and are 1.9 – 5.2 cm long. The lip (labellum) is located below the tepals and is shaped like a large pouch with a slit down the middle, slipper or moccasin-like. It is usually a light pink but occasionally may be magenta and is obovoid in shape, 3 – 6.7 cm long.

Toxicity

The Cypripedium genus all have glandular hairs on the leaves and stems. The compound in the hairs can cause contact dermatitis similar to poison ivy.

Similar Species Frequently Confused With

It is difficult to confuse Cypripedium acaule for most other genera due to the unique moccasin-like lip on the flower. Sometimes people do mistake flowers of other genera, but if you just look for the distinctive labellum or lip that is shaped like a pouch with a slit down the middle, you can tell for certain it is a Cypripedium. So, most misidentifications are with other members of the same genus. Other Cypripedium species can be differentiated as follows:

  • Cypripedium reginae Showy Lady Slipper has a more limited range from Saskatchewan east to Newfoundland Island in Canada and the northern US from Minnesota south to Arkansas and east to Maine, but mostly concentrated in the states that border Canada. It is taller, growing to 100 cm, and usually sprouts many stems from the same rhizome. Showy Lady Slipper has more leaves, with 3 – 5 leaves per stem. It also has larger and much more showy white tepals and usually a brighter pink or magenta lip.
  • Cypripedium parviflorum Yellow Lady Slipper is another largely eastern North American species that extends to the Canadian Rockies and Arctic in the west and the Rocky Mountains in the US states. Otherwise, it has a very similar eastern range as Cypripedium reginae. It is fairly easy to tell apart, however, since its lip is always yellow.
  • Cypripedium candidum Small White Lady Slipper has a limited range and is listed as a Vulnerable species globally. It is rare and endangered in Canada but can be found in the extreme southern parts of eastern Canada from Saskatchewan east to Ontario. In the US, it is also rare and vulnerable and can be found from North Dakota south to Nebraska and east to New York, with some also found from Missouri south to Alabama. It is a much smaller plant, only up to 36 cm tall, and has a smaller lip that is always white.
  • Cypripedium montanum Mountain Lady Slipper is another rare vulnerable orchid that is found in the Pacific Northwest from Alaska south to northern California and east to the Alberta Rocky Mountains in Canada and Montana in the US. It is typically found only at high elevations in open forests and subalpine slopes. It grows to 70 cm tall and has 1 – 3 large flowers on top of its stem, and its pouch is always white.

Distribution of Cypripedium acaule

Moccasin Flower is a central and eastern Canadian and eastern USA species. It is not found anywhere else in the world.

In Canada, Cypripedium acaule is found in Alberta, Saskatchewan, Manitoba, Ontario, Quebec, New Brunswick, Nova Scotia, Prince Edward Island, and Newfoundland Island (not Labrador). It may also be present in the Northwest Territories.

In the USA, Pink Lady Slipper is found in Minnesota, Wisconsin, Illinois, Michigan, Indiana, Michigan, Indiana, Ohio, Kentucky, Tennessee, Alabama, Georgia, North & South Carolina, Virginia, West Virginia, Pennsylvania, Delaware, Maryland, Washington DC, New York, New Jersey, Connecticut, Rhode Island, Massachusetts, Vermont, New Hampshire, and Maine.

Habitat & Growing Conditions of Cypripedium acaule

Cypripedium acaule tolerates a wide range of shade and moisture conditions being found in dry to wet forests, shrublands, and slopes, but it prefers partial shade and well-drained slopes. However, it requires highly acidic soils, which is probably why it is often found in pine forests but can be seen in deciduous and mixed woods as well. It is generally found below 1200 m in elevation.

Growing Pink Lady Slipper in Your Garden

Cypripedium acaule is challenging but not impossible to grow yourself if you don’t already have a highly acidic or coniferous forest to plant them in. For a long time, it was believed that it could not be cultivated due to a belief that a fungal association found in the forest was required for germination and growth. However, Pink Lady Slipper is available commercially but is not easy to find as it is more difficult to grow than other species. You will not be able to grow it from seed as special conditions and fungal associations are still required for germination, but once it has true leaves, it can be moved into a garden.

To grow commercially purchased plants of Cypripedium acaule, the only real critical factor is soil pH. It absolutely must be kept below 5.0 to prevent the plants from rotting from the activity of soil microbes that increase above pH 5.0. Anywhere from 4.0 – 4.5 pH is ideal for Pink Lady Slipper. To keep the pH so low, you must grow them in a container so that you can better control the conditions. Use a significant amount of peat moss or pine duff in the soil and use pH-neutral perlite to improve drainage. Tap water should never be used to water them due to the risk of Calcium in the water that would raise the pH. Instead, use rainwater or distilled water mixed with 2 oz of vinegar per gallon.

Other things to consider are where to place the pots. Ideally, morning sun or dappled sunlight under a tree is best. And be sure to sink the pots into the ground in winter or store them in a cold frame.

Wildlife Values of Cypripedium acaule

Native bees routinely visit the flowers.

Status of Cypripedium acaule

Pink Lady Slipper is currently listed as Least Concern on the IUCN Red List. However, as with most woodland plants, the population of mature individuals being found is decreasing.

NatureServe lists Moccasin Flower as Globally Secure, G5.

In Canada, Pink Lady Slipper is considered Locally Secure S5 in Ontario, Quebec, New Brunswick, Nova Scotia, and Prince Edward Island. It is Apparently Secure S4 in Saskatchewan and Newfoundland Island. In Alberta and Manitoba, it is Vulnerable S3.

In the USA, Cypripedium acaule is considered Locally Secure S5 in West Virginia, Virginia, and North Carolina. It is considered Apparently Secure S4 in Kentucky, Tennessee, New York, New Jersey, Delaware, Connecticut, and Georgia. In Alabama, it is considered Vulnerable S3. It is considered Critically Imperiled S1 in Illinois and Indiana. In all other states where it is found, its status is not yet determined.

Traditional or Other Uses of Pink Lady Slipper

Cypripedium acaule Medicinal Uses

Pink Lady Slipper roots were used by the Algonquin for stomachaches, menstrual disorders, kidney and urinary infections, and venereal diseases. Cherokee used the root as an analgesic, anticonvulsive, to treat colds, flu, stomachaches, kidney problems, worms, and ‘female trouble’. Iroquois used them in a decoction as an analgesic and in a poultice for bite wounds. Menominee used it to treat urinary problems and ‘male disorders’. Micmac and Penobscot used the root as a sedative for nervousness. Rappahannock used the dried roots in whiskey as a panacea for general ailments.

Pink Lady Slipper as an Ornamental

Due to the challenges of growing it this plant is only occasionally grown as an ornamental. But for the most serious and dedicated gardeners or orchid lovers, it is a lovely addition to any woodland garden.

Ethical Wildcrafting of Cypripedium acaule

Check the status in your state before harvesting since it is vulnerable in some areas. See the above section on Status. Alternatively, grow it in your garden but do not attempt to harvest wild plants to transplant to your garden as, most likely, you will simply be killing the plant. Instead, buy commercially available plants that have been germinated in an orchid lab.

Generally speaking, I do not recommend wildcrafting this plant in any form. Transplanting has a poor survival rate, and the only other reason for harvesting is the root to be used medicinally. If using this root is part of your traditional heritage, and you want to harvest some, always use the 1 in 20 rule of Ethical Wildcrafting. Harvest the roots of only one in every plant you see in an area where they are locally common.

Wildcrafting and Processing

Use a digging stick to loosen the roots from the soil. Picked roots can be placed in a basket, bowl, or paper bag and brought home for processing. If you are harvesting multiple products on the same day, be sure to label the roots in a paper bag so that you do not confuse different plants.

Roots should be brushed clean of any dirt and then chopped into more manageable pieces before drying. Dried roots are notoriously difficult to cut into smaller pieces once dried.

Once dried, the roots can be stored in a jar for later use. Label your jar with the species name and the date, and I also usually add the location of the harvest for my own reference. Do not grind or crush the leaves or roots until you are ready to use them. This will keep them as fresh as possible to preserve their medicinal properties. When you pre-grind, even if stored in glass jars, this increases the oxidation rate and rapidly degrades the medicinal properties reducing their useful shelf-life.

References and Resources

Canadensys Plant Search https://data.canadensys.net/vascan/search

Dictionary of Botanical Terms – by Lyrae’s Nature Blog https://lyraenatureblog.com/blog/dictionary-of-botanical-terms/

Flora of North America https://eflora.org

iNaturalist Plant Search https://www.inaturalist.org/home

IUCN Red List https://www.iucnredlist.org/

Lady Bird Johnson Wildflower Center https://www.wildflower.org

Native American Ethnobotany http://naeb.brit.org/

NatureServe Explorer https://explorer.natureserve.org/Search

USDA Plants Database https://plants.sc.egov.usda.gov/home

Willis, Lyrae (? Not yet published).  Plant Families of North America. 

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If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you!


Carduus nutans Musk Thistle - Invasive Species of North America

Carduus nutans floral bug from Knoxville, TN. Photo by Lyrae Willis 2021.
Carduus nutans floral bug from Knoxville, TN. Photo by Lyrae Willis 2021.

Introduction

Carduus nutans also known as Musk Thistle or Nodding thistle is a widespread invasive species found throughout most of southern Canada and almost the entire USA. It is a herbaceous biennial plant in the Carduoideae subfamily of the Asteraceae (Sunflower) family. While in cooler climates it typically takes 2 years to produce seeds before the plant dies in warmer climates it may grow and go to seed in a single season. It is frequently found on roadsides, in pastures, meadows, empty lots, disturbed land, and waste places throughout its range.

Description of Carduus nutans

Leaves & Stems

These biennial plants at maturity will reach 150 – 250 cm tall and typically have multi-branched stems. Occasionally single-stem specimens can be found under poorer conditions or in areas with lots of competition. The stems are covered with spines and a woolly covering of hair with occasional small leaves. Stems and leaves sprout from a stout taproot up to 40 cm long.

Leaves develop from a basal rosette and reach 40 cm long when mature. When young the leaves are somewhat lobed and slightly pubescent hairy and more or less upright but as they mature they become prostrate and more heavily pinnately lobed and have prominent sharp yellowish to whitish spines at the lobe margins. The surface is dark green, waxy, and pubescent (hairy) on top with woolly veins on the lower surface.

Flowers & Fruits

Carduus nutans flower from the Lincoln National Forest, NM. Photo by Lyrae Willis 2022.
Carduus nutans flower from the Lincoln National Forest, NM. Photo by Lyrae Willis 2022.

Flowers form in large globose heads made of hundreds of tiny but showy reddish-purple disk flowers. Rarely, white variants can be seen. Flower heads are 3 – 5(-7) cm in diameter and form at the end of the stems. Upright immature heads begin to droop as they mature at 90° to 120° from the stem. Heads are surrounded by woolly spiny tipped phyllaries that often are purplish. Outer phyllaries reflex at maturity while the rest of them are constricted in the middle with blades that are narrower than their bases.

Fruits are cypselae which are typical for the Asteraceae family. They consist of a small angled seed (frequently mistaken for an achene) 3 – 4.5 mm long that is gold, fawn, or brown with fine wrinkles and a white pappus of numerous hairs up to 2 cm long. These fruits are suitable for dispersal by wind due to the small light seeds and light pappus that catch wind currents. Each flower head produces hundreds up to 1200 cypselae.

Similar Species Frequently Confused With

There are many thistle species in North America that Carduus nutans may be confused with. Some are native and some are introduced. Following is a list of the most similar-looking species and how you can differentiate them.

In the Carduus genus we have:

  • Carduus acanthoides the Broad-Winged Thistle found in the eastern USA and Canada, central USA, and south-western Canada. Native to Eurasia it has similar leaves and growth habits but the flower heads are smaller and not globose or nodding like Carduus nutans. Instead, its mature flowers are more or less erect, 1.3 – 2.5 cm across and its achenes have a pappus that is 1.1 – 1.3 cm long. Hybrids between the two species have been observed in some of the eastern US states which may make identification difficult. However, since no Carduus species are native to North America this should not be a problem for control purposes as long as it is identified to the genus level.

Most of the other less closely-related species that also look similar are in the Cirsium genus. They can be differentiated as follows:

  • Cirsium vulgare the Bull Thistle is also Eurasian and occupies almost the exact same range as Musk Thistle but extends further into Mexico and in the US is found in Vermont and Maine as well. It is fairly easy to tell apart, however, by its much more bulbous-shaped receptacle and smaller phyllaries that have many more layers and are more appressed to the receptacle. The flower heads are less rounded and generally a bit smaller than Nodding Thistle. Leaves are more of a grey-green and are generally smaller to only 30 cm.
  • Cirsium arvense the Creeping Thistle is native to Eurasia and is now found throughout the northern half of the USA and most of southern Canada and some in the northern Territories and eastern Alaska. It tends to grow in clonal colonies by sending up shoots that grow into plants on more slender stems to only 150 cm tall that are smooth and hairless and generally not spiny. The spiny leaves are smaller only 20 cm long. Flower heads are also much smaller only 2.2 cm wide which grow in clusters on the tops of stems instead of singly.
  • Cirsium horridulum the Bristle Thistle is a native plant found in the eastern US from New England south to Florida and west to Texas and Oklahoma as well as parts of Mexico and eastern Central America. It grows to similar heights and has leaves of similar sizes but the often purple-tinged leaves have thicker spines that are more densely placed on the leaves and up the stem. Flower heads are also much spinier and disk flowers may be pink, reddish, white, or even yellow.
  • Cirsium pumilum the Pasture Thistle is native to the north-central and northeastern USA as well as in Ontario, Canada. It only grows to 100 cm tall and its leaves are shorter to 30 cm long. It has a much longer narrowly egg-shaped receptacle with more rows of phyllaries that are not reflexed. Flowers may be pink, purple, or white and flower heads are less dense generally containing fewer than 100 disk flowers.
  • Cirsium pumilum the Wavy Leaf Thistle is native to most of western North America from BC east to Manitoba, Canada, and south to Durango, Mexico. It grows to similar heights but is typically less branched, if at all. Its leaves are much more undulated, more shallowly lobed, and grey rather than dark green. The flower heads are of similar size but generally much lighter pink or even white and the receptacle is much more elongated with more appressed phyllaries.
  • Cirsium ochrocentrum the Yellow Spine Thistle is native to the Central Plains and western deserts of the US and northern Mexico. It grows to only 100 cm tall and its deeply lobed and spiny leaves only grow to 25 cm long. Its phyllaries are appressed to the receptacle and they are tipped with yellow spines. Flower heads contain white, pink, or lavender disk flowers.
  • Cirsium texanum the Texas Thistle is native to the south-central US and northern Mexico. It is much smaller growing to only 80 cm tall and it has light purple or light pink disk flowers. Its phyllaries are not reflexed and are green and white and tipped with spines. The leaves have fewer spines and are dark green above but woolly white below.
  • Cirsium discolor the Field Thistle is native to central and eastern Canada and the US. It grows to 200 cm tall but can easily be distinguished by its always solitary stem and its leaves that are green above but white and woolly below.
  • Cirsium edule the Edible Thistle is native to the Pacific Northwest from Alaska south to Oregon and east to Idaho. Along with its narrow range, it can easily be distinguished by its spiny reflexed arachnoid phyllaries (covered with a mass of downy white hair).
  • Silybum marianum the Milk Thistle is native to Eurasia but can be found in North America mostly in the coastal states on both the Atlantic and Pacific side with fewer plants in the central regions though they are still present. The easiest way to tell this one apart is by its distinctly variegated leaves mottled in pale green and white. To learn more about this invasive species check out my blog at https://lyraenatureblog.com/blog/milk-thistle-silybum-marianum/

Native Distribution of Carduus nutans

Musk or Nodding Thistle was originally native to much of Europe and western Asia as well as northern and eastern Africa where it grew in fields, meadows, and disturbed habitats.

Habitat Types Where Carduus nutans is Found

Carduus nutans typically grow in meadows and grasslands and are often found in heavily grazed pastures. Stout taproots allow it to easily survive in pastures as it can be more resistant to drought than other pasture plants. It is also found in any disturbed open soil such as roadsides, cleared land, and waste sites. Areas subject to natural disturbances such as landslides and flooding are also suitable habitats.

Carduus nutans prefers open sun and will not grow in excessively wet or dry areas or shady sites such as under the forest canopy. It tolerates neutral or acidic soils and grows from sea level to 2500 m elevation.

Human Uses of Musk Thistle

The pith of the stem can be boiled and eaten like asparagus and is said to have a pleasant taste.

Medicinally the flowers are sometimes used to lower fevers and as a blood purifier. In India studies done with the linoleic acid in the seed’s oil have been shown to help prevent atherosclerosis.

Distribution of Carduus nutans in North America

The species was first accidentally brought to eastern North America in the mid-1800s likely in a ship’s ballast water. Since then it has become very widespread and is found in most US states and most of southern Canada.

In Canada, Carduus nutans have been recorded in most of the southern provinces including British Columbia, Alberta, Saskatchewan, Manitoba, Ontario, Quebec, New Brunswick, and Nova Scotia. Its status in Newfoundland is uncertain and it has not been reported in Prince Edward Island or Labrador nor in any of the Arctic territories.

In the USA, Musk Thistle is found in most of the continental USA excluding only Vermont, Maine, and Florida. It is also not currently found in Alaska or Hawaii.

In Mexico, Carduus nutans so far have only been reported in Mexico City and Mexico State. Given its proximity to the northern border, and the as-of-yet under-reporting of invasive species in Mexico this information will change in the near future.

Musk Thistle has been introduced on every continent except for Antarctica. It has been declared a noxious weed in Canada, the USA, Australia, New Zealand, and South Africa.

How Carduus nutans Spreads

Propagation is strictly by seed. Seeds remain viable for only a couple of years on the soil surface. However, if seeds are buried 20 cm they can last up to 80 years in the soil seed bank.

It is currently primarily spread through long distances as a contaminant in uncertified seeds sold out of the country. Within the same country, long-distance dispersal also results from seeds carried as a contaminant in hay as well as on equipment and vehicles.

Short-distance dispersal occurs through seeds dispersed short distances by wind or transported on vehicles, equipment, etc.

Habitats at Risk of Invasion in North America

Carduus nutans prefer meadows, fields, pastures, disturbed ground, farmland, and any open land with good exposure to sunlight. Since it grows up to 2500 m elevation all open areas to this elevation are at risk. Mature forests, permanent wetlands, and deserts are not at risk as they will not grow in the shade or in permanently wet or excessively dry soil.

Impacts of Invasion

In pastures and farmlands, Carduus nutans can negatively impact production by suppressing the growth of more desirable plants through competition. Carduus nutans plants do well in pastures because of the deep thick taproots that allow them to outcompete grasses and other forage plants for water and nutrients. Furthermore, in dense mature stands, they can become a physical barrier to livestock due to their thick sharp spines.

In natural environments, Musk Thistle can outcompete native plants for resources thereby reducing biodiversity. They can also become a barrier to native animals that live there.

They can also be a barrier in recreational areas impeding hiking, swimming, and other recreational activities.

Potential Benefits of Invasion

Musk Thistle is of benefit to bees, hoverflies, and butterflies that visit the flowers. The plant is also a food source for numerous caterpillars in the Lepidoptera order.

Methods to Remove Carduus nutans

As always prevention is the preferred method of control. While Musk Thistle is never intentionally planted by people it can be unintentionally introduced in uncertified seeds. This is particularly common in seeds imported from other countries with less strict guidelines on seed purity and invasive species. Do not buy or plant poor-quality seeds.

An important source of infestation is in hay fields and pasture land. One method to help prevent their infestation is to plant more drought-tolerant species so that they are less likely to be outcompeted by Carduus nutans. Furthermore, avoiding over-grazing during periods of drought will help prevent them from successfully spreading.

Physical Control of Nodding Thistle

Once already established, however, physical control is always the most effective means. Physical control is labor-intensive and time-consuming but it usually causes the least amount of environmental damage.

The best time to remove Carduus nutans is when the plants are young or during flowering but before they have gone to seed. If your plants have seeds it is best to manually cut the seed heads off into a garbage bag to prevent them from spreading. Once the seed heads have been cut the plants can be dealt with by other physical means.

Physical methods to remove Nodding Thistle in less dense stands involve digging the plants up. This can be done with a shovel or hoeing the ground to 10 cm below the soil surface to ensure that the tap root is sufficiently removed or damaged. Fortunately, the plant cannot survive once a sufficient amount of its taproot has been removed. This works for both young and mature plants.

Mowing can be used on larger populations of Carduus nutans just prior to going to seed. The mowing will need to be repeated each month during the growing season due to the variation of maturity in the plants. Young plants that have not started blooming will not be affected by mowing so the area will need to be mowed for at least 2 – 3 years.

Disposal of the Plants Once Removed

If you have removed the seeds or have removed plants that are not yet in seed they can be disposed of in a compost heap or left on the soil where removed. Carduus nutans can only reproduce by seed so the rest of the plant is not a disposal concern. However, if you have plants that have seeds on them they must either be burned or solarized. To solarize put the shrubs under a thick black tarp, or into thick black garbage bags and leave them in the full sun for 8 weeks at least to be sure that all seeds are no longer viable.

Chemical Control of Carduus nutans

Chemical applications are almost never an ideal method of control for any invasive species. That is because chemical alteration of the environment often makes the environment more suitable for invasive species than native species. Furthermore, it is often difficult to keep the chemical control method contained so that it does not directly affect any native species that are there during the application process itself. As a result, plots where chemical control is used usually show a decrease in species richness. On the other hand, in plots where only physical control is used species riches significantly increase.

If using chemical control it must be done when the plants are young, actively growing, and before the basal rosettes get too large. This makes adequate timing in spring imperative. Furthermore, repeated applications in New Zealand are starting to show resistance to herbicides.

Chemical control is not recommended.

Biological Control of Musk Thistle

Biological control involves the use of a predator, herbivore, disease, or some other agent to control an invasive species once it is established in the environment. The problem with biological control is that the agent used must be entirely specific to only the target organism before releasing it into the environment. This is often difficult to determine since the agent of control is also not native to the environment and could behave differently when released there. Take the example of the mongoose and the rat. The mongoose was released in Hawaii in the late 1800s to help control the rat. To this day there are still rats in Hawaii but the mongoose has helped to decimate many native bird populations.

Biological control methods are extremely risky and should only be carried out by professionals after years of rigorous study. The use of biological control methods can never be used alone. They must be part of an integrated pest management approach. However, using biological control in conjunction with physical control and ongoing monitoring can be very effective. Following is a list of biological control methods that have been used in North America in an attempt to help control Carduus nutans.

  • Rhinocyllus conicus is a weevil that has been introduced to fight Carduus and several other invasive thistle species including those in the Cirsium genus. The problem in North America is that we have several native Cirsium species so the use of this method is not recommended by some scientists. Furthermore, the success rate has had mixed results so the benefits may not outweigh the risk.
  • Trichosirocalus mortadelo is perhaps a better choice as it is probably restricted to Carduus nutans and it has been used with some success in Canada, the USA, Australia, and New Zealand. There is still some confusion as there may have also been T. horridus introduced and it is uncertain as of yet which one is effective or more effective. More research is ongoing.
  • Domestic goats are not selective in their choice of forage greens and will eat anything including Carduus nutans. Domestic goats can be used in pasture lands to help control the spread of Nodding Thistle. They can also be penned in smaller areas and used in conjunction with physical removal and ongoing monitoring.

Integrated Pest Management & Ongoing Monitoring

Integrated management is always the best approach. In its simplest and least impactful form this involves physical removal methods, possibly biological control methods, replanting with drought-tolerant species, and ongoing monitoring. Integrated management is required because the area needs to be monitored for new or surviving plants otherwise all the hard work done in removal could be wasted if the invasive species is allowed to regrow.

Replanting is Crucial

In all cases of large-scale physical removal, the site should be replanted immediately because the bare soil will allow the seed bank to germinate and reinvade the patch they were removed from. A replanting program should already be planned and ready to implement immediately upon the removal of the Carduus nutans.

Ongoing Monitoring is Essential

In all cases of invasive Musk Thistle removal, ongoing monitoring is absolutely essential. Since they only reproduce by seed an aggressive monitoring program is not necessary. Apart from the repeated monthly mowings suggested in the physical removal of large patches most monitoring programs can be simply done on a yearly basis. Each year check for any surviving or new individuals and remove them so that they are not allowed to set seed.

References and Resources

CABI on Carduus nutans https://www.cabi.org/isc/datasheet/11259

Canadensys Plant Search https://data.canadensys.net/vascan/search

Dictionary of Botanical Terms – Lyrae’s Nature Blog Dictionary of Botanical Terms

Eflora Plants of North America http://www.efloras.org/browse.aspx?flora_id=1

iNaturalist Plant Search https://www.inaturalist.org/home

Plants For A Future on Carduus nutans https://pfaf.org/user/Plant.aspx?LatinName=Carduus+nutans

USDA Plants Database https://plants.sc.egov.usda.gov/home

Willis, Lyrae (2022).  Plant Families of North America. Not yet published.

Currently Seeking Funding To Continue This Non-Profit, Ad-Free Work

If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you!


Gossypium hirsutum Mexican Cotton - Native Species of the Week

Gossypium hirsutum leaf and fruit in the evening sun Sinaloa, Mazatlan, Mexico
Gossypium hirsutum leaf and fruit in the evening sun Sinaloa, Mazatlan, Mexico

Mexican Upland Cotton Gossypium hirsutum – Native Plant of the Week

Introduction

Gossypium hirsutum has large and beautiful pale-yellow flowers and seeds covered in white cotton. It is both a beautiful and useful plant. The flowers are typical Malvaceae-type flowers with large overlapping petals and a staminal column in the center. It is native to Mexico and possibly southern Florida as well as the West Indies, Central America and northern South America. While it grows throughout the southern USA it is not native in most of it. Mexico is considered to be the center of origin and genetic diversity for Gossypium hirsutum where it was domesticated at least as far back as 3500 BC according to archeological evidence. This species of cotton accounts for about 90% of the worldwide cotton production and about 95% of the cotton production in the Americas.

Description of Upland Cotton Gossypium hirsutum

Stem & Leaves

Herbaceous short-lived perennial or annual from a taproot. It grows 1-2 m tall with widely-branching cylindrical stems that are covered with stellate (star-like) hairs.

Leaves are shallowly 3-5 lobed and 4-10 cm wide and they may be hairy or glabrous. They have a cordate (heart-shaped) base and an acute to acuminate (pointed) tip. Leaves grow on a long cylindrical petiole (leaf stalk) that is about 1/2 to the same length as the leaf itself. Leaves also possess stipules, small leaf-like appendages located at the base of the leaves, that are 5-15(-20) mm long.

Flowers & Fruits

Gossypium hirsutum Upland Cotton Mexican Cotton flowers, photo by Lyrae Willis Mazatlan, Sinaloa, Mexico 2020.
Gossypium hirsutum Upland Cotton Mexican Cotton flowers, photo by Lyrae Willis Mazatlan, Sinaloa, Mexico 2020.

The showy cream to yellowish flowers are 2-5 cm across and may or may not possess a red spot in the center of the flowers at the base of the petals. Flowers have small sepals 5-6 mm long with a truncate or 5-toothed tip.

The reproductive organs are contained in a 15 mm long staminal column characteristic of the Malvaceae family. The staminal column contains both the male and female reproductive organs in a single structure. It encloses the style and 3 to 5 stigmas that protrude past the male reproductive organs. The stamens are arranged around the column itself, located below the stigmas.

Its fruit is a 3 to 5 chambered ovoid or sub-globulose capsule that is 2-4 cm long and smooth and hairless. It contains 8-10 mmm seeds covered with white hairs (cotton).

Gossypium hirsutum Upland Cotton Mexican Cotton fruits, photo by Lyrae Willis Mazatlan, Sinaloa, Mexico 2020.
Gossypium hirsutum Upland Cotton Mexican Cotton fruits, photo by Lyrae Willis Mazatlan, Sinaloa, Mexico 2020.

Similar Species Frequently Confused With

Gossypium hirsutum does not have much for unrelated look-alikes due to its showy Malvaceae type flower and its unique fruits covered in white cotton. Occasionally, however, Hibiscus tiliaceus or Maritime Hibiscus is confused with it. Like Upland Cotton, it is also a member of the Malvaceae family and has large showy flowers but it grows as a tree and its flowers are much more yellow and contain a black spot rather than a red spot in the center. It is found in the southern states near the ocean and throughout much of Mexico.

We have many native Gossypium species that it could be confused with, but these all only grow in subtropical and tropical Americas. They can be differentiated as follows:

  • Gossypium darwinii is a narrow endemic confined to the Galapagos with similar flowers but much more deeply lobed leaves. It is easily differentiated by its narrow range.
  • Gossypium barbadense also known as Pima Cotton is found more in western South America but also grows in Florida, Hawaii, Mexico and the West Indies. It grows as a perennial rather than an annual but is very sensitive to cold so will only grow in subtropical to tropical areas. Its flowers are similar but generally a brighter yellow and the center spot may be red or blackish in color. The seeds are black instead of white.
  • Gossypium herbaceum the African Cotton is rare in North America but is found in southern Florida as well as some locations in Mexico. It grows as a perennial to similar heights with similar flowers but its color ranges from white to yellow to pinkish and the spots in the center are purple instead of red. Its leaves are very wide and maybe entire to deeply lobed.
  • Gossypium harknessii is a rare narrow endemic of the Baja peninsula of western Mexico. It is endangered globally. It is a perennial plant with thick glossy green heart-shaped leaves and grows in desert areas. Its flowers are similar but more bright yellow with petals that have wavy edges and very noticeable bright red dots at their bases in the center of the flower.

Distribution of Upland Cotton Gossypium hirsutum

Upland Cotton is native to Mexican, West Indies, Central America and northern South America. It is possibly also native to southern Florida. It has been introduced around the world for cotton production.

In Canada the climate is not suitable for Upland Cotton and it is not currently found there.

In the USA, Upland Cotton is found in California, Texas, Louisiana, Mississippi, Alabama, Florida, Missouri, Illinois, South Carolina, North Carolina, Virginia, Maryland, New Jersey, Pennsylvania and Massachusetts as well as Hawaii.

Gossypium hirsutum is found throughout all of Mexico but is most prevalent in the southern subtropical zone and less common in the northern desert areas.

Habitat & Growing Conditions of Mexican Cotton Gossypium hirsutum

Upland Cotton is found growing in full sun to partial shade in well-drained soils. While it prefers a moderate amount of moisture it will grow in areas prone to drought, in part due to its taproot that penetrates the soil in search of water. It will not grow well in permanently wet areas.

Growing Mexican Cotton in Your Garden

Make sure that the chosen location in your garden is similar to what it grows in its natural environment. Mexican Cotton is an easy plant to grow that tolerates a variety of conditions as long as certain parameters are met. Any area in full sun with well-drained soil will work perfectly. Commercial crops are generally grown in flat areas in well-drained soil and are often rain-fed for their water unless the area is a heavy drought zone in which case they are irrigated occasionally.

The biggest challenge to growing it in your garden is your climate since it is a subtropical and tropical species. However, it can be grown as an annual in more temperate climates.

Propagation is done by seeds that have a high germination and success rate. Some people use clonal propagation but this is generally not practical for the home gardener, especially with the high success rate of seed propagation.

Wildlife Values of Gossypium hirsutum

Native bees and butterflies routinely visit the flowers and the extrafloral nectaries. Birds frequently use the cotton bolls to build their nests. Seeds are rich in oil and are eaten by various birds and wildlife.

Status of Gossypium hirsutum

Mexican Cotton is considered Apparently Secure, G4 according to NatureServe. The IUCN Red List assessed it as globally Vulnerable in 2017.

Gossypium hirsutum does not grow in Canada.

In the USA Gossypium hirsutum is considered Vulnerable S3 in Florida. It is unranked in Massachusetts but should be listed as Exotic as it did not grow there before humans introduced it. In all other states where it is found it is listed as Exotic.

Gossypium hirsutum is found throughout Mexico, however, Mexico does not currently rank the status of native species on a state-by-state basis. The IUCN Red List has it listed as Globally Vulnerable so one could assume that in Mexico where it is native to it is considered a Vulnerable species in the wild.

Traditional or Other Uses of Upland Cotton

Gossypium hirsutum Medicinal Uses

Koasati peoples used a decoction of the roots to aid in childbirth.

Mexican Cotton Other Uses

Zuni people used the fibers to make ceremonial clothing and ceremonial cordage, particularly in masks and crowns associated with rainmaker ceremonies. They also tied the cotton cords around the wrists and ankles of newborns while making offerings so that the rainmakers would supply enough rain for the child to have a rich and full life. Cotton down was also used by the Zuni people in funeral proceedings for priests to symbolize their duties in this world and the other world.

Pima people used the seeds as a food source.

Gossypium hirsutum is a lovely short-lived perennial or annual plant that produces abundant flowers and lovely cottony fruits so it is occasionally grown as an ornamental in suitable climates.

Ethical Wildcrafting of Gossypium hirsutum

In North America, it is vulnerable in Florida which is the only state it is native in, so any wild plants seen in other states are escaped cultivars and can be harvested. It is, however, considered vulnerable globally so wild harvesting is not recommended. Instead, grow it in your garden for both its lovely leaves and flowers as well as its useful properties.

If you do harvest Gossypium hirsutum from the wild as always use the 1 in 20 rule of Ethical Wildcrafting. Pick one in every 20 mature fruits that you see.

Wildcrafting and Processing

Picked fruits can be placed in a basket, bowl, or paper bag and brought home for processing.

Generally speaking you should wait to pick the fruits until they are already dry and fully mature. If you harvest in the morning or shortly after a rain you should dry them before putting into storage. To dry the fruits simply place them on a rack or screen in a single layer and allow them to dry. Once dried the cotton bolls can easily be plucked from the seeds. Once dried they can be stored in a jar or bag for later use.

References and Resources

Canadensys Plant Search https://data.canadensys.net/vascan/search

Dictionary of Botanical Terms – by Lyrae’s Nature Blog https://lyraenatureblog.com/blog/dictionary-of-botanical-terms/

Eflora.org on Gossypium hirsutum http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=200013695

iNaturalist Plant Search https://www.inaturalist.org/home

IUCN Red List https://www.iucnredlist.org/

Native American Ethnobotany http://naeb.brit.org/

Natureserve Explorer https://explorer.natureserve.org/Search

USDA Plants Database https://plants.sc.egov.usda.gov/home

Willis, Lyrae (Not Yet Published).  Plant Families of North America. 

Currently Seeking Funding To Continue This Non-Profit, Ad-Free Work

If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you!


Convolvulus arvensis Field Bindweed - Invasive Species of North America

Convolvulus arvensis the Field Bindweed - Invasive Species in North America. This picture taken by Lyrae Willis in New Mexico, USA
Convolvulus arvensis the Field Bindweed – Invasive Species in North America. Photo by Lyrae Willis in New Mexico, USA

Introduction

Convolvulus arvensis, commonly known as Field Bindweed, is a well-known plant that has been recognized as being invasive in North America for more than 100 years already. This is because of its detrimental effects on agriculture, which led to its early recognition as an invasive weed. Once established, Field Bindweed is notoriously difficult to eradicate. Instead, most removal programs should focus on control rather than eradication. It is already a significant part of North American ecology, so we will never rid ourselves of this weed. However, new patches can be eradicated before they become too well-established. Any growth in natural areas, particularly nature reserves and parkland, should be eradicated where possible in order to preserve the native species also growing there. Agricultural lands, on the other hand, can focus on controlling this invasive pest to mitigate the damage they are causing.

Description of Convolvulus arvensis

Leaves & Stems

Convolvulus arvensis is a herbaceous perennial trailing or climbing vine. It grows from a very deep persistent root system that develops shoots from buds on the roots down to depths of 1 m. It has a main tap root that can grow from 0.5 – 3 m long. Other vertical roots from the main root may penetrate from 5 – 9 m deep, depending on the soil and site conditions. The roots also spread laterally in all directions from the main tap root allowing it to spread adventitiously throughout all of its length. Lateral roots are shallower and generally only penetrate down to 30 cm deep.

The stems are slender and grow from 20 cm to 2 m long and are either hairless or finely pubescent. Stems twine anticlockwise allowing them to climb over structures and other plants.

Leaves are alternate, have a petiole (leaf stalk), and vary significantly in shape and size. They may be lanceolate, ovate, or narrow-oblong and range in size from 1 – 10 cm long and 0.3 – 6 cm wide. The tips are acute, and the leaves themselves may be entire or hastatesagittate (arrow-shaped) at the base and may be hairless or lightly pubescent.

Flowers & Fruits

Flowers of Convolvulus arvensis are axillary and may be solitary or found in cymes of 2-3 flowers on peduncles (flower stalks). They are subtended by bracteoles 2-4 mm long. The flowers have sepals that are free and obtuse in shape and 2.5 – 4.5 mm long. It has a funnel-shaped corolla with 5 radial pubescent bands. The corolla is entire and not divided into individual lobes, 1 – 2.5 cm in diameter and 1 – 2.5 cm long, and may be found in white or pink varieties. It has 5 stamens that are inserted in the corolla tube and a single style with 2 oblong stigmas.

Field Bindweed produces a capsular fruit that is globular or ovoid with a persistent style at the base. It breaks open irregularly and releases four seeds that are 3 – 5 mm in diameter, dark brown or black, with a granular testa (outer coating).

Toxicity

Convolvulus arvensis is considered mildly toxic. It contains tropane alkaloids that can have toxic effects on the autonomic nervous system and have been found to cause intestinal fibrosis in horses. The seeds are especially toxic compared to the rest of the plant.

Similar Species Frequently Confused With

Many, many plants are often confused with Convolvulus arvensis. By far, most of them belong to the Convolvulaceae family and have similar bell-shaped flowers. Occasionally people mistake the unrelated Oenothera speciosa or Evening Primrose for Field Bindweed with its pink flowers. However, this can easily be differentiated by the fact that its corolla actually has 5 overlapping petals and is not at all entire like Convolvulus arvensis. The other non-related plant that is sometimes confused with it is Fallopia convolvulus Black-bindweed or Wild Buckwheat of the Polygonaceae with similar leaves but reddish stems and small lobed flowers. The more closely related and similar-looking species can be differentiated as follows:

  • Calystegia sepium (sometimes as Convolvulus sepium) Hedge Bindweed of the Convolvulaceae family, has a sub cosmopolitan distribution with several subspecies native throughout North America. It has pale matt green sagittate leaves that are arranged spirally rather than alternately on the stem. When in bud, the flowers are surrounded by green bracts that are tinged with crimson, and its flowers, when open, are much larger, from 3 – 7 cm in diameter. Its fruit is an almost spherical capsule.
  • Calystegia subacaulis of the Convolvulaceae family is a narrow-range species endemic to the North and Central California Coast Ranges and the San Francisco Bay Area, where it is found in woodland and chaparral scrub habitat. In its range, it can be differentiated by its hairy stems and leaves and its short stems that grow no more than 20 cm long.
  • Calystegia macrostegia Bellflower of the Convolvulaceae family also has a narrow range found along the Pacific coasts of southern California and western Mexico. Its leaves are large and triangular and are often more than 10 cm wide. It also produces longer stems to 9 m in length and larger flowers 2 – 6 cm in diameter.
  • Calystegia spithamaea of the Convolvulaceae family is native to eastern North America, primarily the northeastern USA. Its stems are mostly erect rather than trailing or twining, its light green leaves are quite hairy, and its flowers are not entire and instead have 5 shallow lobes.
  • Convolvulus equitans the Wounded Bellflower is native to the southern US and northern Mexico. It can be differentiated by its usually lobed rather than entire leaves and its corolla of similar size but with 5 distinct though shallow lobes, each with a very distinctly acute apex. The flower also has a distinct pink or purple throat giving it the common name of Wounded Bellflower.
  • Calystegia silvatica Greater Bindweed is native to Europe but has been introduced to the eastern and western US and Canada. It can easily be differentiated by its larger leaves that are arrow-shaped and its very large flowers up to 9 cm in diameter that are always white and never pink.
  • Calystegia purpurata Pacific False Bindweed is a narrow endemic found along the Pacific coast of California. It has lobed leaves that are triangular in shape and larger flowers up to 5 cm in diameter that vary in color from white to pink to purple or cream and often have purple stripes.
  • Evolvulus sericeus Silver Dwarf Morning Glory of the Convolvulaceae family is native to the southern USA, Mexico, and parts of tropical Americas. It can easily be differentiated by its very small size, only about 30 cm tall, and the fact that it grows erect rather than a vine. It also has linear to narrowly lanceolate light green or gray-green leaves that are generally hairy.
  • Calystegia soldanella the Dunebell is native to beaches of western North America, Europe, and East Asia. It is found growing only on beach sand dunes making it easy to differentiate by its location. Its leaves are also very thick, glossy, and fleshy, producing large singular pink flowers.
  • Calystegia occidentalis Chaparral False Bindweed is native to California and Oregon, where it grows in foothills and montane habitats. It has small leaves up to 4 cm that are typically hairy, lobed, and arrow-shaped. It produces 1-4 flowers on a single stalk, and each flower is fairly large, from 2 – 5 cm in diameter, and is white, cream, or yellow in color but never pink.
  • Bouchetia erecta of the closely related Solanaceae family is native to the gulf coast of the US. It is a much smaller plant, never more than 30 cm tall, and it has lanceolate leaves and lobed corollas containing bright yellow stamens.
  • Ipomoea imperati  Beach Snowdrop of the Convolvulaceae family is native to the Americas but is restricted to beach sand dunes. Its white flowers are similar in shape but are usually larger and may be tinged with blue. Its leaves are highly variable in shape but are glossy and fleshy.
  • Ipomoea lacunosa  Snowdrop of the Convolvulaceae family is native to eastern North America, mostly in the USA. It has a much smaller taproot, but the vines can grow to similar lengths, 2 m long. Its leaves are always large (8 – 9 c long) and are ovate when young but become cordate (heart-shaped) when mature, and they are always on long petioles (leaf stalks) at least 3 cm long. The similar-sized, usually white but occasionally pink flowers may or may not be shallowly lobed. It has distinctive pinkish or purple anthers on white filaments, and it produces large seed capsules that are spherical and hairy.

Native Distribution of Convolvulus arvensis

Field Bindweed is native throughout much of Europe and Asia in temperate, tropical, and Mediterranean regions.

Habitat Types Where Field Bindweed is Found

Convolvulus arvensis grows successfully in a wide range of temperate, tropical, and Mediterranean climates. It is very problematic for agriculture in the temperate zones between 60°N to 45°S latitude, though it also grows in tropical regions as well. It is a serious pest in wheat, barley, corn, legume, and sugar beet fields as well as vineyards and tree crops.

While it is known mostly as an agricultural weed, it also is frequently found in natural areas, particularly in riparian habitats and open communities such as meadows and grassland. They thrive in areas with cleared and disturbed ground, such as agricultural areas and fallow fields, as well as new construction, land clearing, roadsides, and waste areas.

Human Uses of Field Bindweed

Convolvulus arvensis has a number of medicinal properties. A decoction of the root may act as a diuretic and a laxative. Tea from the flowers and or the leaves has been used to treat fevers and wounds. Caution should be used, however, as taking too much or for too long could potentially cause blood in the urine and toxic effects on the autonomic nervous system. In small doses over short periods of time, it is considered safe.

The stems are sometimes used as twine for tying up plants and other things. While it works well as twine, its usefulness is short-lived, so it should not be used for long-term storage of things, for example. A green dye is sometimes made from the whole plant.

Distribution of Convolvulus arvensis in North America

The species was first brought to the US likely as a contaminant in seed though it was also intentionally planted in baskets. It was first reported in the wild in Virginia in 1739 and, by the 1800s, had already spread throughout the eastern seaboard. By 1838 it had made its way west to California.

In Canada, Convolvulus arvensis has been recorded throughout the southern provinces of British Columbia, Alberta, Saskatchewan, Manitoba, Ontario, Quebec, New Brunswick, Nova Scotia, and Prince Edward Island. So far, it has not been reported in Newfoundland or the northern territories.

In the USA, Field Bindweed is even more widespread. It has been reported in every state in the continental USA as well as in Hawaii. It is absent only in Alaska.

In Mexico, Convolvulus arvensis so far has been reported in Baja California Norte, Baja California Sur, Sonora, Sinaloa, Chihuahua, Coahuila, Nuevo Leon, Durango, Tamaulipas, San Luis Potosi, Jalisco, Zacatecas, Aguascalientes, Guanajuato, Queretaro, Hidalgo, Michoacan, Mexico State, Mexico City, and Puebla. Given that the climate is suitable for it in many other states on the Caribbean side and in the south, it is likely that its range is still spreading.

Eurasian Bindweed has been introduced on every continent except Antarctica.

How Field Bindweed Spreads

Convolvulus arvensis was first introduced to the Americas by accidental introduction as a contaminant in other seeds. It may have also been intentionally planted in flower baskets. Since it spread as a contaminant early in the days of European settlement, it has already invaded much of the habitable zones in North America suitable to its growth.

Short-distance dispersal occurs through contamination of its seeds in other plant materials, where it grows as a weed in fields with agricultural crops. Short-distance dispersal of seeds can also occur via water, clothing, animals, vehicles, and machinery.

Birds can also disperse the seeds short and long distances because the seeds can remain viable in the digestive tract for up to 144 hours making migratory birds an important source of long-distance dispersal.

Dispersal also occurs with poorly disposed of yard waste debris filled with seeds or simply cut plants and roots that can regenerate via fragments. Solarization of removed plant matter is critical to prevent further spread this way.

Habitats at Risk of Invasion in North America

Most habitats in North America at risk of invasion have already been invaded. These include any disturbed areas such as agricultural fields, pastures, roadsides, and waste areas. Its range in North America will likely continue to spread in parts of Mexico where it has not yet reached its potential or has not yet been accurately reported as already being there. As climate change continues its warming trend in northern latitudes, it seems likely that it will continue to spread northwards in the southern provinces of Canada and may, in some locations, particularly near the coast, even reach the southern parts of the Northern Territories.

Impacts of Invasion

Field Bindweed threatens agricultural crops where it has been found to compete for water and nutrients, resulting in reductions in yields anywhere in the range of 0 – 100%. While only studied in crops, it is logical to assume a similar threat to native plant communities. In addition to competition for water and nutrients, it is an aggressive grower and climbs over other vegetation, physically smothering it and reducing biodiversity in that way as well.

It has been reported to be mildly toxic to some grazing animals, horses in particular. However, sheep, pigs, and goats seem more resistant to the toxic effects.

Potential Benefits of Invasion

It does not supply any significant nutritional food sources for native or domestic species and threatens species richness. There are no potential benefits of its invasion.

Methods to Remove Field Bindweed

As always, prevention of invasive species introductions is the preferred method. However, in the case of Convolvulus arvensis it has already been well established throughout North America, so control becomes the next best option. Control, however, is made particularly challenging due to both the longevity of the seeds in the soil seed bank (20 or more years) and the ability of the plant to regenerate via fragmentation of the stems or roots.

Fortunately, no one appears to be selling Convolvulus arvensis, so deliberate introductions of new plants are not likely to happen. Sadly this is not the case with many invasive species. Perhaps because Field Bindweed is such a well-known pest plant and has been known as such for well over 100 years now, people have actually stopped intentionally planting it.

Physical Control of Convolvulus arvensis

Once already established, physical control is always the most effective means. Physical control is labor-intensive and time-consuming, but it usually causes the least amount of environmental damage. Eradication of Field Bindweed once well established is difficult to impossible, but control and mitigation can be achieved.

Physical methods to remove Field Bindweed generally involve removing mature individuals before they go to seed to reduce the amount of seed in the soil. Since seeds are resilient enough to last more than 20 years in the soil, this makes control particularly challenging. Each year ongoing monitoring will be needed to remove any new seedlings as they generate from the soil seed bank.

The other added factors that make the control of this plant particularly challenging are its ability to resprout after being cut down, as well as its ability to generate new plants via fragmentation. When mature individuals are removed, try to pull out as much of their extensive root system as possible. This includes their very long and deep taproot as well as the lateral roots it produces that generally remain in the top 30 cm of the soil. Use a weed puller or a shovel to dig the taproot. For lateral roots, try to trace them from the main taproot along the soil. Heavy or compacted soils make this challenging, but the more root you remove, the better.

Since it can resprout from any remaining fragments, this absolutely must be followed with ongoing monitoring several times throughout the growing season. This can be done via the cutting of new sprouts or the cultivation method. To use the cutting method, simply cut any shoots that are regenerating, being sure to cut about 8 cm below the soil surface and repeat this for several years. Eventually, fewer and fewer sprouts will regenerate until the food reserves in the root system are completely depleted, and they can no longer regenerate. The more frequent the cutting, the less the root system has a chance to restore any energy stores, and the less treatment time is required. At a minimum, 2-3 years of extensive treatments will be required.

Frequent cultivation is a popular method to deal with resprouting mature plants and new seedlings. If the area is cultivated with a hoe or using agricultural equipment every 14 days throughout the growing season, the Field Bindweed can sometimes be controlled in as little as 2 years. It is critical to return to the patch and cultivate every 14 days, however. If left too long, the new green leaves will supply more energy to the extensive root system and prolong the necessary treatment time.

Solarization of large patches is a less labor-intensive method, and it will both deplete the root reserves and destroy any new seedlings as they try to emerge. Cover the area with a large black or other dark tarp and weigh the tarp down, leaving it there for 3 – 5 years. With many invasive plants, 2 years is sufficient, but due to the extensive root systems of Convolvulus arvensis, a minimum of 3 years is absolutely necessary. Prematurely removing the plastic mulch will no doubt result in re-infestation. The area should also still be monitored around the tarp edges throughout the growing season to look for lateral roots that try to sprout new plants outside of the tarped area.

Disposal of Convolvulus arvensis Once Removed

All plant material, whether it has seeds or not, must either be burned or solarized. Even without seeds, Convolvulus arvensis can generate new plants via root and stem fragmentation, so all plant material must be destroyed before disposal to prevent new infestations from carelessly discarded plant waste. Burning is a quick and effective method to dispose of the waste. However, it is not allowed in all areas and certainly not in all seasons. The other option is to solarize the material. To solarize, put the shrubs under a thick black tarp or into thick black garbage bags and leave them in the full sun for a good 8 weeks at least to be sure that all seeds and fragments are no longer viable. Some sources recommend shorter solarization periods, but this varies with latitude, aspect, cloud cover, etc, so leaving as long as possible ensures complete sterilization of the plant debris. Once properly solarized, the material can then be discarded at your local dump but do still inform them of what the material is so that they can properly dispose of it.

Chemical Control of Field Bindweed

Chemical applications are almost never an ideal method of control for any invasive species. That is because chemical alteration of the environment often makes the environment more suitable for invasive species than native species. Furthermore, it is often difficult to keep the chemical control method contained so that it does not directly affect any native species that are there during the application process itself. As a result, plots where chemical control is used usually show a decrease in species richness. On the other hand, in plots where only physical control is used, species riches significantly increases.

Furthermore, there are no chemical control methods that effectively target only Field Bindweed. Also, Field Bindweed has been shown to become herbicide-resistant in some cases after repeated application. Due to the physical nature of Field Bindweed, multiple applications are always needed.

Chemical control is not recommended.

Biological Control of Convolvulus arvensis

Biological control involves the use of a predator, herbivore, disease, or some other agent to control an invasive species once it is established in the environment. The problem with biological control is that the agent used must be entirely specific to only the target organism before releasing it into the environment. This is often difficult to determine since the agent of control is also not native to the environment and could behave differently when released there. Take the example of the mongoose and the rat. The mongoose was released in Hawaii in the late 1800s to help control the rat. To this day, there are still rats in Hawaii, but the mongoose has helped to decimate many native bird populations.

Biological control methods are extremely risky and should only be carried out by professionals after years of rigorous study. The use of biological control methods can never be used alone. They must be part of an integrated pest management approach.

Since we have so many native Convolvulus species in North America, and some of them narrow endemic species such as those in California in particular, biological control is not an option. Multiple biological control agents have been tested for Convolvulus arvensis, but so far, all of them have also impacted our native species.

Grazing can help control Field Bindweed, but cattle will not eat it, and horses should not eat it. Sheep and goats will eat it but prefer other plants, including native species. Pigs, however, seem to love the entire plant and will eat both the above-ground plant matter and the roots if given the time. If you have an area that needs to be controlled, build a fence around it. Then allow the pigs to graze unchecked for 2 years, and they should destroy all of the Convolvulus arvensis growing there. Using goats or sheep in a similar manner can also be effective. However, since sheep and goats do not eat the roots, they would need to remain fenced in the patch for 3-4 years instead of 2.

Integrated Pest Management & Ongoing Monitoring

Integrated management is always the best approach. In its simplest, most effective, and least impactful form, this involves physical removal methods, possibly biological control methods, replanting, and ongoing monitoring. Integrated management is required because the area needs to be monitored for returning sprouts or seedlings; otherwise, all the hard work done in removal could be wasted if the invasive species is allowed to regrow.

Replanting With Native Species is Crucial

In all cases of large patch removal, the site will need to be replanted immediately because the bare soil will allow the seed bank of Field Bindweed and other invasives in the soil to germinate and reinvade the patch they were removed from. A replanting program should already be planned and ready to implement immediately upon successful removal of Convolvulus arvensis. If you are just removing an isolated individual replanting with native species is generally not necessary as the native species around it will simply fill in the spot left behind.

Ongoing Monitoring is Essential

In all cases of invasive Field Bindweed removal, ongoing monitoring is absolutely essential. Regular monitoring programs should be put in place to ensure that any surviving individuals are removed so that the population is not able to recover. In the first 3 years, monitoring should be done every 2 weeks to ensure that resprouting plants are destroyed to deplete the energy reserves of their extensive root systems. After the first 3 years, monitoring can be cut down to 2-3 times per growing season to remove any seedlings that germinate or any rootstock still attempting to recover.

Monitoring is required whether the area is replanted or not. Do not assume that once replanted, Convolvulus arvensis will not still try to grow. Field Bindweed is aggressive and prolific and will outcompete planted vegetation if yearly monitoring is not put in place to remove young native plants before they have a chance to become established. Once the area is well established with mature native plants, a simple once-a-year monitoring program is sufficient to root out any new individuals that may try to pop up. And you get to view the successful rewards of your hard work!

References and Resources

CABI on Convolvulus arvensis https://www.cabi.org/isc/datasheet/15101

Canadensys Plant Search https://data.canadensys.net/vascan/search

Dictionary of Botanical Terms – Lyrae’s Nature Blog Dictionary of Botanical Terms

Fire Effects Information System on Field Bindweed https://www.fs.fed.us/database/feis/plants/vine/conarv/all.html

iNaturalist Plant Search https://www.inaturalist.org/home

USDA Plants Database https://plants.sc.egov.usda.gov/home

Willis, Lyrae (Unpublished).  Plant Families of North America.

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If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you!



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Arisaema triphyllum flower in Virginia, USA
Arisaema triphyllum flower in Virginia, USA

Jack-in-the-Pulpit Arisaema triphyllum – Native Plant of the Week

Introduction

Arisaema triphyllum is easily one of my favorite eastern forest plants. Of course, I have many many favorites, but this one is so much a favorite of mine it is going to be one of my next tattoos! It has beautiful shiny foliage and unique flowers contained in an often colorful spathe. Arisaema triphyllum is a monocot of the Araceae family, an interesting family whose flowers are all tiny little flowers borne on a dense spadix and often enclosed in a colorful bract (modified leaf) known as a spathe. It is hard to misidentify it for anything else when it is in flower in mid-spring on the forest floor.

Description of Jack-in-the-Pulpit Arisaema triphyllum

Stem & Leaves

Herbaceous perennial 30-65 cm tall arising from a corm (underground storage organ).

Arisaema triphyllum has one or two large glossy leaves on long petioles (leaf stalks) 30-65 cm high. Each leaf is 8-15 cm long and 3-7 cm wide and is divided into three entire (not toothed) leaflets whose margins may be smooth or slightly undulate (wavy). The presence of 3 leaflets makes it commonly mistaken for poison ivy before it flowers.

Arisaema triphyllum with purple spathe - Virginia USA
Arisaema triphyllum with purple spathe – Virginia USA

Flowers & Fruits

The unique flower of Arisaema triphyllum blooms on a separate stalk from the leaves from March to June, depending on the location, elevation, aspect, etc. It is produced on a spadix (spike) of many tiny flowers enclosed in a spathe (a large bract that encloses the spadix) that usually folds over the spadix like a hood. At first, the flowers are all male and located on the upper part of the spadix, but as they mature, they often become hermaphroditic, with female flowers appearing on the lower part of the spadix. The spadix and spathe may be green or purple or green and purple striped.

Specimens that remain male-only die back to their corm in summer, but specimens that become hermaphrodites go on to produce clusters of bright red, somewhat oval-shaped berries on their spadix.

Toxicity

All parts of the plant are considered poisonous as they contain calcium oxalate and will seriously irritate the mouth and digestive tract if eaten. Handling the corms with bare hands can irritate the skin.

Similar Species Frequently Confused With

It is difficult to misidentify the Arisaema genus when in flower. However, before it flowers, it is often confused with a number of unrelated species.

  • Panax quinquefolius American Ginseng grows to similar heights and has a similar range. However, it generally has 3 long-stalked leaves from a rhizome, and each leaf is toothed and divided into 5 leaflets rather than smooth and divided into 3 leaflets. It also produces clusters of red berries aiding in the mistaken identification, but when in flower, this species has umbels of numerous small white flowers rather than a spathe and spadix.
  • Toxicodendron radicans Poison Ivy occupies the entire range of Arisaema triphyllum, and then some, and it too has compound leaves with 3 leaflets of similar shape and size. However, it grows more like a vine or a straggly shrub leaning on other plants for support, growing to over 1 m tall. Furthermore, its stems are often a pink or reddish color, and they have multiple racemes of tiny white flowers growing from the leaf axils.
  • Symplocarpus foetidus Eastern Skunk Cabbage also shares a similar range and is sometimes mistaken for Arisaema simply because of the spathe and spadix. However, skunk cabbage spathe and spadix are much larger, very odorous, and surrounded by very large simple (not compound) leaves arising directly from the ground rather than on stalks.
  • Podophyllum peltatum Mayapple has lobed leaves that are much larger and not divided into individual leaflets. It also has a single large white flower that appears below its leaves, and it produces a single large round fruit.

Arisaema triphyllum is recognized as a highly variable species, and many disagree with the number of variants and subspecies . This is further confounded by a high degree of hybridization with Arisaema dracontium. The subspecies or variants typically vary however in small degrees on the size of the spadix and spathe and the degree of fluting of the spathe. Identification to the species level alone is sufficient in most cases. There are two other Arisaema species in their range, however, that vary more significantly, and they can be differentiated as follows:

  • Arisaema dracontium shares the same range as Arisaema triphyllum but can easily be differentiated by its leaves with 7 or 8 leaflets all arising on the same side of a curved petiole and by its much smaller spadix enclosed in a much thinner spathe that tapers to a very long extended and thin point that goes well beyond the spadix.
  • Arisaema quinatum has a flower that looks much more similar to Arisaema triphyllum, but it has leaves divided into 5 instead of 3 leaflets. It also has a narrow range, found only in the southeastern part of the USA.
Arisaema quinatum with 5 foliate leaves - Virginia USA; Some sources call it Arisaema triphyllum ssp quinatum.
Arisaema quinatum with 5 foliate leaves – Virginia USA; Some sources call it Arisaema triphyllum ssp quinatum.

Distribution of Jack-in-the-Pulpit Arisaema triphyllum

Jack-in-the-Pulpit is a northeastern North American species.

In Canada, Arisaema triphyllum is found in Manitoba, Ontario, Quebec, New Brunswick, Nova Scotia and Prince Edward Island.

In the USA, Arisaema triphyllum is found throughout the entire eastern half of the USA, from North Dakota south to Texas, and all states east of that. It is only found in the far eastern sections of North and South Dakota, Nebraska, Kansas, Oklahoma, and Texas and is absent from the plains regions of those states.

Arisaema triphyllum is not found in Mexico. It is only as far south as eastern Texas.

Habitat & Growing Conditions of Jack-in-the-Pulpit Arisaema triphyllum

Jack-in-the-Pulpits are found in moist to wet woodlands, forest edges, and meadows with partial shade. While they will grow in a variety of conditions, they thrive best in moist shady forests with a seasonal wet period.

They require a high amount of water and prefer moist, humus-rich soil with a significant leaf litter layer. While they will grow in sunny conditions, they perform best in part to full shade.

Growing Jack-in-the-Pulpit in Your Garden

Arisaema triphyllum is an easy-to-grow native perennial that requires little to no maintenance, providing the conditions are suitable. Make sure that the chosen location in your garden is similar to what it grows in its natural environment. If you have shaded to partially shaded woods with leaf litter, that would be ideal. Otherwise, growing in shade or partial shade will work as well but be sure to amend the soil with lots of organic matter and top dress with leaf litter.

Propagate by root division by separating the cormlets (baby corms) from the parent corm in the fall. Alternatively, you can grow it from seeds collected from mature fruits in late August or September. Seeds may take up to 2 years to germinate. To grow from seed, remove them from the pulpy berries collected in late summer and stratify them in moist moss placed in the fridge for a minimum of 60 days before planting in the late fall, placing them approximately 1.75 cm deep in the soil. Alternatively, they can be sown early the following spring.

Wildlife Values of Jack-in-the-Pulpit

Native birds and mammals will eat the berries. They are not recommended for human consumption, however, due to the presence of calcium oxalate crystals that will seriously irritate the mouth and digestive tract if ingested.

Native gnats and flies routinely visit the flowers and are the primary pollinators.

Status of Arisaema triphyllum

Arisaema triphyllum is considered Globally Secure, G5.

In Canada, Jack-in-the-Pulpit is considered Locally Secure S5 in Ontario and New Brunswick, Apparently Secure S4 in Nova Scotia and Prince Edward Island, and Critically Imperilled S1 in Manitoba. It is unranked in Quebec.

In the USA, Arisaema triphyllum is considered Locally Secure S5 in Illinois, Indiana, Kentucky, West Virginia, Virginia, North Carolina, and Georgia. It is considered Apparently Secure S4 in Iowa. In all other states where it is found, its status is not yet determined.

Arisaema triphyllum is not found in Mexico.

Traditional or Other Uses of Jack-in-the-Pulpit

Arisaema triphyllum Medicinal Uses

The Cherokee used a poultice of the root for headaches, boils, ringworm, and diseases of the skin. They also used it for colds and coughs and as a carminative, expectorant, liniment, and diaphoretic. The Chippewa used a decoction of the roots for an eyewash, and the Iroquois steamed the roots for sore eyes. Iroquois also used it internally for headaches, pains, cramps, diarrhea, colds, tuberculosis, and as a febrifuge and a blood medicine to induce temporary sterility in women. Iroquois used it externally in a liniment for sore joints, bruises, and lameness and in a snuff for catarrh. Choctaw also used it as a blood medicine.

The Malecite and Micmac used it in a poultice for abscesses, pains, and boils. The Menominee and Ojibwa used a poultice for sore eyes and the Menominee in a lip incision to ward against witchcraft. The Meskwaki used a powdered form of the root as a poison, sedative, and snakebite treatment and in ceremonies to predict recovery or death. Mohegan is also used it for pain in small doses, poison in larger doses, and diluted as a gargle for sore throats. The Pawnee used the powdered root as an analgesic and anti-rheumatic and used the seeds in gourd shells as a rattle.

Jack-in-the-Pulpit as an Ornamental

With its unique flowers, Arisaema triphyllum is often a popular garden ornamental and is planted in temperate gardens throughout North America.

Ethical Wildcrafting of Arisaema triphyllum

Check the status in your state before harvesting since it is imperiled in some areas. See the above section on Status. Alternatively, grow it in your garden for unique flowers.

If you are harvesting Arisaema triphyllum from the wild, as always, use the 1 in 20 rule of Ethical Wildcrafting. Pick one in every 20 flowers, leaves, or plants that you see. If you are harvesting cormlets to propagate simply pick one out of 20 plants, dig up the corm, and harvest the baby corms from the parent corm. Then be sure to replant the parent corm so that it can grow the following year again.

Wildcrafting and Processing

Picked roots can be placed in a basket, bowl, or paper bag and brought home for processing. If you are harvesting multiple products on the same day, be sure to label the roots in a paper bag so that you do not confuse different plants. Wear gloves when harvesting due to the poisonous side effects of the plant, especially its roots.

To dry the corms or cormlets, brush off the dirt and place them on a drying rack to dry. If you harvested them to grow in your garden, do not dry them. Instead, simply plant them in your garden right away, about 3 cm deep.

Once dried, corms can be stored in a jar for later use. Label your jar with the species’ name and the date of harvest. I also usually add the location of the harvest for my own reference. Do not grind or crush the roots until you are ready to use them to keep them as fresh as possible and preserve their medicinal properties. When you pre-grind, even if stored in glass jars, this increases the oxidation rate and rapidly degrades the medicinal properties so that they are rendered ineffective in a shorter amount of time than if left as whole as possible.

References and Resources

Canadensys Plant Search https://data.canadensys.net/vascan/search

Dictionary of Botanical Terms – by Lyrae’s Nature Blog https://lyraenatureblog.com/blog/dictionary-of-botanical-terms/

eflora.org on Arisaema triphyllum http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=222000013

iNaturalist Plant Search https://www.inaturalist.org/home

IUCN Red List https://www.iucnredlist.org/

Lady Bird Johnson Wildflower Center on Arisaema triphyllum https://www.wildflower.org/plants/result.php?id_plant=artr

Native American Ethnobotany http://naeb.brit.org/

NatureServe Explorer https://explorer.natureserve.org/Search

USDA Plants Database https://plants.sc.egov.usda.gov/home

Willis, Lyrae (Unpublished).  Plant Families of North America. 

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Wisteria sinensis & W. floribunda Chinese & Japanese Wisteria - Invasive Species of North America

Arisaema quinatum with 5 foliate leaves - Virginia USA; Some sources call it Arisaema triphyllum ssp quinatum.
Wisteria sinensis or Chinese Wisteria an invasive species in North America, these ones were in Marietta, Georgia.

Introduction

Wisteria sinensis Chinese Wisteria and Wisteria floribunda Japanese Wisteria are both popular garden ornamentals that have become invasive throughout North America. In Canada, it only grows in the warmest regions near the coast and so far has not become invasive there. However, it is becoming very invasive throughout the eastern USA, where the climate is similar to its native range in China. There you can see it growing over the top of trees and old buildings and smothering native vegetation. Even in areas where it has not yet become invasive, it is well known to be aggressive and difficult to keep in check, requiring routine maintenance in your garden to keep it from growing over your other plants. There are so many lovely native vines in North America that are not aggressive and invasive, do yourself a favor and research what is native to your area and grow those instead. For instance, if you live in the eastern USA, why not grow American Wisteria or Trumpet Creeper that are native to this area?

Many of the populations found to be invasive in the eastern US have recently been discovered to be hybrids of Wisteria sinensis and Wisteria floribunda. While this article focuses on Chinese Wisteria Wisteria sinensis, keep in mind that many invasive plants are hybrids. Furthermore, both of these species appear very similar and, once hybridized, are difficult to distinguish. Finally, both are not native to North America, so for the purposes of control and eradication, identification to simply Asian Wisteria (as opposed to the less aggressive and native American Wisteria Wisteria frutescens) is sufficient in most cases.

Description of Wisteria sinensis

Leaves & Stems

Chinese Wisteria is a climbing, twining, or trailing perennial vine of the Fabaceae (Legume) family. While it also occasionally can be seen growing more as a shrub, it typically is a vine, and its stems can reach 20 m up into the canopy and reach 38 cm in diameter. They twine clockwise, and they occasionally branch alternately along the stem.

The leaves of Wisteria sinensis are compound and about 0.3 m in length, and these, too, alternate along the stem. It has from 7 to 13 leaflets that are themselves attached opposite along the stock of the compound leaf. Leaflets are oblong in shape, have wavy edges and long tapering tops, and are from 2-6 cm long. When young, the leaflets are covered in silky hairs, but they become mostly hairless as they mature.

Flowers & Fruits

The flowers or Wisteria sinensis are borne on showing dangling racemes 10-50 cm long and 7-10 cm wide. Its flowers have the typical pea family morphology, are very fragrant, and are usually vibrant blue to lavender or violet in color though occasional white varieties are seen. The flowers all mature and open around the same time, unlike Japanese Wisteria (see Similar Species below).

Fruits of Chinese Wisteria are typical legume-type pods that are velvety brown and 10-15 cm long. The pods are narrowed toward the base and have constrictions between the seeds in the pods. The pods each contain from 1 to 8 round but flattened brown seeds roughly 1.2 – 2.5 cm in diameter. 

Toxicity

Many sources consider the flowers, leaves, fruits, and seeds of the Chinese Wisteria to be poisonous. This is not uncommon in the Fabaceae family. Accidental or purposeful ingestion may cause symptoms such as nausea, vomiting, and diarrhea. 

Similar Species Frequently Confused With

Occasionally people mistake Chinese Wisteria for non-related species such as:

  • Paulownia tomentosa with its purple flowers, but its flowers are large and tubular and not legume-like, and it has massive simple leaves rather than compound leaves.
  • Dermatophyllum secundiflorum is native to the southern US and northern Mexico and has similar legume-like racemes of purple blossoms. But it can easily be differentiated by its evergreen tree habit and its 6-15 cm long compound leaves of oval leaflets 2-6 cm long with rounded or indented tips rather than tapering.
  • Apios americana is a vine of the Fabaceae family, also native to the eastern US, with compound leaves with leaflets that have tapering tips. However, its flowers are not at all similar. Instead, they are red-brown to somewhat purple and occur in dense clusters rather than elongated racemes. This plant is also not poisonous and was an important food source for native peoples before the arrival of the Europeans.
  • Syringa vulgaris Common Lilac is also occasionally mistaken for Chinese Wisteria, but these are usually shrubs and can easily be differentiated by the distinctive fragrance of its blossoms and its 4 petal flowers that look nothing like a legume flower.

There are two species in the same genus that are more often confused with Chinese Wisteria.

  • Wisteria floribunda Japanese Wisteria looks very similar to Chinese Wisteria, and both are closely related. They are also both found throughout the eastern US though Chinese Wisteria does have a broader distribution and is also occasionally found in the West. Chinese and Japanese Wisteria have often been found hybridizing throughout this range at times, making the actual distribution of each problematic to determine. However, given that both are non-native, when found, identification as an Asian and not American Wisteria is sufficient for management purposes. The two Asian Wisterias can be differentiated by their leaves and their twining habit. Japanese Wisteria has 13 to 19 leaflets on its compound leaves, while Chinese wisteria has 7 to 13 leaflets. Japanese Wisteria twines counter-clockwise while Chinese Wisteria twines clockwise. Furthermore, the flowers of Japanese Wisteria bloom sequentially from the bottom up, while Chinese Wisteria blooms all at the same time.
  • Wisteria frutescens American Wisteria is native to the eastern USA, sharing much of the same range as Chinese Wisteria. It can usually be easily differentiated by its much smaller racemes, only 5-15 cm in length, which is why it is less popular as a garden ornamental despite its native status and non-invasive habit. It also does not grow as tall, it has a shorter bloom time, its blossoms are not fragrant, and the seed pods are smooth instead of velvety when they mature.

Native Distribution of Wisteria sinensis

Wisteria sinensis is native to China in Guangxi, Guizhou, Hebei, Henan, Hubei, Shaanxi, and Yunnan provinces. It has been widely introduced around the world outside of its native range.

Habitat Types Where Chinese Wisteria is Found

Chinese Wisteria tends to escape cultivation vegetatively when left unchecked. From there, they spread into forest edges, disturbed areas, roadsides, ditches, and riparian habitats. They tend to become invasive in warm temperate climates but remain more controllable in cool temperate climates, where they will suffer dieback from harsh winter temperatures. They will not survive in the colder climates in most of Canada, for example.

Wisteria sinensis prefer deep rich soils but have been found growing in a range of sites. They will not tolerate extended periods of drought but can tolerate some seasonal flooding. While they grow best in full sun, they also tolerate full shade and are capable of spreading in these conditions.

Human Uses of Chinese Wisteria

Chinese Wisteria is widely used as a popular garden ornamental for its abundant fragrant blossoms. They are used on porches, gazebos, fences, walls, and in gardens and parks when also pruned as a shrub.

Since it is poisonous, it is not used as a food source or medicinally.

Distribution of Wisteria sinensis or their hybrids in North America

The species was first brought to the Americas in 1816 as a garden ornamental, where it quickly became popular.

In Canada, Wisteria sinensis has not been recorded outside of cultivation. It is cultivated in the southern coastal regions, however, and may one day escape cultivation there, particularly with climate change and the warming of southern Canada.

In the USA, Chinese Wisteria is found outside of cultivation in Texas, Missouri, Arkansas, Illinois, Kentucky, Tennessee, Mississippi, Alabama, Georgia, Florida, North & South Carolina, Virginia, West Virginia, Delaware, Washington DC, Michigan, Pennsylvania, New Jersey, New York, Massachusetts, Connecticut, Vermont, and Hawaii. It has also been reported in the Pacific coastal states, but it is not known if it has truly escaped cultivation there yet or not.

In Mexico, Wisteria sinensis so far has only been reported in Baja California Norte.

Chinese Knotweed is cultivated in temperate climates on every continent except Antarctica and has been reported as naturalized in New Zealand, Argentina, Chile, Brazil, South Africa, and many European countries.

How Chinese & Japanese Wisteria Spreads

It is primarily spread over long-distance by deliberate human introductions as garden ornamentals that then escape from cultivation when left unchecked in a suitable climate.

Short-distance dispersal occurs primarily from unchecked garden specimens that escape cultivation through vegetative spread. Rooting occurs at nodes in the vines and easily produces new plants this way. Another source of short-distance dispersal is carelessly discarded yard waste dumped in waste sites or forest edges that regenerate and produce new populations. Seeds are produced when the conditions are right and short-distance dispersal can occur this way as well. Since nothing eats the seeds, this is limited to locations right next to the parent plant.

Habitats at Risk of Invasion in North America

All warm temperate areas with adequate rainfall in North America will be at risk of invasion. Cool temperate areas are also at risk but less so as the vine is less aggressive in those regions. It invades forests and riparian areas, in particular throughout these regions but is also a threat to forest edges and shrub meadows, providing there is adequate moisture. Given that these similar conditions exist on the west coast, but Wisteria sinensis so far has not been reported as invasive there, this is the region most at current risk of new invasion in North America. People in those areas should not plant Chinese Wisteria in their gardens and should control or remove any existing plants there before they become a problem.

Desert areas (except valley bottoms with permanent rivers) and montane regions about 1000 m in elevation are not at risk of invasion.

Impacts of Invasion

Wisteria sinensis invades forest edges and penetrates the forest reducing all the biodiversity within it. Chinese Wisteria climbs by twining around trees and killing them by girdling or simply smothering them so they are unable to photosynthesize. They are currently being managed in many state and national parks in the eastern USA because of their detrimental effects on native tree populations and biodiversity. Chinese Wisteria also smothers the native understory vegetation by growing over it and reducing the availability of light. In areas where it gains a foothold in the ecosystem, biodiversity is always significantly reduced.

Potential Benefits of Invasion

Other than the fact that bees still visit their flowers, the rest of the plant is poisonous and provides no wildlife value.

Methods to Remove Chinese Wisteria

As always, prevention is the preferred method of control. It, like most invasive species, is still widely sold online and in most local garden stores. Do not buy or transport any Chinese Wisteria. Do not plant it in your yard. If it is already growing there, and especially if you live on the west coast, where it has not yet gained a foothold but certainly will if left unchecked, then please destroy it and replace it with a native vine.

If you see them being sold online or in your local garden stores, please inform them of their invasive status and ask them to do their part and cease selling them. Ask them to instead sell more native species as ecologically friendly garden alternatives to invasive species.

Physical Control of Chinese and Japanese Wisteria

Once already established, however, physical control is always the most effective means. Physical control is labor-intensive and time-consuming, but it usually causes the least amount of environmental damage.

Physical methods to remove populations of Chinese Wisteria generally involve the physical pulling of all above-ground vines from the soil and the trees. Then this is followed by repeated cutting of the vine down to its rootstock. Since they can resprout many times from their rootstock, this should be repeated every 2-3 weeks from spring until early fall. This will exhaust the rootstock, and it will no longer be able to resprout. It will, of course, need to be monitored the following spring for any surviving rootstocks.

Juvenile plants or isolated individuals can be controlled by digging out the entire rootstock, all roots, and runners. This can be done with a weeding tool on juvenile plants or a pulaski (ax with an adze in one tool that is used for both chopping and digging) on more mature isolated plants.

Since seeds are only responsible for a small portion of the spread of Chinese Wisteria, the plant can be removed at any time. However, it is always best to remove any invasive plant early in the spring or when in flower. If cut while in flower, as long as the flowers are not already turning into fruit, they will not go to seed after being cut.

Disposal of the Shrubs Once Removed

Since the vines can resprout from any nodes, they should either be burned or solarized. Burning is fast and effective. However, it is not allowed in all areas and certainly not in all seasons. To solarize, put the vines under a thick black tarp or into thick black garbage bags and leave them in the full sun for a good 8 weeks to be sure that all seeds are no longer viable. Some sources recommend shorter solarization periods, but in my experience, differences in exposure, latitude, cloud cover, etc, can all lead to differential success. Leave it to solarize as long as possible to ensure they are no longer viable, and then they can be disposed of accordingly.

Chemical Control of Chinese Wisteria

Chemical applications are almost never an ideal method of control for any invasive species. That is because chemical alteration of the environment often makes the environment more suitable for invasive species than native species. Furthermore, it is often difficult to keep the chemical control method contained so that it does not directly affect any native species that are there during the application process itself. As a result, plots where chemical control is used usually show a decrease in species richness. On the other hand, in plots where only physical control is used, species riches significantly increases.

Furthermore, there are no chemical control methods that effectively target only Chinese Wisteria. Chinese Wisteria is known to resist herbicide treatments, so multiple applications are always needed. While this may eventually kill the Wisteria, the chemical side effects often pave the way for other herbicide-resistant invaders to come in.

Chemical control is not recommended.

Biological Control of Chinese Wisteria

Biological control involves the use of a predator, herbivore, disease, or some other agent to control an invasive species once it is established in the environment. The problem with biological control is that the agent used must be entirely specific to only the target organism before releasing it into the environment. This is often difficult to determine since the agent of control is also not native to the environment and could behave differently when released there. Biological control methods are extremely risky and should only be carried out by professionals after years of rigorous study.

Currently, there are no known biological control methods for Chinese Wisteria. No information could be found on even the use of goats which will eat just about anything. Since all parts of the plants are considered poisonous, it is not recommended. Physical control is the preferred method.

Integrated Pest Management & Ongoing Monitoring

Integrated management is always the best approach. In its simplest and least impactful form, this involves physical removal methods, possibly biological control methods, replanting, and ongoing monitoring. Integrated management is required because the area needs to be monitored for returning sprouts or seedlings. Otherwise, all the hard work done in removal could be wasted if the invasive species is allowed to regrow.

Replanting With Native Species is Crucial

In all cases of large patch removal, the site will need to be replanted immediately because the bare soil will allow the seed bank in the soil to germinate and reinvade the patch they were removed from. A replanting program should already be planned and ready to implement immediately upon the removal of the Chinese Wisteria. The area will need to be monitored for any surviving fragments that resprout.

In the case of isolated individuals replating is probably not necessary. Simply remove the individual(s) and monitor over the following 2-5 years to ensure that no rootstock is remaining and resprouting.

Ongoing Monitoring is Essential

In all cases of invasive Chinese Wisteria removal, ongoing monitoring is absolutely essential. In the first year, monitoring should be done every 2-3 weeks to remove the aggressive regrowth from the rootstock. Then in the following 2-5 years, the area should be monitored at least every spring, summer, and early fall to ensure that any surviving individuals are removed so that the population is not able to recover. This is required whether the area is replanted or not. Monitoring will prevent the re-establishment of invasive species and prevent all your hard work in removal from being wasted. And the added bonus is that you get to watch the ecosystem recover.

References and Resources

CABI on Wisteria sinensis https://www.cabi.org/isc/datasheet/56852

Canadensys Plant Search https://data.canadensys.net/vascan/search

Dictionary of Botanical Terms – Lyrae’s Nature Blog Dictionary of Botanical Terms

Fire Effects Information System on Chinese Wisteria https://www.fs.fed.us/database/feis/plants/vine/wisspp/all.html

iNaturalist Plant Search https://www.inaturalist.org/home

USDA Plants Database https://plants.sc.egov.usda.gov/home

Wikipedia on Wisteria frutescens https://en.wikipedia.org/wiki/Wisteria_frutescens

Willis, Lyrae (Unpublished).  Plant Families of North America.

Currently Seeking Funding To Continue This Non-Profit, Ad-Free Work

If you are able to donate so that I can continue this non-profit work of supplying people with scientific information on the plant families, native plants, and invasive species found throughout North America, please donate using the GoFundMe link below. Thank you!